Cladistia is a subclass of bony fishes whose only living members are the bichirs of tropical Africa. Their major synapomorphies are a dorsal fin with independent rays, and a posteriorly elongated parasphenoid.
Cladistia are the earliest diverging branch of living Actinopterygii, and are thought to have diverged from the Actinopteri, the group which includes all other living ray finned fish, by the Carboniferous. However, the fossil range for the only extant order (Polypteriformes) is comparatively young, only reaching as far back as the mid-Cretaceous of South America and Africa, and the two extant genera of bichir only diverged around the Miocene.
Aside from bichirs, other extinct fish groups thought to be members of the group include the Scanilepiformes, known from Triassic (and possibly Permian) of the Northern Hemisphere. The Guildayichthyiformes of Carboniferous North America are also sometimes considered cladistians, but this is thought to be dubious, with other authorities placing them as a stem-group to Neopterygii.
Taxonomy
Based on work done by Near et al (2017) and Lund (2000):
- Order Polypteriformes <small>Bleeker 1859</small>
- Genus â Latinopollia <small>Meunier and Gayet 1998</small>
- Species â Latinopollia suarezi <small>(Meunier & Gayet 1996) Meunier & Gayet 1998</small>
- Genus â Dagetella <small>Gayet & Meunier 1991</small>
- Species â Dagetella sudamericana <small>Gayet & Meunier 1991</small>
- Genus â Bartschichthys <small>Gayet & Meunier 1996</small> [Bartschia <small>Gayet & Meunier 1996 non Rehder 1943</small>]
- Species â B. arnoulti <small>(Gayet & Meunier 1996)</small> [Bartschia arnoulti <small>Gayet & Meunier 1996</small>]
- Species â B. napatensis <small>Werner & Gayet 1997</small>
- Species â B. tubularis <small>(Gayet & Meunier 1996)</small> [Bartschia tubularis <small>Gayet & Meunier 1996</small>]
- Genus â Sudania <small>Werner & Gayet 1997</small>
- Species â S. gracilis <small>Werner & Gayet 1997</small>
- Species â S. oblonga <small>Werner & Gayet 1997</small>
- Genus â Saharichthys <small>Werner & Gayet 1997</small>
- Species â S. nigeriensis <small>(Gayet & Meunier 1996)</small> [Sainthilairia nigeriensis <small>Gayet & Meunier 1996</small>]
- Species â S. africanus <small>(Gayet & Meunier 1996)</small> [Sainthilairia africana <small>Gayet & Meunier 1996</small>]
- Genus â Sainthilairia <small>Gayet & Meunier 1996</small>
- Species â S. beccussiformis <small>Gayet & Meunier 1996</small>
- Species â S. elongata <small>Werner & Gayet 1997</small>
- Species â S. falciformis <small>Gayet & Meunier 1996</small>
- Species â S. grandis <small>Gayet & Meunier 1996</small>
- Species â S. intermedia <small>Werner & Gayet 1997</small>
- Genus â Inbecetemia <small>Werner & Gayet 1997</small>
- Species â I. torta <small>(Gayet & Meunier 1996)</small> [Sainthilairia torta <small>Gayet & Meunier 1996</small>]
- Species â I. tortissima <small>(Gayet & Meunier 1996)</small> [Sainthilairia tortissima <small>Gayet & Meunier 1996</small>]
- Genus â Nagaia <small>Werner & Gayet 1997</small>
- Species â Nagaia extrema <small>Werner & Gayet 1997</small>
- Family Polypteridae <small>Lacépède 1803 sensu stricto</small>
- Genus â Bawitius <small>Grandstaff et al. 2012</small>
- Species â Bawitius bartheli <small>(Schaal 1984) Grandstaff et al. 2012</small> [Polypterus bartheli <small>Schaal 1984</small>]
- Genus â Serenoichthys <small>Dutheil 1999</small>
- Species â Serenoichthys kemkemensis <small>Dutheil 1999</small>
- Genus Erpetoichthys <small>Smith 1865</small> [Calamoichthys <small>Smith 1866</small>]
- Species Erpetoichthys calabaricus
- Genus Polypterus <small>Lacepède 1803</small>
- Species Polypterus ansorgii <small>Boulenger, 1910</small>
- Species Polypterus bichir <small>Lacépède, 1803</small>
- Species Polypterus congicus <small>Boulenger, 1898</small>
- Species Polypterus delhezi <small>Boulenger, 1899</small>
- Species Polypterus endlicherii <small>Heckel, 1847</small>
- Species Polypterus mokelembembe <small>Schliewen & Schäfer, 2006</small>
- Species Polypterus ornatipinnis <small>Boulenger, 1902</small>
- Species Polypterus palmas <small>Ayres, 1850</small>
- Species Polypterus polli <small>J. P. Gosse, 1988</small>
- Species Polypterus retropinnis <small>Vaillant, 1899</small>
- Species Polypterus senegalus <small>Cuvier, 1829</small>
- Species Polypterus teugelsi <small>Britz, 2004</small>
- Species Polypterus weeksii <small>Boulenger, 1898</small>
- Order â Scanilepiformes <small>Sytchevskaya, 1999</small>
- Genus â Beishanichthys <small>Xu and Gao, 2011</small>
- Species â Beishanichthys brevicaudalis <small>Xu and Gao, 2011</small>
- Genus â Fukangichthys <small>Su, 1978</small>
- Species â Fukangichthys longidorsalis <small>Su, 1978</small>
- Genus â Mizhilepis <small>Liu & Shen, 2006</small>
- Species â Mizhilepis zhangyaensis <small>Liu & Shen, 2006</small>
- Genus â Tanaocrossus <small>Schaeffer, 1967</small>
- Species â Tanaocrossus kalliokoskii <small>Schaeffer, 1967</small>
- Family â Scanilepididae <small>Aldinger, 1935</small>
- Genus â Scanilepis <small>Aldinger, 1935</small>
- Species â Scanilepis dubius <small>(Woodward, 1893)</small>
- ?Species â Scanilepis spitzbergensis <small>Aldinger, 1935</small>
- Family â Evenkiidae <small>Selezneva, 1985</small>
- Genus â Evenkia <small>Berg, 1941</small>
- Species â Evenkia eunotoptera <small>Berg, 1941</small>
- Genus â Oshiaichthys <small>Schultze et al., 2021</small>
- Species â Oshiaichthys ferganica <small>Schultze et al., 2021</small>
- Genus â Toyemia <small>Minikh, 1990</small>
- Species â Toyemia blumentalis <small>Minikh, 1995</small>
- Species â Toyemia tverdochlebovi <small>Minikh, 1990</small>
Disputed members
References