List of Onthophagus species

This is a list of 2257 species in the genus Onthophagus. The species and subgenera reflect the Catalog of Life and the World Scarabaeidae Database (2023).

Onthophagus species

Genus Onthophagus <small>Latreille, 1802</small>

Subgenus Afrostrandius <small>Moretto, 2009</small>

Onthophagus adspersus <small>Orbigny, 1908</small>

Onthophagus bimetallicus <small>Orbigny, 1907</small>

Onthophagus demeyeri <small>Moretto, 2009</small>

Onthophagus granulipennis <small>Lansberge, 1886</small>

Onthophagus loxodontaphilus <small>Moretto, 2009</small>

Onthophagus plebejus <small>Klug, 1855</small>

Onthophagus pseudoplebejus <small>Moretto, 2009</small>

Onthophagus tigrinus <small>Orbigny, 1908</small>

Subgenus Altonthophagus <small>Kabakov, 1990</small>

Onthophagus cervenkai <small>Kabakov, 2008</small>

Onthophagus concolor <small>Sharp, 1878</small>

Onthophagus cupreiceps <small>Arrow, 1907</small>

Onthophagus inelegans <small>Balthasar, 1935</small>

Onthophagus kozlovi <small>Kabakov, 1990</small>

Onthophagus kukunorensis <small>Kabakov, 1990</small>

Onthophagus marmotae <small>Kabakov, 1990</small>

Onthophagus sibiricus <small>Harold, 1877</small>

Onthophagus tibetanus <small>Arrow, 1907</small>

Onthophagus turpidoides <small>Kabakov, 2008</small>

Onthophagus turpidus <small>Reitter, 1887</small>

Onthophagus uniformis <small>Heyden, 1886</small>

Subgenus Amphionthophagus <small>Martín-Piera & Zunino, 1983</small>

Onthophagus falzonii <small>Goidanich, 1926</small>

Onthophagus melitaeus <small>(Fabricius, 1798)</small>

Onthophagus numidicus <small>Orbigny, 1908</small>

Subgenus Bicornonthophagus <small>Tagliaferri & Moretto, 2012</small>

Onthophagus auriculatus <small>Klug, 1855</small>

Onthophagus beiranus <small>Péringuey, 1908</small>

Onthophagus cornifrons <small>Thomson, 1858</small>

Onthophagus decorsei <small>Orbigny, 1908</small>

Onthophagus emeritus <small>Péringuey, 1901</small>

Onthophagus latestriatus <small>Orbigny, 1908</small>

Onthophagus reticulatus <small>Orbigny, 1902</small>

Onthophagus rotundibasis <small>Orbigny, 1902</small>

Subgenus Colobonthophagus <small>Balthasar, 1935</small>

Onthophagus aenescens <small>(Wiedemann, 1823)</small>

Onthophagus agnus <small>Gillet, 1925</small>

Onthophagus armatus <small>Blanchard, 1853</small>

Onthophagus arunensis <small>Scheuern, 1995</small>

Onthophagus bengalensis <small>Harold, 1886</small>

Onthophagus bison <small>Boucomont, 1919</small>

Onthophagus caprai <small>Frey, 1956</small>

Onthophagus dama <small>(Fabricius, 1798)</small>

Onthophagus ephippioderus <small>Arrow, 1907</small>

Onthophagus hindu <small>Arrow, 1931</small>

Onthophagus lahorensis <small>Kabakov, 2008</small>

Onthophagus lunatus <small>Harold, 1868</small>

Onthophagus metalliceps <small>Arrow, 1931</small>

Onthophagus nagasawai <small>Matsumura, 1938</small>

Onthophagus neocolobus <small>Scheuern, 1996</small>

Onthophagus occipitalis <small>Lansberge, 1885</small>

Onthophagus paliceps <small>Arrow, 1931</small>

Onthophagus pardalis <small>(Fabricius, 1798)</small>

Onthophagus piceorufulus <small>Kabakov, 1994</small>

Onthophagus piffli <small>Petrovitz, 1961</small>

Onthophagus poggii <small>Scheuern, 1996</small>

Onthophagus quadridentatus <small>(Fabricius, 1798)</small>

Onthophagus ramosellus <small>Bates, 1891</small>

Onthophagus ramosus <small>(Wiedemann, 1823)</small>

Onthophagus shillongensis <small>Scheuern, 1995</small>

Onthophagus thai <small>Kabakov, 1994</small>

Onthophagus tragoides <small>Boucomont, 1914</small>

Onthophagus tragus <small>(Fabricius, 1792)</small>

Onthophagus transquadridentatus <small>Scheuern, 1995</small>

Onthophagus triceratops <small>Arrow, 1913</small>

Onthophagus urellus <small>Boucomont, 1919</small>

Subgenus Cryptonthophagus <small>Dierkens, 2022</small>

Onthophagus convexipennis <small>Dierkens, 2022</small>

Onthophagus flavipleurus <small>Dierkens, 2022</small>

Onthophagus granulopygus <small>Dierkens, 2022</small>

Onthophagus iringaensis <small>Dierkens, 2022</small>

Onthophagus kapangaensis <small>Dierkens, 2022</small>

Onthophagus lilongweensis <small>Dierkens, 2022</small>

Onthophagus mwenseensis <small>Dierkens, 2022</small>

Onthophagus planidorsis <small>Dierkens, 2022</small>

Onthophagus ugandaensis <small>Dierkens, 2022</small>

Subgenus Drepanonthophagus <small>Dierkens, 2022</small>

Onthophagus bellus <small>Orbigny, 1905</small>

Onthophagus gradivus <small>Balthasar, 1966</small>

Onthophagus renejeanneli <small>Dierkens, 2022</small>

Subgenus Endrodius <small>Balthasar, 1959</small>

Onthophagus baeri <small>Boucomont, 1914</small>

Onthophagus gaesatus <small>Boucomont, 1925</small>

Onthophagus praedatus <small>Harold, 1862</small>

Subgenus Eremonthophagus <small>Zunino, 1979</small>

Onthophagus abeillei <small>Orbigny, 1897</small>

Onthophagus heydeni <small>Harold, 1875</small>

Onthophagus infuscatus <small>Klug, 1845</small>

Onthophagus oberthueri <small>Orbigny, 1898</small>

Onthophagus semicinctus <small>Orbigny, 1897</small>

Onthophagus sticticus <small>Harold, 1867</small>

Onthophagus transcaspicus <small>Koenig, 1889</small>

Subgenus Exonthophagus <small>Kabakov, 2006</small>

Onthophagus haroldi <small>Ballion, 1871</small>

Onthophagus viriditinctus <small>Reitter, 1892</small>

Subgenus Furconthophagus <small>Zunino, 1979</small>

Onthophagus aethiopicus <small>Orbigny, 1902</small>

Onthophagus amicus <small>(Gillet, 1925)</small>

Onthophagus boucomonti <small>Paulian, 1931</small>

Onthophagus dapcauensis <small>Boucomont, 1921</small>

Onthophagus flaviclava <small>Orbigny, 1902</small>

Onthophagus frontalis <small>Raffray, 1877</small>

Onthophagus furcatoides <small>Lansberge, 1886</small>

Onthophagus furcatus <small>(Fabricius, 1781)</small>

Onthophagus karenensis <small>Masumoto, Ochi & Hanboonsong, 2008</small>

Onthophagus khonkaenus <small>Masumoto, Ochi & Hanboonsong, 2008</small>

Onthophagus lamelliger <small>Gerstaecker, 1871</small>

Onthophagus liwondeensis <small>Josso, 2019</small>

Onthophagus monforti <small>Josso & Prévost, 2006</small>

Onthophagus papulatorius <small>Kabakov, 2008</small>

Onthophagus papulatus <small>Boucomont, 1914</small>

Onthophagus parvulus <small>(Fabricius, 1798)</small>

Onthophagus promontorii <small>Boucomont, 1924</small>

Onthophagus rugulipennis <small>Fairmaire, 1887</small>

Onthophagus schoolmeestersi <small>Ochi & Kon, 2007</small>

Onthophagus sellatus <small>Klug, 1845</small>

Onthophagus shaykh <small>Montanaro & Ziani, 2022</small>

Onthophagus suffusus <small>Klug, 1855</small>

Onthophagus sugillatus <small>Klug, 1855</small>

Onthophagus troglodyta <small>(Wiedemann, 1823)</small>

Onthophagus ulula <small>Balthasar, 1966</small>

Onthophagus valentineae <small>Josso, 2019</small>

Onthophagus variegatus <small>(Fabricius, 1798)</small>

Onthophagus verae <small>Josso, 2013</small>

Onthophagus versutus <small>Péringuey, 1901</small>

Subgenus Gibbonthophagus <small>Balthasar, 1935</small>

Onthophagus amamiensis <small>Nomura, 1965</small>

Onthophagus apicetinctus <small>Orbigny, 1898</small>

Onthophagus asiaticus <small>Endrödi, 1973</small>

Onthophagus atripennis <small>Waterhouse, 1875</small>

Onthophagus balthasari <small>Všetečka, 1939</small>

Onthophagus bisscrutator <small>Krikken & Huijbregts, 2017</small>

Onthophagus cervicapra <small>Boucomont, 1914</small>

Onthophagus chineicus <small>Kabakov, 1998</small>

Onthophagus denticornis <small>Boucomont, 1914</small>

Onthophagus dubernardi <small>Boucomont, 1914</small>

Onthophagus duporti <small>Boucomont, 1914</small>

Onthophagus euryceros <small>Kabakov, 1998</small>

Onthophagus fujiii <small>Ochi & Kon, 1995</small>

Onthophagus hiabunicus <small>Kabakov, 1998</small>

Onthophagus incollaris <small>Kabakov, 2008</small>

Onthophagus kentingensis <small>Nomura, 1973</small>

Onthophagus kimioi <small>Ochi, Kon & Kawahara, 2011</small>

Onthophagus kiyoshii <small>Ochi & Kon, 2007</small>

Onthophagus laichauensis <small>Ochi, Kon & Pham, 2021</small>

Onthophagus lenis <small>Kabakov, 1998</small>

Onthophagus limbatus <small>(Herbst, 1789)</small>

Onthophagus luridipennis <small>Boheman, 1858</small>

Onthophagus nasalis <small>Arrow, 1931</small>

Onthophagus nigriobscurior <small>Ochi, Kon & Tsubaki, 2009</small>

Onthophagus obscurior <small>Boucomont, 1914</small>

Onthophagus palawanicus <small>Ochi & Kon, 2004</small>

Onthophagus parviobscurior <small>Ochi, Kon & Tsubaki, 2009</small>

Onthophagus penmani <small>Masumoto, Ochi & Hanboonsong, 2002</small>

Onthophagus privus <small>Kabakov, 1998</small>

Onthophagus proletarius <small>Harold, 1875</small>

Onthophagus quangnamensis <small>Ochi, Kon & Pham, 2021</small>

Onthophagus rectecornutus <small>Lansberge, 1883</small>

Onthophagus remotus <small>Harold, 1879</small>

Onthophagus rufiobscurior <small>Ochi, Kon & Tsubaki, 2009</small>

Onthophagus sasajii <small>Ochi & Kon, 2001</small>

Onthophagus schillhammeri <small>Kabakov, 2006</small>

Onthophagus scrutator <small>Harold, 1877</small>

Onthophagus semipersonatus <small>Ochi, Kon & Tsubaki, 2009</small>

Onthophagus solivagus <small>Harold, 1886</small>

Onthophagus subcornutus <small>Boucomont, 1914</small>

Onthophagus sunantaae <small>Masumoto, 1989</small>

Onthophagus susterai <small>Balthasar, 1952</small>

Onthophagus taeniatus <small>Boucomont, 1914</small>

Onthophagus taurinus <small>White, 1844</small>

Onthophagus viduus <small>Harold, 1875</small>

Onthophagus viridicervicapra <small>Ochi, Kon & Tsubaki, 2009</small>

Subgenus Gonocyphus <small>Lansberge, 1885</small>

Onthophagus obliquus <small>(Olivier, 1789)</small>

Onthophagus shimba <small>Cambefort, 1980</small>

Subgenus Hikidaeus <small>Ochi & Kon, 2016</small>

Onthophagus azusae <small>Ochi & Kon, 2006</small>

Onthophagus cupreopastillatus <small>Ochi & Kon, 2006</small>

Onthophagus kanyaayonus <small>Masumoto, 1992</small>

Onthophagus leknamnaous <small>Masumoto & Ochi, 2015</small>

Onthophagus pastillatus <small>Boucomont, 1919</small>

Onthophagus peninsularis <small>Boucomont, 1914</small>

Onthophagus sauteri <small>Gillet, 1924</small>

Onthophagus shirakii <small>Nakane, 1960</small>

Onthophagus simboroni <small>Ochi & Kon, 2006</small>

Onthophagus singhaakhomus <small>Masumoto, 1992</small>

Onthophagus tsutomui <small>Ochi & Kon, 2016</small>

Onthophagus yangmunensis <small>Masumoto & Ochi, 2015</small>

Subgenus Indachorius <small>Balthasar, 1941</small>

Onthophagus aereopictus <small>Boucomont, 1914</small>

Onthophagus arai <small>Masumoto, 1989</small>

Onthophagus baenzigeri <small>Masumoto, Ochi & Hanboonsong, 2007</small>

Onthophagus baykanus <small>Kabakov, 1994</small>

Onthophagus bonengus <small>Kabakov, 1994</small>

Onthophagus chetroiensis <small>Masumoto, Ochi & Sakchoowong, 2012</small>

Onthophagus cheyi <small>Ochi & Kon, 2006</small>

Onthophagus chinbucephalus <small>Ochi & Kon, 2018</small>

Onthophagus chinensis <small>(Balthasar, 1952)</small>

Onthophagus chumphonensis <small>Masumoto, Ochi & Hanboonsong, 2008</small>

Onthophagus clermonti <small>Paulian, 1931</small>

Onthophagus danumensis <small>Ochi, Kon & Barclay, 2009</small>

Onthophagus doisuthepensis <small>Masumoto, 1989</small>

Onthophagus heterorrhinus <small>Lansberge, 1885</small>

Onthophagus hikidai <small>Ochi & Kon, 2006</small>

Onthophagus hsui <small>Masumoto, Chen & Ochi, 2004</small>

Onthophagus jingping <small>Masumoto, Ochi & Hanboonsong, 2007</small>

Onthophagus koshunensis <small>Balthasar, 1941</small>

Onthophagus kyokoae <small>Masumoto & Ochi, 2015</small>

Onthophagus lamyi <small>Paulian, 1945</small>

Onthophagus lannamiibun <small>Masumoto, Ochi & Hanboonsong, 2002</small>

Onthophagus lekmaengtibanphrao <small>Masumoto & Ochi, 2015</small>

Onthophagus liwagensis <small>Ochi & Kon, 2006</small>

Onthophagus maephaluangus <small>Masumoto, Ochi & Hanboonsong, 2007</small>

Onthophagus maesaensis <small>Masumoto, Ochi & Hanboonsong, 2002</small>

Onthophagus magnini <small>Paulian, 1933</small>

Onthophagus maruchanus <small>Masumoto, Ochi & Sakchoowong, 2012</small>

Onthophagus maruyamai <small>Ochi, Kon & Masumoto, 2014</small>

Onthophagus masaoi <small>Ochi, 1992</small>

Onthophagus mendeli <small>Ochi, Kon & Barclay, 2009</small>

Onthophagus mongkhoni <small>Masumoto, Hanboonsong & Ochi, 2002</small>

Onthophagus namnaonus <small>Masumoto, Ochi & Higurashi, 2020</small>

Onthophagus nefarius <small>Balthasar, 1963</small>

Onthophagus nongkaiensis <small>Masumoto, Ochi & Hanboonsong, 2002</small>

Onthophagus octonaevus <small>Kabakov, 1994</small>

Onthophagus paramasaoi <small>Ochi, Kon & Barclay, 2009</small>

Onthophagus perakensis <small>Ochi, Kon & Masumoto, 2014</small>

Onthophagus phahompokus <small>Masumoto & Ochi, 2015</small>

Onthophagus phakuaiensis <small>Masumoto, Ochi & Hanboonsong, 2013</small>

Onthophagus phetchabunensis <small>Masumoto, Ochi & Hanboonsong, 2002</small>

Onthophagus phuquoci <small>Paulian, 1945</small>

Onthophagus platypus <small>Zhang, 1997</small>

Onthophagus pseudoarai <small>Ochi & Kon, 2018</small>

Onthophagus pseudoworoae <small>Ochi, Kon & Barclay, 2009</small>

Onthophagus pumilus <small>Kabakov, 2014</small>

Onthophagus ranongensis <small>Masumoto, Ochi & Hanboonsong, 2008</small>

Onthophagus sabai <small>Masumoto, Ochi & Sakchoowong, 2012</small>

Onthophagus scotti <small>Masumoto, Ochi & Hanboonsong, 2002</small>

Onthophagus semiaereopictus <small>Ochi, Kawahara & Cabras, 2021</small>

Onthophagus semidanumensis <small>Ochi, Kon & Barclay, 2009</small>

Onthophagus semiperakensis <small>Ochi, Kon & Masumoto, 2014</small>

Onthophagus semiworoae <small>Ochi, Kon & Masumoto, 2014</small>

Onthophagus spathatus <small>Boucomont, 1914</small>

Onthophagus subsapaensis <small>Kabakov, 1994</small>

Onthophagus suginoi <small>Ochi, 1984</small>

Onthophagus thungphraensis <small>Masumoto, Ochi & Higurashi, 2020</small>

Onthophagus tiamicus <small>Kabakov, 1994</small>

Onthophagus tongbantumi <small>Masumoto, Ochi & Hanboonsong, 2002</small>

Onthophagus trochilus <small>Arrow, 1931</small>

Onthophagus uedai <small>Ochi & Kon, 2006</small>

Onthophagus ulugombakensis <small>Ochi, Kon & Masumoto, 2014</small>

Onthophagus viridiperakensis <small>Ochi, Kon & Masumoto, 2014</small>

Onthophagus wangtaichieni <small>Masumoto, Ochi & Higurashi, 2020</small>

Onthophagus woroae <small>Ochi & Kon, 2006</small>

Onthophagus yaoi <small>Masumoto, Ochi & Lee, 2014</small>

Onthophagus yumotoi <small>Ochi & Kon, 2006</small>

Subgenus Indonthophagus <small>Kabakov, 2006</small>

Onthophagus ensifer <small>Boucomont, 1914</small>

Onthophagus hastifer <small>Lansberge, 1885</small>

Onthophagus mopsus <small>(Fabricius, 1792)</small>

Onthophagus nitidulus <small>Klug, 1845</small>

Onthophagus turbatus <small>Walker, 1858</small>

Subgenus Macronthophagus <small>Ochi, 2003</small>

Onthophagus cludtsi <small>Ochi, 2003</small>

Onthophagus curvicarinatus <small>Boucomont, 1914</small>

Onthophagus diabolicus <small>Harold, 1877</small>

Onthophagus manipurensis <small>Arrow, 1907</small>

Onthophagus menieri <small>Ochi, 2003</small>

Onthophagus nilgirensis <small>Gillet, 1922</small>

Onthophagus rubricollis <small>Hope, 1831</small>

Onthophagus shinichii <small>Ochi & Kon, 2018</small>

Onthophagus uenoi <small>Ochi, 1995</small>

Subgenus Matashia <small>Matsumura, 1938</small>

Onthophagus apomontanus <small>Ochi, Kon & Cabras, 2023</small>

Onthophagus fossor <small>Arrow, 1931</small>

Onthophagus galeatus <small>Boucomont, 1919</small>

Onthophagus gracilipes <small>Boucomont, 1914</small>

Onthophagus kuluensis <small>Bates, 1891</small>

Onthophagus lutosopictus <small>Fairmaire, 1897</small>

Onthophagus negrosensis <small>Ochi, Kon & Cabras, 2023</small>

Onthophagus ohbayashii <small>Nomura, 1939</small>

Onthophagus samarensis <small>Ochi, Kon & Cabras, 2023</small>

Onthophagus troniceki <small>Balthasar, 1933</small>

Onthophagus yubarinus <small>Matsumura, 1937</small>

Subgenus Micronthophagus <small>Balthasar, 1963</small>

Onthophagus arayai <small>Ochi & Kon, 2007</small>

Onthophagus cavia <small>Boucomont, 1914</small>

Onthophagus chinvigilans <small>Ochi & Kon, 2018</small>

Onthophagus collinsi <small>Krikken & Huijbregts, 1987</small>

Onthophagus depressipennis <small>Cambefort, 1975</small>

Onthophagus drescheri <small>Paulian, 1939</small>

Onthophagus falsivigilans <small>Masumoto, 1995</small>

Onthophagus fukuyamai <small>Ochi, Kon & Ueda, 2018</small>

Onthophagus gigantivigilans <small>Masumoto, Hanboonsong & Ochi, 2002</small>

Onthophagus gulo <small>Arrow, 1931</small>

Onthophagus hystrix <small>Boucomont, 1914</small>

Onthophagus incertus <small>Orbigny, 1897</small>

Onthophagus konoi <small>Matsumura, 1938</small>

Onthophagus liui <small>Masumoto, Ochi & Lee, 2014</small>

Onthophagus melanocephalus <small>Klug, 1845</small>

Onthophagus mogo <small>Krikken & Huijbregts, 2008</small>

Onthophagus ochreatus <small>Orbigny, 1897</small>

Onthophagus oculatus <small>Arrow, 1931</small>

Onthophagus paroculus <small>Krikken & Huijbregts, 1987</small>

Onthophagus phillippsorum <small>Krikken & Huijbregts, 1987</small>

Onthophagus rotundicollis <small>Lansberge, 1883</small>

Onthophagus setoculus <small>Krikken & Huijbregts, 1987</small>

Onthophagus sideki <small>Krikken & Huijbregts, 1987</small>

Onthophagus sinagai <small>Krikken & Huijbregts, 2008</small>

Onthophagus taoi <small>Ochi & Kon, 2006</small>

Onthophagus vanasseni <small>Krikken & Huijbregts, 2008</small>

Onthophagus vanderblomi <small>Krikken & Huijbregts, 2008</small>

Onthophagus vanofwegeni <small>Krikken & Huijbregts, 2008</small>

Onthophagus vigilans <small>Boucomont, 1921</small>

Onthophagus wangi <small>Masumoto, Chen & Ochi, 2004</small>

Onthophagus watuwila <small>Krikken & Huijbregts, 2008</small>

Subgenus Onthophagiellus <small>Balthasar, 1935</small>

Onthophagus abei <small>Ochi & Kon, 2015</small>

Onthophagus andonarensis <small>Boucomont, 1914</small>

Onthophagus andreji <small>Prokofiev, 2014</small>

Onthophagus crassicollis <small>Boucomont, 1913</small>

Onthophagus deliensis <small>Lansberge, 1885</small>

Onthophagus falculatus <small>Boucomont, 1914</small>

Onthophagus hanboonsongae <small>Masumoto, Ochi & Higurashi, 2021</small>

Onthophagus hidakai <small>Ochi & Kon, 1995</small>

Onthophagus inermivertex <small>Boucomont, 1921</small>

Onthophagus jugicola <small>Orbigny, 1902</small>

Onthophagus kangeanus <small>Paulian, 1936</small>

Onthophagus kapitensis <small>Frey, 1971</small>

Onthophagus kawaharai <small>Ochi & Kon, 2007</small>

Onthophagus kondaoensis <small>Kabakov, 1994</small>

Onthophagus millingeni <small>Orbigny, 1898</small>

Onthophagus opacifalculatus <small>Ochi, Kon & Tsubaki, 2009</small>

Onthophagus parafalculatus <small>Ochi, Kon & Tsubaki, 2009</small>

Onthophagus popovi <small>Kabakov, 1983</small>

Onthophagus solmani <small>Stebnicka, 1975</small>

Onthophagus suginokoichii <small>Ochi & Kon, 2008</small>

Onthophagus sumatramontanus <small>Ochi & Kon, 2008</small>

Onthophagus taiyaruensis <small>Masumoto, 1977</small>

Onthophagus thanwaakhomus <small>Masumoto, 1992</small>

Onthophagus tridentitibialis <small>Ochi & Kon, 2008</small>

Onthophagus unguiculatus <small>Kabakov, 1994</small>

Subgenus Onthophagus <small>Latreille, 1802</small>

Onthophagus abas <small>Balthasar, 1946</small>

Onthophagus abmisibilus <small>Krikken & Huijbregts, 2012</small>

Onthophagus abreui <small>Arrow, 1931</small>

Onthophagus abruptus <small>Orbigny, 1913</small>

Onthophagus absyrtus <small>Balthasar, 1946</small>

Onthophagus abyssinicus <small>Gillet, 1925</small>

Onthophagus academus <small>Balthasar, 1946</small>

Onthophagus acernorus <small>Moctezuma, Hernández & Sánchez-Huerta, 2021</small>

Onthophagus aceroides <small>Krikken & Huijbregts, 2013</small>

Onthophagus acerus <small>Gillet, 1930</small>

Onthophagus aciculatulus <small>Blatchley, 1928</small>

Onthophagus acrisius <small>Balthasar, 1946</small>

Onthophagus acuminatus <small>Harold, 1880</small>

Onthophagus adelaidae <small>Hope, 1846</small>

Onthophagus adelphus <small>Gillet, 1930</small>

Onthophagus admetus <small>Balthasar, 1946</small>

Onthophagus adornatus <small>Orbigny, 1904</small>

Onthophagus aegrotus <small>Balthasar, 1967</small>

Onthophagus aemulus <small>Gillet, 1930</small>

Onthophagus aeneoniger <small>Orbigny, 1913</small>

Onthophagus aeneopiceus <small>Orbigny, 1902</small>

Onthophagus aequepubens <small>Orbigny, 1905</small>

Onthophagus aereidorsis <small>Orbigny, 1902</small>

Onthophagus aeremicans <small>Orbigny, 1904</small>

Onthophagus aereomaculatus <small>Boucomont, 1914</small>

Onthophagus aerestriatus <small>Orbigny, 1913</small>

Onthophagus aeruginosus <small>Roth, 1851</small>

Onthophagus aesopus <small>Lansberge, 1882</small>

Onthophagus affinis <small>Gillet, 1930</small>

Onthophagus africanus <small>Lansberge, 1886</small>

Onthophagus agaricophilus <small>Arrow, 1931</small>

Onthophagus ahenicollis <small>Orbigny, 1902</small>

Onthophagus ahenomicans <small>Orbigny, 1902</small>

Onthophagus akamineae <small>Moretto & Génier, 2020</small>

Onthophagus akhaus <small>Masumoto, Ochi & Hanboonsong, 2007</small>

Onthophagus alaindrumonti <small>Krikken & Huijbregts, 2012</small>

Onthophagus alainmonforti <small>Josso, 2022</small>

Onthophagus albicomus <small>Orbigny, 1908</small>

Onthophagus albicornis <small>(Palisot de Beauvois, 1805)</small>

Onthophagus albipennis <small>Péringuey, 1908</small>

Onthophagus albipodex <small>Orbigny, 1902</small>

Onthophagus alfuricus <small>Huijbregts & Krikken, 2012</small>

Onthophagus aliceae <small>Josso, 2019</small>

Onthophagus alienus <small>Frey, 1958</small>

Onthophagus allojavanus <small>Huijbregts & Krikken, 2011</small>

Onthophagus alluaudi <small>Orbigny, 1902</small>

Onthophagus alluvius <small>Howden & Cartwright, 1963</small>

Onthophagus aloysiellus <small>Zunino, 1977</small>

Onthophagus alquirta <small>Matthews, 1972</small>

Onthophagus alternans <small>Raffray, 1877</small>

Onthophagus altilamina <small>Orbigny, 1905</small>

Onthophagus altivagans <small>Howden & Génier, 2004</small>

Onthophagus ambang <small>Krikken & Huijbregts, 2011</small>

Onthophagus amoenus <small>Orbigny, 1908</small>

Onthophagus amphicoma <small>Boucomont, 1914</small>

Onthophagus amphinasus <small>Arrow, 1931</small>

Onthophagus amphioxus <small>Arrow, 1933</small>

Onthophagus amplipennis <small>Orbigny, 1905</small>

Onthophagus amycoides <small>Kabakov, 2014</small>

Onthophagus anchommatus <small>Lea, 1923</small>

Onthophagus andersoni <small>Howden & Gill, 1987</small>

Onthophagus andrewesi <small>Arrow, 1931</small>

Onthophagus andrewsmithi <small>Krikken & Huijbregts, 2012</small>

Onthophagus androgynus <small>Orbigny, 1905</small>

Onthophagus anewtoni <small>Howden & Génier, 2004</small>

Onthophagus angolensis <small>Paulian, 1937</small>

Onthophagus angularis <small>Orbigny, 1908</small>

Onthophagus angulicornis <small>Orbigny, 1908</small>

Onthophagus anisocerus <small>Erichson, 1842</small>

Onthophagus annoyeri <small>Moretto, 2010</small>

Onthophagus annulopunctatus <small>Krikken & Huijbregts, 2017</small>

Onthophagus anogeissii <small>Cambefort, 1984</small>

Onthophagus anomalicollis <small>Frey, 1973</small>

Onthophagus anomalipes <small>Orbigny, 1902</small>

Onthophagus antecornutus <small>Josso, 2019</small>

Onthophagus antennalis <small>Frey, 1961</small>

Onthophagus anthracinus <small>Harold, 1873</small>

Onthophagus antillarum <small>Arrow, 1903</small>

Onthophagus antilocapra <small>Balthasar, 1937</small>

Onthophagus antoinei <small>Walter, 1990</small>

Onthophagus antoineianus <small>Josso, 2022</small>

Onthophagus aphodioides <small>Lansberge, 1883</small>

Onthophagus apiciosus <small>Orbigny, 1902</small>

Onthophagus apterus <small>Matthews, 1972</small>

Onthophagus apunneea <small>Masumoto, Ochi & Hanboonsong, 2007</small>

Onthophagus arboreus <small>Arrow, 1931</small>

Onthophagus arcifer <small>Orbigny, 1904</small>

Onthophagus areolatus <small>Orbigny, 1905</small>

Onthophagus arkoola <small>Storey & Weir, 1990</small>

Onthophagus arnetti <small>Howden & Cartwright, 1963</small>

Onthophagus arrilla <small>Matthews, 1972</small>

Onthophagus arunachalensis <small>Biswas & Chatterjee, 1985</small>

Onthophagus aschenborni <small>Frey, 1975</small>

Onthophagus ashanticola <small>Frey, 1973</small>

Onthophagus asimilis <small>Péringuey, 1901</small>

Onthophagus asper <small>MacLeay, 1864</small>

Onthophagus aspericeps <small>Orbigny, 1908</small>

Onthophagus aspericollis <small>Lansberge, 1885</small>

Onthophagus asperipennis <small>Orbigny, 1902</small>

Onthophagus aspernatus <small>Orbigny, 1907</small>

Onthophagus asperodorsatus <small>Howden & Gill, 1993</small>

Onthophagus asperulus <small>Orbigny, 1905</small>

Onthophagus astigma <small>Orbigny, 1904</small>

Onthophagus athiensis <small>Orbigny, 1913</small>

Onthophagus atricolor <small>Orbigny, 1907</small>

Onthophagus atridorsis <small>Orbigny, 1902</small>

Onthophagus atriglabrus <small>Howden & Gill, 1987</small>

Onthophagus atrofasciatus <small>Orbigny, 1905</small>

Onthophagus atronitidus <small>Orbigny, 1902</small>

Onthophagus atrosericeus <small>Boucomont, 1932</small>

Onthophagus atrosplendens <small>Moretto, 2017</small>

Onthophagus atrostriatus <small>Orbigny, 1913</small>

Onthophagus atrovirens <small>Orbigny, 1905</small>

Onthophagus atrox <small>Harold, 1867</small>

Onthophagus aureofuscus <small>Bates, 1887</small>

Onthophagus aureopilosus <small>Boucomont, 1914</small>

Onthophagus auritus <small>Erichson, 1842</small>

Onthophagus australis <small>Guérin-Méneville, 1830</small> - (Southern Dung Beetle)

Onthophagus axillaris <small>Boheman, 1860</small>

Onthophagus aztecus <small>Zunino & Halffter, 1988</small>

Onthophagus babaulti <small>Orbigny, 1915</small>

Onthophagus babirussa <small>(Eschscholtz, 1822)</small>

Onthophagus baiyericus <small>Krikken & Huijbregts, 2013</small>

Onthophagus bajacalifornianus <small>Moctezuma & Halffter, 2019</small>

Onthophagus bakweri <small>Moretto, 2014</small>

Onthophagus balawaicus <small>Scheuern, 1995</small>

Onthophagus baloghi <small>Balthasar, 1967</small>

Onthophagus bambra <small>Matthews, 1972</small>

Onthophagus bandamai <small>Cambefort, 1984</small>

Onthophagus baoule <small>Cambefort, 1984</small>

Onthophagus barretti <small>Génier & Howden, 1999</small>

Onthophagus barriorum <small>Walter, 1990</small>

Onthophagus bartosi <small>Balthasar, 1966</small>

Onthophagus basakata <small>Walter & Cambefort, 1977</small>

Onthophagus basicarinatus <small>Rossini, Vaz-de-Mello & Zunino, 2018</small>

Onthophagus basilewskyi <small>Frey, 1961</small>

Onthophagus bassariscus <small>Zunino & Halffter, 1988</small>

Onthophagus bateke <small>Walter & Cambefort, 1977</small>

Onthophagus batesi <small>Howden & Cartwright, 1963</small>

Onthophagus batillifer <small>Harold, 1875</small>

Onthophagus batui <small>Huijbregts & Krikken, 2009</small>

Onthophagus bayeri <small>Balthasar, 1942</small>

Onthophagus bechynei <small>Frey, 1953</small>

Onthophagus beelarong <small>Storey & Weir, 1990</small>

Onthophagus beesoni <small>Arrow, 1931</small>

Onthophagus begoniophilus <small>Krikken & Huijbregts, 2017</small>

Onthophagus belinga <small>Walter, 1989</small>

Onthophagus belorhinus <small>Bates, 1887</small>

Onthophagus benedictorum <small>Walter & Cambefort, 1977</small>

Onthophagus bengali <small>Gordon & Oppenheimer, 1977</small>

Onthophagus benguellianus <small>Paulian, 1937</small>

Onthophagus bennigseni <small>Paulian, 1937</small>

Onthophagus bequaerti <small>Orbigny, 1915</small>

Onthophagus bergeri <small>Frey, 1975</small>

Onthophagus bernaudi <small>Cambefort & Nicolas, 1991</small>

Onthophagus betschuanus <small>Frey, 1975</small>

Onthophagus bicallifrons <small>Orbigny, 1902</small>

Onthophagus bicarinaticeps <small>Lea, 1923</small>

Onthophagus bicarinatus <small>Gomes Alves, 1944</small>

Onthophagus bicavicollis <small>Lea, 1923</small>

Onthophagus bicavifrons <small>Orbigny, 1902</small>

Onthophagus bicolensis <small>Ochi & Kon, 2006</small>

Onthophagus bicolor <small>Raffray, 1877</small>

Onthophagus biconifer <small>Orbigny, 1905</small>

Onthophagus bicristatus <small>Orbigny, 1905</small>

Onthophagus bicristiger <small>Orbigny, 1913</small>

Onthophagus bicuneus <small>Kabakov, 2008</small>

Onthophagus bidens <small>(Olivier, 1789)</small>

Onthophagus bidentatus <small>Drapiez, 1819</small>

Onthophagus bidentifrons <small>Orbigny, 1902</small>

Onthophagus bifidicornis <small>Orbigny, 1902</small>

Onthophagus bifidus <small>Reiche, 1850</small>

Onthophagus bifrons <small>Orbigny, 1905</small>

Onthophagus bilingula <small>Balthasar, 1965</small>

Onthophagus bimarginatus <small>Orbigny, 1902</small>

Onthophagus bindaree <small>Storey & Weir, 1990</small>

Onthophagus binodis <small>(Thunberg, 1818)</small> - (Humpbacked Dung Beetle)

Onthophagus binodosus <small>Orbigny, 1908</small>

Onthophagus binodulus <small>Orbigny, 1913</small>

Onthophagus binyana <small>Storey & Weir, 1990</small>

Onthophagus biplagiatus <small>Thomson, 1858</small>

Onthophagus birugatus <small>Orbigny, 1902</small>

Onthophagus birugifer <small>Orbigny, 1908</small>

Onthophagus bisbicornis <small>Orbigny, 1909</small>

Onthophagus biscarinulatus <small>Huijbregts & Krikken, 2012</small>

Onthophagus bisectus <small>Arrow, 1931</small>

Onthophagus bisignatus <small>Orbigny, 1913</small>

Onthophagus bistiniocelloides <small>Krikken, 1986</small>

Onthophagus bituberans <small>Orbigny, 1905</small>

Onthophagus bituberifrons <small>Josso, 2018</small>

Onthophagus bituberoculus <small>Krikken & Huijbregts, 2013</small>

Onthophagus bivertex <small>Heyden, 1887</small>

Onthophagus blackburni <small>Shipp, 1895</small>

Onthophagus blackwoodensis <small>Blackburn, 1892</small>

Onthophagus blanchardi <small>Harold, 1869</small>

Onthophagus bocandei <small>Orbigny, 1904</small>

Onthophagus bokiaunus <small>Masumoto, 1995</small>

Onthophagus bolivari <small>Moctezuma, Rossini & Zunino, 2016</small>

Onthophagus bomberaianus <small>Balthasar, 1969</small>

Onthophagus bongkudai <small>Krikken & Huijbregts, 2017</small>

Onthophagus bonorae <small>Zunino, 1976</small>

Onthophagus boops <small>Orbigny, 1905</small>

Onthophagus borassi <small>Cambefort, 1984</small>

Onthophagus bordati <small>Moretto, 2014</small>

Onthophagus borneensis <small>Harold, 1877</small>

Onthophagus bornemisszai <small>Matthews, 1972</small>

Onthophagus bornemisszanus <small>Matthews, 1972</small>

Onthophagus boucomontianus <small>Balthasar, 1935</small>

Onthophagus bourgognei <small>Paulian, 1945</small>

Onthophagus bovinus <small>Péringuey, 1892</small>

Onthophagus brachypterus <small>Zunino & Halffter, 1997</small>

Onthophagus brazzavillianus <small>Balthasar, 1974</small>

Onthophagus breviceps <small>Orbigny, 1902</small>

Onthophagus brevicollis <small>Arrow, 1907</small>

Onthophagus breviconus <small>Génier & Howden, 1999</small>

Onthophagus brevifrons <small>Horn, 1881</small>

Onthophagus brevigena <small>Orbigny, 1902</small>

Onthophagus brevipennis <small>Orbigny, 1902</small>

Onthophagus brevisetis <small>Orbigny, 1915</small>

Onthophagus brittoni <small>Paulian, 1948</small>

Onthophagus brivioi <small>Frey, 1973</small>

Onthophagus bronzeus <small>Arrow, 1907</small>

Onthophagus brooksi <small>Matthews, 1972</small>

Onthophagus browni <small>Howden & Cartwright, 1963</small>

Onthophagus brunellii <small>Müller, 1942</small>

Onthophagus brutus <small>Arrow, 1931</small>

Onthophagus buculus <small>Mannerheim, 1828</small>

Onthophagus bufonidius <small>Josso, 2019</small>

Onthophagus bufulus <small>Arrow, 1941</small>

Onthophagus bulga <small>Reid & Runagall-McNaull, 2022</small>

Onthophagus bunamin <small>Matthews, 1972</small>

Onthophagus bundara <small>Storey & Weir, 1990</small>

Onthophagus burchelli <small>Orbigny, 1908</small>

Onthophagus busiris <small>Balthasar, 1969</small>

Onthophagus caelator <small>Balthasar, 1967</small>

Onthophagus caesariatus <small>Boucomont, 1921</small>

Onthophagus calamophilus <small>Krikken, 1977</small>

Onthophagus calcaratus <small>Boucomont, 1919</small>

Onthophagus calchas <small>Balthasar, 1967</small>

Onthophagus calliger <small>Orbigny, 1913</small>

Onthophagus callosipennis <small>Boucomont, 1930</small>

Onthophagus cambrai <small>Delgado & Curoe, 2014</small>

Onthophagus camerunicus <small>Orbigny, 1905</small>

Onthophagus camilleae <small>Josso, 2022</small>

Onthophagus campestris <small>Masumoto, Kiuchi & Ho, 2018</small>

Onthophagus cancer <small>Lansberge, 1886</small>

Onthophagus canelasensis <small>Howden & Génier, 2004</small>

Onthophagus canescens <small>Zunino & Halffter, 1988</small>

Onthophagus capella <small>Kirby, 1818</small>

Onthophagus capelliformis <small>Gillet, 1925</small>

Onthophagus capellinus <small>Frey, 1963</small>

Onthophagus capitatus <small>Castelnau, 1840</small>

Onthophagus capitosus <small>Harold, 1867</small>

Onthophagus carayoni <small>Cambefort, 1986</small>

Onthophagus carcharias <small>Harold, 1875</small>

Onthophagus carinicollis <small>Raffray, 1877</small>

Onthophagus carinidorsis <small>Orbigny, 1908</small>

Onthophagus carinulatus <small>Harold, 1877</small>

Onthophagus carmodensis <small>Blackburn, 1907</small>

Onthophagus carpophilus <small>Pereira & Halffter, 1961</small>

Onthophagus cartwrighti <small>Howden, 1973</small>

Onthophagus castetsi <small>Lansberge, 1887</small>

Onthophagus catenatus <small>Lansberge, 1883</small>

Onthophagus catharinensis <small>Paulian, 1936</small>

Onthophagus catherineae <small>Josso, 2022</small>

Onthophagus cavernicollis <small>Howden & Cartwright, 1963</small>

Onthophagus caviceps <small>Frey, 1962</small>

Onthophagus cavifrons <small>Harold, 1886</small>

Onthophagus cavivertex <small>Orbigny, 1913</small>

Onthophagus centricornis <small>(Fabricius, 1798)</small>

Onthophagus centurio <small>Lansberge, 1885</small>

Onthophagus cephalophi <small>Cambefort, 1984</small>

Onthophagus cervus <small>(Fabricius, 1798)</small>

Onthophagus ceylonicus <small>Harold, 1859</small>

Onthophagus chacoensis <small>Rossini, Vaz-de-Mello & Zunino, 2018</small>

Onthophagus chaiyaphumensis <small>Masumoto, Ochi & Hanboonsong, 2002</small>

Onthophagus championi <small>Bates, 1887</small>

Onthophagus chebaicus <small>Boucomont, 1919</small>

Onthophagus cheesmanae <small>Arrow, 1941</small>

Onthophagus chepara <small>Matthews, 1972</small>

Onthophagus chevrolati <small>Harold, 1869</small>

Onthophagus chiangdaolekus <small>Masumoto, Ochi & Higurashi, 2020</small>

Onthophagus chiapanecus <small>Zunino & Halffter, 1988</small>

Onthophagus chilapensis <small>Gasca-Álvarez, Zunino & Deloya, 2018</small>

Onthophagus chimalapensis <small>Delgado & Mora-Aguilar, 2019</small>

Onthophagus chinantecus <small>Moctezuma & Halffter, 2019</small>

Onthophagus chirindaensis <small>Josso, 2022</small>

Onthophagus chirindanus <small>Orbigny, 1913</small>

Onthophagus chitipaensis <small>Josso, 2019</small>

Onthophagus chlorophanus <small>Orbigny, 1902</small>

Onthophagus choanicus <small>Orbigny, 1904</small>

Onthophagus chremes <small>Balthasar, 1969</small>

Onthophagus chryses <small>Bates, 1887</small>

Onthophagus chrysurus <small>Arrow, 1931</small>

Onthophagus cicer <small>Balthasar, 1964</small>

Onthophagus cincticollis <small>Orbigny, 1902</small>

Onthophagus cinctifrons <small>Orbigny, 1902</small>

Onthophagus cinctipennis <small>Quedenfeldt, 1884</small>

Onthophagus cineraceus <small>Orbigny, 1902</small>

Onthophagus circulator <small>Reitter, 1891</small>

Onthophagus circulifer <small>Arrow, 1931</small>

Onthophagus circumdatus <small>Orbigny, 1913</small>

Onthophagus citreum <small>Boucomont, 1919</small>

Onthophagus civettae <small>Cambefort, 1984</small>

Onthophagus civitasorum <small>Moctezuma & Halffter, 2020</small>

Onthophagus clavijeroi <small>Moctezuma, Rossini & Zunino, 2016</small>

Onthophagus clavisetis <small>Orbigny, 1905</small>

Onthophagus clitellarius <small>Orbigny, 1908</small>

Onthophagus clivimerus <small>Huijbregts & Krikken, 2011</small>

Onthophagus clusifrons <small>Orbigny, 1905</small>

Onthophagus clypealis <small>Lea, 1923</small>

Onthophagus clypeatus <small>Blanchard, 1845</small>

Onthophagus coahuilae <small>Zunino & Halffter, 1988</small>

Onthophagus cochisus <small>Brown, 1927</small>

Onthophagus coeruleicollis <small>Arrow, 1907</small>

Onthophagus cognatus <small>Boucomont, 1921</small>

Onthophagus coiffaiti <small>Frey, 1970</small>

Onthophagus colasi <small>Paulian, 1937</small>

Onthophagus colffsi <small>Lansberge, 1883</small>

Onthophagus collaris <small>Kabakov, 1983</small>

Onthophagus comatulus <small>Orbigny, 1905</small>

Onthophagus compactus <small>Arrow, 1933</small>

Onthophagus comperei <small>Blackburn, 1903</small>

Onthophagus compositus <small>Lea, 1923</small>

Onthophagus compressus <small>Guérin-Méneville, 1855</small>

Onthophagus concinnus <small>Castelnau, 1840</small>

Onthophagus confertus <small>Péringuey, 1908</small>

Onthophagus confluens <small>Orbigny, 1902</small>

Onthophagus confusus <small>Boucomont, 1932</small>

Onthophagus conradsi <small>Balthasar, 1937</small>

Onthophagus consentaneus <small>Harold, 1867</small> - (Comon Northern Dung Beetle)

Onthophagus conspicuus <small>MacLeay, 1864</small>

Onthophagus contiguicornis <small>Orbigny, 1913</small>

Onthophagus contrapositus <small>Génier, 2017</small>

Onthophagus convexicollis <small>Boheman, 1858</small>

Onthophagus convexus <small>Orbigny, 1908</small>

Onthophagus cooloola <small>Storey & Weir, 1990</small>

Onthophagus coomani <small>Paulian, 1931</small>

Onthophagus coorgensis <small>Arrow, 1931</small>

Onthophagus coprimorphoides <small>Huijbregts & Krikken, 2012</small>

Onthophagus coprimorphus <small>Gillet, 1930</small>

Onthophagus coproides <small>Horn, 1881</small>

Onthophagus coptorhinodes <small>Péringuey, 1901</small>

Onthophagus coracinoides <small>Kabakov, 1983</small>

Onthophagus coracinus <small>Boucomont, 1914</small>

Onthophagus coriaceoumbrosus <small>Kohlmann & Solis, 2001</small>

Onthophagus corniculiger <small>Orbigny, 1913</small>

Onthophagus cornutus <small>Ferreira, 1971</small>

Onthophagus coronatus <small>Orbigny, 1902</small>

Onthophagus corrosus <small>Bates, 1887</small>

Onthophagus coscineus <small>Bates, 1887</small>

Onthophagus costatus <small>Orbigny, 1913</small>

Onthophagus costifer <small>Gillet, 1930</small>

Onthophagus costiger <small>Orbigny, 1915</small>

Onthophagus costilatus <small>Orbigny, 1910</small>

Onthophagus costulicollis <small>Frey, 1975</small>

Onthophagus crantor <small>Balthasar, 1967</small>

Onthophagus cratippus <small>Balthasar, 1969</small>

Onthophagus creber <small>Orbigny, 1905</small>

Onthophagus cretus <small>Péringuey, 1901</small>

Onthophagus cribellum <small>Orbigny, 1902</small>

Onthophagus cribratus <small>Lansberge, 1883</small>

Onthophagus cribripennis <small>Orbigny, 1902</small>

Onthophagus criniger <small>Orbigny, 1904</small>

Onthophagus crinitus <small>Harold, 1869</small>

Onthophagus cristatus <small>Orbigny, 1905</small>

Onthophagus croesulus <small>Bates, 1888</small>

Onthophagus crotchi <small>Harold, 1871</small>

Onthophagus crucenotatus <small>Orbigny, 1905</small>

Onthophagus cruciger <small>MacLeay, 1888</small>

Onthophagus cruentatus <small>Klug, 1855</small>

Onthophagus cryptodicranius <small>Kohlmann & Solis, 2001</small>

Onthophagus cryptogenus <small>Boucomont, 1914</small>

Onthophagus cuboidalis <small>Bates, 1887</small>

Onthophagus cuevensis <small>Howden, 1973</small>

Onthophagus cuneus <small>Boucomont, 1924</small>

Onthophagus cuniculus <small>MacLeay, 1864</small>

Onthophagus cupreopinguis <small>Josso, 2022</small>

Onthophagus cupreovirens <small>Orbigny, 1904</small>

Onthophagus cupreus <small>Harold, 1880</small>

Onthophagus cupricollis <small>Péringuey, 1888</small>

Onthophagus curtulus <small>Frey, 1961</small>

Onthophagus curvicornis <small>Latreille, 1812</small>

Onthophagus curvifrons <small>Orbigny, 1906</small>

Onthophagus curvilamina <small>Müller, 1942</small>

Onthophagus curvispina <small>Reitter, 1892</small>

Onthophagus cyaneiceps <small>Orbigny, 1913</small>

Onthophagus cyanellus <small>Bates, 1887</small>

Onthophagus cyaneoniger <small>Orbigny, 1902</small>

Onthophagus cyanochlorus <small>Orbigny, 1902</small>

Onthophagus cyclographus <small>Bates, 1887</small>

Onthophagus cynomysi <small>Brown, 1927</small>

Onthophagus cyobioides <small>Boucomont, 1921</small>

Onthophagus dandalu <small>Matthews, 1972</small>

Onthophagus danumicus <small>Huijbregts & Krikken, 2012</small>

Onthophagus dapitanensis <small>Boucomont, 1919</small>

Onthophagus darlingtoni <small>Matthews, 1972</small>

Onthophagus davisi <small>Frey, 1975</small>

Onthophagus dayacus <small>Boucomont, 1914</small>

Onthophagus daymanus <small>Krikken & Huijbregts, 2013</small>

Onthophagus debilis <small>Orbigny, 1905</small>

Onthophagus deccanensis <small>Frey, 1962</small>

Onthophagus decedens <small>Péringuey, 1904</small>

Onthophagus decens <small>Balthasar, 1964</small>

Onthophagus declivicollis <small>Orbigny, 1902</small>

Onthophagus declivis <small>Harold, 1869</small>

Onthophagus decolor <small>Orbigny, 1902</small>

Onthophagus decoratus <small>Orbigny, 1905</small>

Onthophagus dedecor <small>Wallengren, 1881</small>

Onthophagus deflexicollis <small>Lansberge, 1883</small>

Onthophagus deflexus <small>Orbigny, 1908</small>

Onthophagus delicatulus <small>Raffray, 1877</small>

Onthophagus delicatus <small>Orbigny, 1915</small>

Onthophagus delphinensis <small>Orbigny, 1913</small>

Onthophagus demarzi <small>Frey, 1959</small>

Onthophagus densatus <small>Frey, 1963</small>

Onthophagus densepunctatus <small>Frey, 1963</small>

Onthophagus densipilis <small>Orbigny, 1902</small>

Onthophagus denticollis <small>Lansberge, 1883</small>

Onthophagus denticulatus <small>Orbigny, 1902</small>

Onthophagus denudatus <small>Orbigny, 1902</small>

Onthophagus depilatus <small>Orbigny, 1913</small>

Onthophagus depilis <small>Orbigny, 1902</small>

Onthophagus deplanatus <small>Lansberge, 1883</small>

Onthophagus depressicollis <small>Boucomont, 1914</small>

Onthophagus depressifrons <small>Orbigny, 1915</small>

Onthophagus derasus <small>Orbigny, 1902</small>

Onthophagus desaegeri <small>Frey, 1961</small>

Onthophagus desectus <small>MacLeay, 1871</small>

Onthophagus deterrens <small>Péringuey, 1901</small>

Onthophagus devagiriensis <small>Schoolmeesters & Thomas, 2006</small>

Onthophagus devexicornis <small>Gillet, 1930</small>

Onthophagus devexus <small>MacLeay, 1888</small>

Onthophagus dhanjuricus <small>Frey, 1973</small>

Onthophagus diadematus <small>Orbigny, 1902</small>

Onthophagus dicax <small>Balthasar, 1966</small>

Onthophagus dicella <small>Bates, 1888</small>

Onthophagus dicranius <small>Bates, 1887</small>

Onthophagus dicranocerus <small>Gillet, 1925</small>

Onthophagus dicranoides <small>Balthasar, 1939</small>

Onthophagus difficilis <small>Le Guillou, 1844</small>

Onthophagus digitifer <small>Boucomont, 1932</small>

Onthophagus dignus <small>Gillet, 1930</small>

Onthophagus dilutus <small>Orbigny, 1905</small>

Onthophagus dinjeera <small>Storey & Weir, 1990</small>

Onthophagus dinoderus <small>Orbigny, 1913</small>

Onthophagus discolor <small>Hope, 1841</small>

Onthophagus discovirens <small>Orbigny, 1902</small>

Onthophagus discretus <small>Péringuey, 1901</small>

Onthophagus dissentaneus <small>Balthasar, 1964</small>

Onthophagus dissidentatus <small>Krikken & Huijbregts, 2013</small>

Onthophagus distichus <small>Roth, 1851</small>

Onthophagus ditus <small>Péringuey, 1901</small>

Onthophagus diversus <small>Reiche, 1850</small>

Onthophagus dlabolai <small>Balthasar, 1964</small>

Onthophagus dohertyi <small>Orbigny, 1902</small>

Onthophagus doiinthanonensis <small>Masumoto, 1989</small>

Onthophagus doipuiensis <small>Masumoto, 1989</small>

Onthophagus doitungensis <small>Masumoto, Ochi & Hanboonsong, 2002</small>

Onthophagus doriae <small>Harold, 1877</small>

Onthophagus dorsipilulus <small>Howden & Gill, 1987</small>

Onthophagus dorsuosus <small>Orbigny, 1902</small>

Onthophagus drumonti <small>Moretto & Génier, 2020</small>

Onthophagus dryander <small>Monteith & Storey, 2013</small>

Onthophagus dubitabilis <small>Howden & Génier, 2004</small>

Onthophagus duboulayi <small>Waterhouse, 1894</small>

Onthophagus ducorpsi <small>Orbigny, 1913</small>

Onthophagus dudleyianus <small>Josso, 2019</small>

Onthophagus dummal <small>Matthews, 1972</small>

Onthophagus dunningi <small>Harold, 1869</small>

Onthophagus durangoensis <small>Balthasar, 1939</small>

Onthophagus duvivieri <small>Orbigny, 1904</small>

Onthophagus ebenicolor <small>Orbigny, 1902</small>

Onthophagus eburneus <small>Orbigny, 1913</small>

Onthophagus echinus <small>Boucomont, 1914</small>

Onthophagus ecopas <small>Josso & Prévost, 2006</small>

Onthophagus egenus <small>Harold, 1877</small>

Onthophagus elegans <small>Klug, 1834</small>

Onthophagus eliptaminus <small>Balthasar, 1969</small>

Onthophagus elongatus <small>Frey, 1954</small>

Onthophagus embersoni <small>Masumoto, Hanboonsong & Ochi, 2002</small>

Onthophagus embrikianus <small>Paulian, 1936</small>

Onthophagus endota <small>Matthews, 1972</small>

Onthophagus endroedianus <small>Walter & Cambefort, 1977</small>

Onthophagus enuguensis <small>Frey, 1975</small>

Onthophagus epilamprus <small>Bates, 1888</small>

Onthophagus erectinasus <small>Orbigny, 1902</small>

Onthophagus erichsoni <small>Hope, 1841</small>

Onthophagus escalerai <small>Orbigny, 1902</small>

Onthophagus eschscholtzi <small>Boucomont, 1924</small>

Onthophagus estelleae <small>Josso, 2022</small>

Onthophagus etlaensis <small>Kohlmann, Escobar-Hernández & Arriaga-Jiménez, 2019</small>

Onthophagus eulophus <small>Bates, 1887</small>

Onthophagus euzeti <small>Walter & Cambefort, 1977</small>

Onthophagus evae <small>Balthasar, 1944</small>

Onthophagus evanidus <small>Harold, 1869</small>

Onthophagus exasperatus <small>Gerstaecker, 1871</small>

Onthophagus excisiceps <small>Orbigny, 1902</small>

Onthophagus exiguus <small>Raffray, 1877</small>

Onthophagus exilis <small>Orbigny, 1913</small>

Onthophagus expansicornis <small>Bates, 1891</small>

Onthophagus exquisitus <small>Arrow, 1931</small>

Onthophagus extensicollis <small>Orbigny, 1907</small>

Onthophagus extraneus <small>Josso, 2021</small>

Onthophagus fabricianus <small>Goidanich, 1926</small>

Onthophagus fabricii <small>Waterhouse, 1894</small>

Onthophagus falcarius <small>Balthasar, 1966</small>

Onthophagus falcifer <small>Harold, 1880</small>

Onthophagus fallaciosus <small>Raffray, 1877</small>

Onthophagus falsus <small>Gillet, 1925</small>

Onthophagus fasciatus <small>Boucomont, 1914</small>

Onthophagus fasciculiger <small>Orbigny, 1902</small>

Onthophagus fasciolatus <small>Boucomont, 1924</small>

Onthophagus favrei <small>Boucomont, 1914</small>

Onthophagus feai <small>Orbigny, 1905</small>

Onthophagus felix <small>Arrow, 1931</small>

Onthophagus ferox <small>Harold, 1867</small> - (Western Dung Beetle)

Onthophagus ferrari <small>Matthews, 1972</small>

Onthophagus filicornis <small>Harold, 1873</small>

Onthophagus fimetarius <small>Roth, 1851</small>

Onthophagus fissiceps <small>MacLeay, 1888</small>

Onthophagus fitiniensis <small>Cambefort, 1984</small>

Onthophagus flavibasis <small>Orbigny, 1904</small>

Onthophagus flavicornis <small>Germar, 1823</small>

Onthophagus flavipennis <small>Orbigny, 1905</small>

Onthophagus flavoapicalis <small>Lea, 1923</small>

Onthophagus flavocinctus <small>Frey, 1975</small>

Onthophagus flavolimbatus <small>Klug, 1855</small>

Onthophagus flavomaculatus <small>Gillet, 1930</small>

Onthophagus flavorufus <small>Orbigny, 1904</small>

Onthophagus fletcheri <small>Blackburn, 1903</small>

Onthophagus flexifrons <small>Orbigny, 1908</small>

Onthophagus foedus <small>Boucomont, 1914</small>

Onthophagus foliaceus <small>Lansberge, 1886</small> - (Scarab beetle)

Onthophagus foliiceps <small>Quedenfeldt, 1884</small>

Onthophagus foraminosus <small>Orbigny, 1902</small>

Onthophagus fossibasis <small>Orbigny, 1910</small>

Onthophagus fossifrons <small>Orbigny, 1902</small>

Onthophagus fossulatus <small>Orbigny, 1908</small>

Onthophagus foulliouxi <small>Cambefort, 1971</small>

Onthophagus foveatus <small>Frey, 1957</small>

Onthophagus foveicollis <small>Boucomont, 1930</small>

Onthophagus fradei <small>Gomes Alves, 1956</small>

Onthophagus fragosus <small>Génier & Howden, 2014</small>

Onthophagus francoisgenieri <small>Krikken & Huijbregts, 2012</small>

Onthophagus francoisi <small>Josso, 2013</small>

Onthophagus frankenbergeri <small>Balthasar, 1933</small>

Onthophagus frankrozendaali <small>Huijbregts & Krikken, 2012</small>

Onthophagus frenchi <small>Blackburn, 1903</small>

Onthophagus fritschi <small>Orbigny, 1902</small>

Onthophagus froggattellus <small>Monteith & Rossini, 2017</small>

Onthophagus fugitivus <small>Péringuey, 1901</small>

Onthophagus fuliginosus <small>Erichson, 1842</small>

Onthophagus fulvocinctus <small>Orbigny, 1902</small>

Onthophagus fulvus <small>Sharp, 1875</small>

Onthophagus funestus <small>Moretto & Génier, 2010</small>

Onthophagus fungicola <small>Orbigny, 1908</small>

Onthophagus fungus <small>Josso, 2022</small>

Onthophagus furcaticeps <small>Masters, 1886</small>

Onthophagus furcicollis <small>Arrow, 1931</small>

Onthophagus furcillifer <small>Bates, 1891</small>

Onthophagus furculifer <small>Orbigny, 1905</small>

Onthophagus furculus <small>(Fabricius, 1798)</small>

Onthophagus fuscatus <small>Orbigny, 1908</small>

Onthophagus fuscidorsis <small>Orbigny, 1902</small>

Onthophagus fuscivestis <small>Orbigny, 1902</small>

Onthophagus fuscopunctatus <small>(Fabricius, 1798)</small>

Onthophagus fuscostriatus <small>Boucomont, 1914</small>

Onthophagus fuscus <small>Boucomont, 1932</small>

Onthophagus gabonensis <small>Walter, 1989</small>

Onthophagus gagatinus <small>Gillet, 1930</small>

Onthophagus gaillardi <small>Orbigny, 1911</small>

Onthophagus gajo <small>Krikken, 1977</small>

Onthophagus ganalensis <small>Gestro, 1895</small>

Onthophagus gandju <small>Matthews, 1972</small>

Onthophagus gangeticus <small>Gillet, 1925</small>

Onthophagus gangulu <small>Matthews, 1972</small>

Onthophagus garambae <small>Frey, 1961</small>

Onthophagus gazellinus <small>Bates, 1887</small>

Onthophagus geelongensis <small>Blackburn, 1892</small>

Onthophagus gemellatus <small>Raffray, 1877</small>

Onthophagus geminatus <small>Orbigny, 1907</small>

Onthophagus geminifrons <small>Orbigny, 1905</small>

Onthophagus gemma <small>(Sharp, 1875)</small>

Onthophagus genuinus <small>Kohlmann & Solis, 2001</small>

Onthophagus georgwerneri <small>Josso, 2022</small>

Onthophagus germanus <small>Gillet, 1927</small>

Onthophagus geryon <small>Balthasar, 1969</small>

Onthophagus gestroi <small>Harold, 1877</small>

Onthophagus ghanensis <small>Balthasar, 1966</small>

Onthophagus gibbidorsis <small>Orbigny, 1902</small>

Onthophagus gibbifrons <small>Orbigny, 1902</small>

Onthophagus gibbus <small>Orbigny, 1913</small>

Onthophagus gibsoni <small>Howden & Génier, 2004</small>

Onthophagus gidju <small>Matthews, 1972</small>

Onthophagus gilli <small>Delgado & Howden, 2000</small>

Onthophagus giraffa <small>(Hausmann, 1807)</small>

Onthophagus girardinae <small>Walter, 1989</small>

Onthophagus giuseppecarpanetoi <small>Tagliaferri & Moretto, 2012</small>

Onthophagus glabratus <small>Hope, 1841</small>

Onthophagus glaucinus <small>Gillet, 1930</small>

Onthophagus gnu <small>Frey, 1955</small>

Onthophagus godarra <small>Storey & Weir, 1990</small>

Onthophagus gonipa <small>Krikken & Huijbregts, 2011</small>

Onthophagus gonopygus <small>Orbigny, 1902</small>

Onthophagus gonzaloi <small>Moctezuma, Hernández & Sánchez-Huerta, 2022</small>

Onthophagus gorochovi <small>Kabakov, 1998</small>

Onthophagus gorokae <small>Paulian, 1972</small>

Onthophagus gothicus <small>Gillet, 1930</small>

Onthophagus grandidorsis <small>Orbigny, 1913</small>

Onthophagus grandifrons <small>Orbigny, 1907</small>

Onthophagus grandivigilans <small>Masumoto, 1995</small>

Onthophagus graniceps <small>Orbigny, 1908</small>

Onthophagus granosus <small>Orbigny, 1913</small>

Onthophagus granulatus <small>Boheman, 1858</small> - (Granulose Dung Beetle)

Onthophagus granulifer <small>Harold, 1886</small>

Onthophagus granulifrons <small>Frey, 1961</small>

Onthophagus granulum <small>Orbigny, 1904</small>

Onthophagus granum <small>Lansberge, 1885</small>

Onthophagus grassei <small>Walter, 1989</small>

Onthophagus grataehelenae <small>Kohlmann & Solis, 2001</small>

Onthophagus gratus <small>Arrow, 1931</small>

Onthophagus gravearmatus <small>Balthasar, 1944</small>

Onthophagus gravis <small>Walker, 1858</small>

Onthophagus gravoti <small>Orbigny, 1908</small>

Onthophagus griseoaeneus <small>Lansberge, 1885</small>

Onthophagus griseosetosus <small>Arrow, 1931</small>

Onthophagus guatemalensis <small>Bates, 1887</small>

Onthophagus gulmarri <small>Matthews, 1972</small>

Onthophagus gurburra <small>Storey & Weir, 1990</small>

Onthophagus guttatus <small>Boheman, 1860</small>

Onthophagus guttiger <small>Orbigny, 1905</small>

Onthophagus haafi <small>Frey, 1963</small>

Onthophagus haagi <small>Harold, 1867</small>

Onthophagus haematopus <small>Harold, 1875</small>

Onthophagus hageni <small>Lansberge, 1883</small>

Onthophagus hagenmontis <small>Krikken & Huijbregts, 2012</small>

Onthophagus hajimei <small>Masumoto, 1984</small>

Onthophagus halffteri <small>Zunino, 1981</small>

Onthophagus hamaticeps <small>Arrow, 1931</small>

Onthophagus hamaticornis <small>Gillet, 1930</small>

Onthophagus hartiniae <small>Ochi & Kon, 2009</small>

Onthophagus hayashii <small>Masumoto, 1991</small>

Onthophagus hecate <small>(Panzer, 1794)</small>

Onthophagus helciatus <small>Harold, 1871</small>

Onthophagus hemichalceus <small>Orbigny, 1915</small>

Onthophagus hemipygus <small>Frey, 1964</small>

Onthophagus hericiniformis <small>Moretto, 2010</small>

Onthophagus hericius <small>Orbigny, 1908</small>

Onthophagus herus <small>Péringuey, 1901</small>

Onthophagus heteroclitus <small>Orbigny, 1911</small>

Onthophagus heurni <small>Gillet, 1930</small>

Onthophagus heyrovskyi <small>Všetečka, 1943</small>

Onthophagus hidalgus <small>Zunino & Halffter, 1988</small>

Onthophagus hielkemai <small>Krikken & Huijbregts, 2011</small>

Onthophagus hilaridis <small>Cambefort, 1984</small>

Onthophagus hilaris <small>Orbigny, 1905</small>

Onthophagus hildebrandti <small>Harold, 1878</small>

Onthophagus hippopotamus <small>Harold, 1869</small>

Onthophagus hircus <small>Billberg, 1815</small>

Onthophagus hirsutulus <small>Lansberge, 1883</small>

Onthophagus hirsutus <small>Orbigny, 1902</small>

Onthophagus hirtellus <small>Frey, 1957</small>

Onthophagus hirticulus <small>Orbigny, 1915</small>

Onthophagus hirtipodex <small>Orbigny, 1904</small>

Onthophagus hirtuosus <small>Gillet, 1930</small>

Onthophagus histeriformis <small>Boucomont, 1914</small>

Onthophagus histrio <small>Orbigny, 1902</small>

Onthophagus hoberlandti <small>Balthasar, 1966</small>

Onthophagus hoepfneri <small>Harold, 1869</small>

Onthophagus hollowayi <small>Krikken & Huijbregts, 2017</small>

Onthophagus holosericus <small>Harold, 1877</small>

Onthophagus holzi <small>Boucomont, 1914</small>

Onthophagus hoogstraali <small>Saylor, 1943</small>

Onthophagus hoplocerus <small>Lea, 1923</small>

Onthophagus hoplothorax <small>Gillet, 1930</small>

Onthophagus horii <small>Ochi, Kon & Tsubaki, 2009</small>

Onthophagus horrens <small>Orbigny, 1908</small>

Onthophagus horribilis <small>Balthasar, 1969</small>

Onthophagus horridus <small>Orbigny, 1908</small>

Onthophagus hosomai <small>Ochi & Kon, 2014</small>

Onthophagus howdeni <small>Zunino & Halffter, 1988</small>

Onthophagus howdenorum <small>Zunino & Halffter, 1988</small>

Onthophagus hulstaerti <small>Boucomont, 1932</small>

Onthophagus humboldti <small>Kohlmann, Solis & Alvarado, 2019</small>

Onthophagus humpatensis <small>Lansberge, 1886</small>

Onthophagus hyaena <small>(Fabricius, 1801)</small>

Onthophagus ibex <small>(Fabricius, 1792)</small>

Onthophagus ieti <small>Cambefort, 1984</small>

Onthophagus ifugaoensis <small>Ochi & Kon, 2006</small>

Onthophagus igualensis <small>Bates, 1887</small>

Onthophagus ikelengeensis <small>Josso, 2018</small>

Onthophagus illotus <small>Péringuey, 1901</small>

Onthophagus illyricus <small>(Scopoli, 1763)</small>

Onthophagus imberbis <small>Orbigny, 1902</small>

Onthophagus imbutus <small>Sharp, 1875</small>

Onthophagus immundus <small>Boheman, 1858</small>

Onthophagus importunus <small>Péringuey, 1901</small>

Onthophagus impressicollis <small>Boheman, 1860</small>

Onthophagus impunctatus <small>Orbigny, 1904</small>

Onthophagus impurus <small>Harold, 1868</small>

Onthophagus inaequalis <small>Orbigny, 1902</small>

Onthophagus incantatus <small>Balthasar, 1967</small>

Onthophagus incanus <small>MacLeay, 1888</small>

Onthophagus incensus <small>Say, 1835</small> - (Scarab beetle)

Onthophagus incisus <small>Harold, 1877</small>

Onthophagus includens <small>Orbigny, 1905</small>

Onthophagus inclusus <small>Orbigny, 1905</small>

Onthophagus incornutus <small>MacLeay, 1871</small>

Onthophagus indigus <small>Péringuey, 1901</small>

Onthophagus indosinicus <small>Boucomont, 1921</small>

Onthophagus indutus <small>Orbigny, 1902</small>

Onthophagus inecolorum <small>Moctezuma & Halffter, 2020</small>

Onthophagus inediapterus <small>Kohlmann & Solis, 2001</small>

Onthophagus ineptus <small>Harold, 1871</small>

Onthophagus inermiceps <small>Orbigny, 1905</small>

Onthophagus inermicollis <small>Orbigny, 1908</small>

Onthophagus infaustus <small>Cambefort, 1984</small>

Onthophagus inflaticollis <small>Bates, 1889</small>

Onthophagus inflatus <small>Walter & Cambefort, 1977</small>

Onthophagus infucatus <small>Harold, 1877</small>

Onthophagus insignis <small>Péringuey, 1896</small>

Onthophagus insularis <small>Boheman, 1858</small>

Onthophagus insulindicus <small>Boucomont, 1914</small>

Onthophagus insulsus <small>Péringuey, 1901</small>

Onthophagus intermixtus <small>Orbigny, 1902</small>

Onthophagus interruptus <small>Raffray, 1877</small>

Onthophagus interstitialis <small>FÃ¥hraeus, 1857</small>

Onthophagus intonsus <small>Orbigny, 1913</small>

Onthophagus intricatus <small>Moretto, 2010</small>

Onthophagus investigator <small>Lansberge, 1885</small>

Onthophagus investis <small>Orbigny, 1904</small>

Onthophagus iodiellus <small>Bates, 1887</small>

Onthophagus ioramaculatus <small>Krikken & Huijbregts, 2012</small>

Onthophagus ioranus <small>Krikken & Huijbregts, 2012</small>

Onthophagus irianus <small>Balthasar, 1969</small>

Onthophagus iris <small>Sharp, 1875</small>

Onthophagus isanus <small>Masumoto, Ochi & Hanboonsong, 2008</small>

Onthophagus ishiii <small>Ochi & Kon, 1995</small>

Onthophagus istmenus <small>Moctezuma, Sánchez-Huerta & Halffter, 2020</small>

Onthophagus itonoborui <small>Masumoto, Ochi & Sakchoowong, 2012</small>

Onthophagus iulicola <small>Cambefort, 1984</small>

Onthophagus iumienus <small>Masumoto, Ochi & Hanboonsong, 2007</small>

Onthophagus ixtepecorum <small>Moctezuma & Halffter, 2020</small>

Onthophagus ixtlanensis <small>Moctezuma & Halffter, 2020</small>

Onthophagus iyengari <small>Arrow, 1931</small>

Onthophagus jacksoni <small>Orbigny, 1913</small>

Onthophagus jacobeus <small>Boucomont, 1924</small>

Onthophagus jalamari <small>Matthews, 1972</small>

Onthophagus jalapensis <small>Balthasar, 1939</small>

Onthophagus jaliscensis <small>Zunino & Halffter, 1988</small>

Onthophagus jangga <small>Matthews, 1972</small>

Onthophagus janssensi <small>Frey, 1958</small>

Onthophagus janthinus <small>Harold, 1875</small>

Onthophagus javaecola <small>Balthasar, 1959</small>

Onthophagus javanensis <small>Balthasar, 1963</small>

Onthophagus javanoides <small>Huijbregts & Krikken, 2011</small>

Onthophagus jocquei <small>Josso, 2019</small>

Onthophagus joliveti <small>Paulian, 1972</small>

Onthophagus jonathani <small>Moretto & Génier, 2020</small>

Onthophagus jubatus <small>Harold, 1869</small>

Onthophagus juheli <small>Josso, 2022</small>

Onthophagus juncticornis <small>Orbigny, 1908</small>

Onthophagus juniperensis <small>Josso, 2019</small>

Onthophagus justei <small>Walter, 1989</small>

Onthophagus jwalae <small>Karimbumkara & Priyadarsanan, 2016</small>

Onthophagus kachinicus <small>Kabakov, 2006</small>

Onthophagus kakadu <small>Storey & Weir, 1990</small>

Onthophagus kanarensis <small>Arrow, 1931</small>

Onthophagus kaomaensis <small>Josso, 2022</small>

Onthophagus kaosoidowensis <small>Masumoto, Ochi & Hanboonsong, 2008</small>

Onthophagus kapuri <small>Endrödi, 1974</small>

Onthophagus karlwerneri <small>Josso, 2022</small>

Onthophagus kashizakii <small>Kon & Ochi, 2005</small>

Onthophagus kashmirensis <small>Balthasar, 1966</small>

Onthophagus kassaicus <small>Orbigny, 1908</small>

Onthophagus katangensis <small>Frey, 1961</small>

Onthophagus kavirondus <small>Orbigny, 1915</small>

Onthophagus kchatriya <small>Boucomont, 1914</small>

Onthophagus keikoae <small>Ochi & Kon, 2014</small>

Onthophagus keiseri <small>Frey, 1956</small>

Onthophagus keralensis <small>Frey, 1975</small>

Onthophagus keralicus <small>Biswas & Chatterjee, 1986</small>

Onthophagus kiambram <small>Storey, 1977</small>

Onthophagus kindianus <small>Frey, 1953</small>

Onthophagus kingstoni <small>Cambefort, 1980</small>

Onthophagus kinhthaicus <small>Kabakov, 1998</small>

Onthophagus kirki <small>Frey, 1975</small>

Onthophagus kirokanus <small>Frey, 1975</small>

Onthophagus kiuchii <small>Masumoto, 1995</small>

Onthophagus kivuensis <small>Walter, 2018</small>

Onthophagus kleinei <small>Balthasar, 1935</small>

Onthophagus knapperti <small>Krikken, 1981</small>

Onthophagus knausi <small>Brown, 1927</small>

Onthophagus koebelei <small>Blackburn, 1903</small>

Onthophagus koechlei <small>Masumoto, Ochi & Kon, 2011</small>

Onthophagus kohlmanni <small>Génier, 2017</small>

Onthophagus kokereka <small>Matthews, 1972</small>

Onthophagus kokocellosus <small>Krikken & Huijbregts, 2013</small>

Onthophagus kokodanus <small>Krikken & Huijbregts, 2012</small>

Onthophagus kokodentatus <small>Krikken & Huijbregts, 2013</small>

Onthophagus kokoiorus <small>Krikken & Huijbregts, 2012</small>

Onthophagus kokopygus <small>Krikken & Huijbregts, 2013</small>

Onthophagus kokosquamatus <small>Krikken & Huijbregts, 2012</small>

Onthophagus kolaka <small>Huijbregts & Krikken, 2009</small>

Onthophagus koma <small>Matsumura, 1937</small>

Onthophagus komareki <small>Balthasar, 1935</small>

Onthophagus kongkaewensis <small>Masumoto, Ochi & Sakchoowong, 2012</small>

Onthophagus konsarnensis <small>Masumoto, Ochi & Hanboonsong, 2008</small>

Onthophagus kontumicus <small>Kabakov, 1983</small>

Onthophagus kora <small>Storey, 1977</small>

Onthophagus kouassii <small>Cambefort, 1984</small>

Onthophagus kraatzeanus <small>Lansberge, 1883</small>

Onthophagus krakadaakhomus <small>Masumoto, 1992</small>

Onthophagus kudrnaianus <small>Josso, 2022</small>

Onthophagus kukali <small>Krikken & Huijbregts, 2013</small>

Onthophagus kulti <small>Balthasar, 1952</small>

Onthophagus kulzeri <small>Frey, 1958</small>

Onthophagus kumaonensis <small>Arrow, 1931</small>

Onthophagus kumbaingeri <small>Matthews, 1972</small>

Onthophagus kyleensis <small>Frey, 1975</small>

Onthophagus labdacus <small>Balthasar, 1969</small>

Onthophagus laborans <small>Arrow, 1931</small>

Onthophagus lacustris <small>Harold, 1877</small>

Onthophagus laetus <small>Orbigny, 1913</small>

Onthophagus laeviceps <small>Orbigny, 1902</small>

Onthophagus laevidorsis <small>Orbigny, 1907</small>

Onthophagus laevigatus <small>(Fabricius, 1798)</small>

Onthophagus laevissimus <small>Orbigny, 1905</small>

Onthophagus lagnyi <small>Paulian, 1937</small>

Onthophagus lamellicornis <small>Frey, 1953</small>

Onthophagus lamgalio <small>Matthews, 1972</small>

Onthophagus laminatus <small>MacLeay, 1864</small>

Onthophagus laminicornis <small>Lansberge, 1886</small>

Onthophagus laminidorsis <small>Orbigny, 1902</small>

Onthophagus laminosus <small>Orbigny, 1905</small>

Onthophagus lamnifer <small>Orbigny, 1902</small>

Onthophagus lamotellus <small>Cambefort, 1980</small>

Onthophagus lamottei <small>Frey, 1962</small>

Onthophagus lamtoi <small>Cambefort, 1980</small>

Onthophagus landolti <small>Harold, 1880</small>

Onthophagus langkawiensis <small>Ochi & Kon, 2015</small>

Onthophagus lapillus <small>Arrow, 1931</small>

Onthophagus laratinus <small>Arrow, 1916</small>

Onthophagus lassulus <small>Balthasar, 1964</small>

Onthophagus latenasutus <small>Arrow, 1941</small>

Onthophagus latepunctatus <small>Orbigny, 1913</small>

Onthophagus latevittatus <small>Orbigny, 1905</small>

Onthophagus latigibber <small>Orbigny, 1902</small>

Onthophagus latipennis <small>Orbigny, 1897</small>

Onthophagus latissimus <small>Orbigny, 1902</small>

Onthophagus latro <small>Harold, 1877</small>

Onthophagus leai <small>Blackburn, 1895</small>

Onthophagus leanus <small>Goidanich, 1926</small>

Onthophagus lebasi <small>Boucomont, 1932</small>

Onthophagus lecontei <small>Harold, 1871</small>

Onthophagus lefebvrei <small>Orbigny, 1913</small>

Onthophagus lefiniensis <small>Balthasar, 1967</small>

Onthophagus legendrei <small>Walter & Cambefort, 1977</small>

Onthophagus leichhardti <small>Monteith & Storey, 2013</small>

Onthophagus lemagneni <small>Orbigny, 1902</small>

Onthophagus lemekensis <small>Orbigny, 1915</small>

Onthophagus lemniscatus <small>Gillet, 1924</small>

Onthophagus leomontanus <small>Boucomont, 1914</small>

Onthophagus leroyi <small>Orbigny, 1902</small>

Onthophagus leucopygus <small>Harold, 1867</small>

Onthophagus leusermontis <small>Huijbregts & Krikken, 2011</small>

Onthophagus limonensis <small>Kohlmann & Solis, 2001</small>

Onthophagus lindaae <small>Masumoto, 1989</small>

Onthophagus lindu <small>Huijbregts & Krikken, 2009</small>

Onthophagus liodermus <small>Orbigny, 1913</small>

Onthophagus lioides <small>Orbigny, 1902</small>

Onthophagus liopterus <small>Harold, 1880</small>

Onthophagus liothorax <small>Koshantschikov, 1894</small>

Onthophagus lituratus <small>Roth, 1851</small>

Onthophagus lobi <small>Cambefort, 1984</small>

Onthophagus lojanus <small>Balthasar, 1939</small>

Onthophagus lomii <small>Müller, 1942</small>

Onthophagus longecarinatus <small>Zunino & Halffter, 1988</small>

Onthophagus longegranus <small>Frey, 1957</small>

Onthophagus longiceps <small>Orbigny, 1904</small>

Onthophagus longimanus <small>Bates, 1887</small>

Onthophagus longipes <small>Paulian, 1937</small>

Onthophagus longipilis <small>Orbigny, 1905</small>

Onthophagus lopeensis <small>Josso, 2022</small>

Onthophagus lore <small>Huijbregts & Krikken, 2009</small>

Onthophagus lorianus <small>Gillet, 1930</small>

Onthophagus loroi <small>Balthasar, 1941</small>

Onthophagus loudetiae <small>Cambefort, 1984</small>

Onthophagus loxodontae <small>Cambefort, 1984</small>

Onthophagus luctuosus <small>Boucomont, 1914</small>

Onthophagus ludicrus <small>Balthasar, 1969</small>

Onthophagus ludio <small>Boucomont, 1914</small>

Onthophagus lugens <small>FÃ¥hraeus, 1857</small>

Onthophagus lugubris <small>FÃ¥hraeus, 1857</small>

Onthophagus luismargaritorum <small>Delgado, 1995</small>

Onthophagus lumareti <small>Walter & Cambefort, 1977</small>

Onthophagus lunulifer <small>Boucomont, 1914</small>

Onthophagus lutaticollis <small>Orbigny, 1907</small>

Onthophagus luteosignatus <small>Lansberge, 1883</small>

Onthophagus luzonwallacei <small>Ochi, Kawahara & Cabras, 2021</small>

Onthophagus mabuensis <small>Daniel, Strümpher & Josso, 2023</small>

Onthophagus macleayi <small>Blackburn, 1903</small>

Onthophagus macrocephalus <small>Kirby, 1818</small>

Onthophagus macrothorax <small>Orbigny, 1902</small>

Onthophagus maculosipennis <small>Gillet, 1930</small>

Onthophagus maculosus <small>Orbigny, 1908</small>

Onthophagus madoqua <small>Arrow, 1931</small>

Onthophagus magnioculus <small>Ochi & Kon, 2006</small>

Onthophagus magnipygus <small>Boucomont, 1914</small>

Onthophagus makokou <small>Walter, 1989</small>

Onthophagus malabarensis <small>Boucomont, 1919</small>

Onthophagus malangensis <small>Boucomont, 1914</small>

Onthophagus malasiacus <small>Gillet, 1927</small>

Onthophagus maleengnaafon <small>Masumoto, 1990</small>

Onthophagus maleengnoi <small>Masumoto, 1990</small>

Onthophagus malevolus <small>Balthasar, 1964</small>

Onthophagus malthinus <small>Gillet, 1930</small>

Onthophagus mamillatus <small>Lea, 1923</small>

Onthophagus manantlanensis <small>Sánchez-Huerta, Moctezuma & Hernández, 2021</small>

Onthophagus manguliensis <small>Boucomont, 1914</small>

Onthophagus maniti <small>Masumoto, 1989</small>

Onthophagus mankonoensis <small>Balthasar, 1966</small>

Onthophagus manya <small>Matthews, 1972</small>

Onthophagus marahouensis <small>Cambefort, 1984</small>

Onthophagus margaretensis <small>Blackburn, 1903</small>

Onthophagus margaritifer <small>Orbigny, 1898</small>

Onthophagus marginatus <small>Castelnau, 1840</small>

Onthophagus marginicollis <small>Harold, 1880</small>

Onthophagus marginifer <small>Frey, 1953</small>

Onthophagus marginipennis <small>Josso, 2018</small>

Onthophagus mariozuninoi <small>Delgado, Navarrete & Blackaller-Bages, 1993</small>

Onthophagus marshalli <small>Orbigny, 1908</small>

Onthophagus martellii <small>Frey, 1972</small>

Onthophagus martialis <small>Boucomont, 1914</small>

Onthophagus martinpierai <small>Moctezuma, Rossini & Zunino, 2016</small>

Onthophagus masaicus <small>Orbigny, 1905</small>

Onthophagus masumotoi <small>Ochi, 1985</small>

Onthophagus matae <small>Cambefort, 1984</small>

Onthophagus matanyo <small>Huijbregts & Krikken, 2009</small>

Onthophagus mateui <small>Moctezuma & Halffter, 2020</small>

Onthophagus matsudai <small>Ochi & Kon, 2004</small>

Onthophagus matsuii <small>Ochi & Kon, 2006</small>

Onthophagus mauritii <small>Boucomont, 1919</small>

Onthophagus maxwellianus <small>Moretto, 2013</small>

Onthophagus maya <small>Zunino, 1981</small>

Onthophagus mayeri <small>Harold, 1876</small>

Onthophagus mcclevei <small>Howden & Génier, 2004</small>

Onthophagus medorensis <small>Brown, 1929</small>

Onthophagus megapacificus <small>Ochi & Kon, 2006</small>

Onthophagus megathorax <small>Gillet, 1921</small>

Onthophagus mekara <small>Huijbregts & Krikken, 2009</small>

Onthophagus mendicus <small>Gillet, 1924</small>

Onthophagus menkaoensis <small>Walter & Cambefort, 1977</small>

Onthophagus merdrignaci <small>Walter & Cambefort, 1977</small>

Onthophagus meruanus <small>Orbigny, 1910</small>

Onthophagus merus <small>Péringuey, 1901</small>

Onthophagus metalliger <small>Orbigny, 1913</small>

Onthophagus mexicanus <small>Bates, 1887</small>

Onthophagus mextexus <small>Howden & Cartwright, 1970</small>

Onthophagus micropterus <small>Zunino & Halffter, 1981</small>

Onthophagus micros <small>Orbigny, 1902</small>

Onthophagus mije <small>Storey & Weir, 1990</small>

Onthophagus miles <small>Orbigny, 1905</small>

Onthophagus militaris <small>Boucomont, 1914</small>

Onthophagus millamilla <small>Matthews, 1972</small>

Onthophagus mimikanus <small>Balthasar, 1969</small>

Onthophagus mimus <small>Orbigny, 1913</small>

Onthophagus minettianus <small>Josso, 2019</small>

Onthophagus minotaurus <small>Arrow, 1941</small>

Onthophagus minutissimus <small>Orbigny, 1908</small>

Onthophagus minutulus <small>Harold, 1875</small>

Onthophagus minutus <small>(Hausmann, 1807)</small>

Onthophagus mirabilis <small>Bates, 1887</small>

Onthophagus mirandus <small>Arrow, 1931</small>

Onthophagus miricollis <small>Frey, 1962</small>

Onthophagus miricornis <small>Orbigny, 1902</small>

Onthophagus mirifrons <small>Orbigny, 1905</small>

Onthophagus miscellaneus <small>Orbigny, 1908</small>

Onthophagus miscellus <small>Orbigny, 1905</small>

Onthophagus misellus <small>Orbigny, 1907</small>

Onthophagus mixticeps <small>Orbigny, 1905</small>

Onthophagus mixtidorsis <small>Orbigny, 1905</small>

Onthophagus mjobergi <small>Gillet, 1925</small>

Onthophagus mniszechi <small>Harold, 1869</small> - (Threehorned Dung Beetle)

Onthophagus moajat <small>Huijbregts & Krikken, 2009</small>

Onthophagus mocquerysi <small>Orbigny, 1902</small>

Onthophagus modestus <small>Harold, 1862</small>

Onthophagus mokwamensis <small>Krikken & Huijbregts, 2012</small>

Onthophagus monardi <small>Boucomont, 1936</small>

Onthophagus monardiellus <small>Frey, 1963</small>

Onthophagus mongana <small>Storey & Weir, 1990</small>

Onthophagus monodon <small>FÃ¥hraeus, 1857</small>

Onthophagus monteithi <small>Matthews, 1972</small>

Onthophagus montishannoniae <small>Krikken & Huijbregts, 2008</small>

Onthophagus montivagus <small>Orbigny, 1902</small>

Onthophagus montreuili <small>Moretto & Génier, 2010</small>

Onthophagus mordindangus <small>Masumoto, Ochi & Hanboonsong, 2008</small>

Onthophagus moreleti <small>Baraud, 1980</small>

Onthophagus morenoi <small>Moretto, 2013</small>

Onthophagus morettoi <small>Josso, 2022</small>

Onthophagus moroni <small>Zunino & Halffter, 1988</small>

Onthophagus morosus <small>Gerstaecker, 1871</small>

Onthophagus mpassa <small>Walter, 1982</small>

Onthophagus mucronatus <small>Thomson, 1858</small>

Onthophagus mucronifer <small>Orbigny, 1905</small>

Onthophagus mugheseensis <small>Josso, 2019</small>

Onthophagus mulgravei <small>Paulian, 1937</small>

Onthophagus multipunctatus <small>Boucomont, 1914</small>

Onthophagus mundill <small>Matthews, 1972</small>

Onthophagus murchisoni <small>Blackburn, 1892</small>

Onthophagus murgon <small>Monteith & Storey, 2013</small>

Onthophagus murphyi <small>Josso, 2019</small>

Onthophagus musculus <small>Frey, 1963</small>

Onthophagus mutatus <small>Harold, 1859</small>

Onthophagus muticus <small>MacLeay, 1864</small>

Onthophagus mycetorum <small>Zunino & Halffter, 1988</small>

Onthophagus naaroon <small>Masumoto, 1990</small>

Onthophagus naevuliger <small>Orbigny, 1908</small>

Onthophagus nagpurensis <small>Arrow, 1931</small>

Onthophagus nammuldi <small>Matthews, 1972</small>

Onthophagus namnaoensis <small>Masumoto, Ochi & Hanboonsong, 2002</small>

Onthophagus namnaolekus <small>Masumoto, Ochi & Higurashi, 2020</small>

Onthophagus namnaous <small>Masumoto, Ochi & Hanboonsong, 2013</small>

Onthophagus nanus <small>Harold, 1878</small>

Onthophagus nasicornis <small>Harold, 1869</small>

Onthophagus nasicus <small>Gillet, 1930</small>

Onthophagus naso <small>FÃ¥hraeus, 1857</small>

Onthophagus nasonis <small>Kabakov, 1998</small>

Onthophagus nasutus <small>Guérin-Méneville, 1855</small>

Onthophagus natronis <small>Cambefort, 1984</small>

Onthophagus navarretorum <small>Delgado & Capistan, 1996</small>

Onthophagus nchilaensis <small>Josso, 2022</small>

Onthophagus neboissi <small>Frey, 1970</small>

Onthophagus necrophagus <small>Arrow, 1931</small>

Onthophagus neervoorti <small>Boucomont, 1914</small>

Onthophagus negligens <small>Walker, 1858</small>

Onthophagus nemorivagus <small>Kohlmann & Solis, 2001</small>

Onthophagus neofurcatus <small>Goidanich, 1926</small>

Onthophagus neofuscus <small>Moctezuma & Halffter, 2020</small>

Onthophagus neomirabilis <small>Howden, 1973</small>

Onthophagus neostenocerus <small>Goidanich, 1926</small>

Onthophagus neyo <small>Moretto, 2013</small>

Onthophagus nicobaricus <small>Biswas, Chatterjee & Sengupta, 1999</small>

Onthophagus nigerianus <small>Orbigny, 1913</small>

Onthophagus nigriceps <small>Raffray, 1877</small>

Onthophagus nigrinus <small>Paulian, 1937</small>

Onthophagus nigripennis <small>Orbigny, 1908</small>

Onthophagus nigriventris <small>Orbigny, 1902</small> - (Coastal Dung Beetle)

Onthophagus nigrivestis <small>Orbigny, 1902</small>

Onthophagus nigroflavus <small>Josso, 2022</small>

Onthophagus nilicola <small>Orbigny, 1902</small>

Onthophagus niloticus <small>Harold, 1879</small>

Onthophagus nimbatus <small>Orbigny, 1905</small>

Onthophagus niokolokoba <small>Moretto, 2007</small>

Onthophagus nitefactus <small>Harold, 1877</small>

Onthophagus nitidifrons <small>Orbigny, 1905</small>

Onthophagus nitidior <small>Bates, 1887</small>

Onthophagus nodieri <small>Orbigny, 1908</small>

Onthophagus nodulifer <small>Harold, 1867</small>

Onthophagus nonstriatus <small>Orbigny, 1905</small>

Onthophagus notatus <small>Orbigny, 1902</small>

Onthophagus notiodes <small>Solis & Kohlmann, 2003</small>

Onthophagus novaeirlandiae <small>Balthasar, 1969</small>

Onthophagus nubilus <small>Kohlmann & Solis, 2001</small>

Onthophagus nudatus <small>Orbigny, 1913</small>

Onthophagus nudifrons <small>Balthasar, 1939</small>

Onthophagus nudus <small>Orbigny, 1908</small>

Onthophagus nummelini <small>Moretto & Génier, 2020</small>

Onthophagus nurubuan <small>Matthews, 1972</small>

Onthophagus nyctopus <small>Bates, 1887</small>

Onthophagus nyikaensis <small>Josso, 2018</small>

Onthophagus nymani <small>Gillet, 1930</small>

Onthophagus oaxacanus <small>Zunino & Halffter, 1988</small>

Onthophagus obliviosus <small>Balthasar, 1935</small>

Onthophagus obtusicornis <small>FÃ¥hraeus, 1857</small>

Onthophagus obtutus <small>Péringuey, 1901</small>

Onthophagus ocellidorsis <small>Orbigny, 1915</small>

Onthophagus ocellifer <small>Orbigny, 1902</small>

Onthophagus ocelliger <small>Harold, 1877</small>

Onthophagus ochii <small>Masumoto, 1988</small>

Onthophagus ochromerus <small>Harold, 1877</small>

Onthophagus ochropygus <small>Orbigny, 1902</small>

Onthophagus octogonus <small>Frey, 1962</small>

Onthophagus ofianus <small>Krikken & Huijbregts, 2013</small>

Onthophagus ohkuboi <small>Ochi & Kon, 2006</small>

Onthophagus okahandjanus <small>Balthasar, 1974</small>

Onthophagus oklahomensis <small>Brown, 1927</small> - (Scarab beetle)

Onthophagus olidus <small>Balthasar, 1964</small>

Onthophagus omiltemius <small>Bates, 1889</small>

Onthophagus omostigma <small>Orbigny, 1902</small>

Onthophagus onorei <small>Zunino & Halffter, 1997</small>

Onthophagus onthochromus <small>Arrow, 1913</small>

Onthophagus ontosatu <small>Huijbregts & Krikken, 2012</small>

Onthophagus opacihartiniae <small>Ochi & Kon, 2015</small>

Onthophagus opacotaurus <small>Krikken & Huijbregts, 2012</small>

Onthophagus ophion <small>Erichson, 1847</small>

Onthophagus ophtalmicus <small>Frey, 1958</small>

Onthophagus orbicularis <small>Lansberge, 1885</small>

Onthophagus orbus <small>Boucomont, 1924</small>

Onthophagus orientalis <small>Harold, 1868</small>

Onthophagus orissanus <small>Arrow, 1931</small>

Onthophagus orizabensis <small>Moctezuma, Joaqui & Sánchez-Huerta, 2019</small>

Onthophagus ornaticollis <small>Gillet, 1930</small>

Onthophagus ornatulus <small>Orbigny, 1908</small>

Onthophagus orpheus <small>(Panzer, 1794)</small>

Onthophagus orphnoides <small>Bates, 1887</small>

Onthophagus orthocerus <small>Thomson, 1858</small>

Onthophagus osculatii <small>Guérin-Méneville, 1855</small>

Onthophagus otai <small>Ochi & Kon, 2006</small>

Onthophagus otjivarongus <small>Balthasar, 1967</small>

Onthophagus ouratita <small>Matthews, 1972</small>

Onthophagus ovigranosus <small>Frey, 1971</small>

Onthophagus ovulum <small>Gerstaecker, 1871</small>

Onthophagus pacificus <small>Lansberge, 1885</small>

Onthophagus padrinoi <small>Delgado, 1999</small>

Onthophagus palamoui <small>Biswas & Chatterjee, 1986</small>

Onthophagus palatus <small>Boucomont, 1914</small>

Onthophagus palavanus <small>Balthasar, 1959</small>

Onthophagus pallidipennis <small>FÃ¥hraeus, 1857</small>

Onthophagus pallidus <small>Orbigny, 1905</small>

Onthophagus paluma <small>Matthews, 1972</small>

Onthophagus pangmaphanus <small>Masumoto, Ochi & Higurashi, 2022</small>

Onthophagus papuater <small>Krikken & Huijbregts, 2012</small>

Onthophagus papuensis <small>Harold, 1877</small>

Onthophagus papuplicatus <small>Krikken & Huijbregts, 2012</small>

Onthophagus papurugosus <small>Krikken & Huijbregts, 2012</small>

Onthophagus paracentricornis <small>Ochi & Kon, 2014</small>

Onthophagus parafasciatus <small>Balthasar, 1974</small>

Onthophagus parafuscus <small>Zunino & Halffter, 2005</small>

Onthophagus parallelicornis <small>MacLeay, 1887</small>

Onthophagus parapalatus <small>Krikken & Huijbregts, 1988</small>

Onthophagus parapedisequus <small>Balthasar, 1969</small>

Onthophagus paravinctus <small>Tagliaferri & Moretto, 2012</small>

Onthophagus parceguttatus <small>Orbigny, 1902</small>

Onthophagus parcepictus <small>Orbigny, 1908</small>

Onthophagus parcepilosus <small>Balthasar, 1966</small>

Onthophagus parenthesis <small>Boucomont, 1913</small>

Onthophagus parisii <small>Balthasar, 1941</small>

Onthophagus parrumbal <small>Matthews, 1972</small>

Onthophagus parryi <small>Harold, 1869</small>

Onthophagus parumnotatus <small>FÃ¥hraeus, 1857</small>

Onthophagus parvus <small>Blanchard, 1853</small>

Onthophagus paucigranosus <small>Lansberge, 1886</small>

Onthophagus paulianellus <small>Frey, 1958</small>

Onthophagus pauliani <small>Balthasar, 1937</small>

Onthophagus pauper <small>Boucomont, 1914</small>

Onthophagus pauxillus <small>Orbigny, 1902</small>

Onthophagus pavidus <small>Harold, 1877</small>

Onthophagus pedator <small>Sharp, 1875</small>

Onthophagus pedester <small>Howden & Génier, 2004</small>

Onthophagus pedisequus <small>Balthasar, 1969</small>

Onthophagus penedwardsae <small>Monteith & Storey, 2013</small>

Onthophagus peninsulomerus <small>Huijbregts & Krikken, 2011</small>

Onthophagus pennsylvanicus <small>Harold, 1871</small>

Onthophagus pentacanthus <small>Harold, 1867</small> - (Fivehorned Dung Beetle)

Onthophagus peotoxus <small>Krikken & Huijbregts, 2012</small>

Onthophagus peramelinus <small>(Lea, 1923)</small>

Onthophagus peringueyi <small>Shipp, 1895</small>

Onthophagus perniger <small>Boucomont, 1930</small>

Onthophagus perpilosus <small>MacLeay, 1871</small>

Onthophagus personatus <small>Boucomont, 1914</small>

Onthophagus petenensis <small>Howden & Gill, 1993</small>

Onthophagus pexatus <small>Harold, 1869</small>

Onthophagus peyrierasi <small>Paulian, 1985</small>

Onthophagus phanaeomorphus <small>Janssens, 1954</small>

Onthophagus phatoensis <small>Masumoto, Ochi & Hanboonsong, 2008</small>

Onthophagus philippemorettoi <small>Josso, 2022</small>

Onthophagus philippinensis <small>Boucomont, 1919</small>

Onthophagus phoenicocerus <small>Lea, 1923</small>

Onthophagus phrixus <small>Balthasar, 1969</small>

Onthophagus phrutsaphaakhomus <small>Masumoto, 1992</small>

Onthophagus phukhieoensis <small>Masumoto, Ochi & Hanboonsong, 2007</small>

Onthophagus picatus <small>Orbigny, 1902</small>

Onthophagus piceiceps <small>Orbigny, 1904</small>

Onthophagus picipennis <small>Hope, 1841</small>

Onthophagus pictipennis <small>Orbigny, 1913</small>

Onthophagus pictipodex <small>Orbigny, 1902</small>

Onthophagus picturatus <small>Orbigny, 1908</small>

Onthophagus pictus <small>Reitter, 1892</small>

Onthophagus pierrei <small>Josso, 2022</small>

Onthophagus pilicollis <small>Orbigny, 1902</small>

Onthophagus pilipodex <small>Orbigny, 1913</small>

Onthophagus pillara <small>Matthews, 1972</small>

Onthophagus pilosus <small>FÃ¥hraeus, 1857</small>

Onthophagus pimpasaleei <small>Masumoto, Ochi & Hanboonsong, 2008</small>

Onthophagus pinaroo <small>Storey & Weir, 1990</small>

Onthophagus pinguis <small>Gerstaecker, 1871</small>

Onthophagus pipitzi <small>Ancey, 1883</small>

Onthophagus pisciphagus <small>Walter & Cambefort, 1977</small>

Onthophagus pithankithae <small>Karimbumkara & Priyadarsanan, 2016</small>

Onthophagus placens <small>Péringuey, 1904</small>

Onthophagus planaticeps <small>Orbigny, 1913</small>

Onthophagus planiceps <small>MacLeay, 1886</small>

Onthophagus planicollis <small>Harold, 1880</small>

Onthophagus planifrons <small>Frey, 1962</small>

Onthophagus platalea <small>Arrow, 1941</small>

Onthophagus pleurogonus <small>Orbigny, 1913</small>

Onthophagus plicatifrons <small>Orbigny, 1908</small>

Onthophagus politus <small>(Fabricius, 1798)</small>

Onthophagus pollicatus <small>Harold, 1879</small>

Onthophagus polyedrus <small>Orbigny, 1905</small>

Onthophagus polyodon <small>Orbigny, 1913</small>

Onthophagus polyphemi <small>Hubbard, 1894</small> - (Onthophagus tortoise commensal scarab)

Onthophagus polystigma <small>Orbigny, 1902</small>

Onthophagus pooensis <small>Cambefort & Nicolas, 1991</small>

Onthophagus porcus <small>Arrow, 1931</small>

Onthophagus possoi <small>Walter, 1982</small>

Onthophagus posticicornis <small>Orbigny, 1913</small>

Onthophagus posticus <small>Erichson, 1842</small>

Onthophagus potosinus <small>Moctezuma & Halffter, 2020</small>

Onthophagus praecellens <small>Bates, 1887</small>

Onthophagus praelaminatus <small>Frey, 1957</small>

Onthophagus praestans <small>Péringuey, 1901</small>

Onthophagus prehensilis <small>(Arrow, 1920)</small>

Onthophagus probus <small>Péringuey, 1901</small>

Onthophagus profanus <small>Frey, 1958</small>

Onthophagus pronocarinatus <small>Josso, 2019</small>

Onthophagus pronus <small>Erichson, 1842</small>

Onthophagus propinquus <small>MacLeay, 1888</small>

Onthophagus propraecellens <small>Howden & Gill, 1987</small>

Onthophagus proteus <small>Orbigny, 1902</small>

Onthophagus proximus <small>Orbigny, 1908</small>

Onthophagus pseudoaeneus <small>Orbigny, 1908</small>

Onthophagus pseudobidens <small>Orbigny, 1915</small>

Onthophagus pseudobrutus <small>Kabakov, 1983</small>

Onthophagus pseudobufonidius <small>Josso, 2019</small>

Onthophagus pseudoconvexicollis <small>Bai & Yang, 2016</small>

Onthophagus pseudocoracinus <small>Kabakov, 1983</small>

Onthophagus pseudocostifer <small>Krikken & Huijbregts, 2012</small>

Onthophagus pseudocribripennis <small>Josso, 2019</small>

Onthophagus pseudofimetarius <small>Balthasar, 1946</small>

Onthophagus pseudofuscus <small>Zunino & Halffter, 1988</small>

Onthophagus pseudoguatemalensis <small>Moctezuma & Halffter, 2021</small>

Onthophagus pseudohystrix <small>Masumoto, 1995</small>

Onthophagus pseudopilosus <small>Frey, 1958</small>

Onthophagus pseudosanguineus <small>Walter & Cambefort, 1977</small>

Onthophagus pseudosellatus <small>Balthasar, 1964</small>

Onthophagus pseudoundulans <small>Zunino & Halffter, 1988</small>

Onthophagus pseudovinctus <small>Tagliaferri & Moretto, 2012</small>

Onthophagus ptox <small>Erichson, 1847</small>

Onthophagus puberulus <small>Orbigny, 1902</small>

Onthophagus pugionatus <small>FÃ¥hraeus, 1857</small>

Onthophagus pugnacior <small>Blackburn, 1903</small>

Onthophagus pugnax <small>Harold, 1868</small>

Onthophagus pulchellus <small>Orbigny, 1905</small>

Onthophagus pullatus <small>Orbigny, 1905</small>

Onthophagus pullus <small>Roth, 1851</small>

Onthophagus punthari <small>Storey, 1977</small>

Onthophagus pupillatus <small>Kolbe, 1886</small>

Onthophagus purifrons <small>Orbigny, 1908</small>

Onthophagus purpurascens <small>Boucomont, 1914</small>

Onthophagus purpureicollis <small>MacLeay, 1864</small>

Onthophagus pusillus <small>(Fabricius, 1798)</small>

Onthophagus pusio <small>FÃ¥hraeus, 1857</small>

Onthophagus pygidialis <small>Lansberge, 1883</small>

Onthophagus pygmaeus <small>(Schaller, 1783)</small>

Onthophagus quadricallosus <small>Orbigny, 1902</small>

Onthophagus quadrilunatus <small>Orbigny, 1902</small>

Onthophagus quadrimaculatus <small>Raffray, 1877</small>

Onthophagus quadrinodosus <small>FÃ¥hraeus, 1857</small>

Onthophagus quadrinotatus <small>Orbigny, 1905</small>

Onthophagus quadripustulatus <small>(Fabricius, 1775)</small>

Onthophagus queenslandicus <small>Blackburn, 1903</small>

Onthophagus quetzalis <small>Howden & Gill, 1993</small>

Onthophagus quinquetuberculatus <small>MacLeay, 1871</small>

Onthophagus quiproquo <small>Moretto & Génier, 2010</small>

Onthophagus ragazzii <small>Orbigny, 1904</small>

Onthophagus rana <small>Arrow, 1931</small>

Onthophagus ranunculus <small>Arrow, 1913</small>

Onthophagus rasipennis <small>Orbigny, 1908</small>

Onthophagus ratchasimaensis <small>Masumoto, Hanboonsong & Ochi, 2002</small>

Onthophagus ratchataniensis <small>Masumoto, Ochi & Hanboonsong, 2013</small>

Onthophagus rectestriatus <small>Orbigny, 1915</small>

Onthophagus rectilamina <small>Orbigny, 1902</small>

Onthophagus rectorispauliani <small>Cambefort, 1980</small>

Onthophagus refulgens <small>Arrow, 1931</small>

Onthophagus regalis <small>Arrow, 1907</small>

Onthophagus reticollis <small>MacLeay, 1886</small>

Onthophagus reticuliger <small>Frey, 1960</small>

Onthophagus reticulundaticeps <small>Josso, 2022</small>

Onthophagus retusus <small>Harold, 1869</small>

Onthophagus reyesi <small>Zunino & Halffter, 1988</small>

Onthophagus rhinocerus <small>Gomes Alves, 1944</small>

Onthophagus rhinolophus <small>Harold, 1869</small>

Onthophagus rhinophyllus <small>Harold, 1868</small>

Onthophagus rhodesianus <small>Frey, 1975</small>

Onthophagus rhynchophorus <small>Péringuey, 1904</small>

Onthophagus riparius <small>Lansberge, 1885</small>

Onthophagus rizali <small>Ochi, Kawahara & Cabras, 2021</small>

Onthophagus robertminettii <small>Josso, 2022</small>

Onthophagus robertopoggii <small>Ochi & Kon, 2006</small>

Onthophagus rohwedderi <small>Moretto, 2017</small>

Onthophagus rojkoffianus <small>Josso, 2018</small>

Onthophagus rorarius <small>Harold, 1877</small>

Onthophagus rosemarieae <small>Josso, 2022</small>

Onthophagus rosenbergi <small>Krikken & Huijbregts, 2011</small>

Onthophagus rosettae <small>Frey, 1958</small>

Onthophagus rostratus <small>Harold, 1869</small>

Onthophagus rotundatus <small>Orbigny, 1905</small>

Onthophagus roubali <small>Balthasar, 1935</small>

Onthophagus rouyeri <small>Boucomont, 1914</small>

Onthophagus royi <small>Biswas & Chatterjee, 1985</small>

Onthophagus ruandanus <small>Frey, 1971</small>

Onthophagus rubefactus <small>Orbigny, 1913</small>

Onthophagus rubellus <small>Orbigny, 1908</small>

Onthophagus rubens <small>Orbigny, 1902</small>

Onthophagus rubenticollis <small>Orbigny, 1902</small>

Onthophagus rubescens <small>MacLeay, 1888</small>

Onthophagus rubicundulus <small>MacLeay, 1871</small>

Onthophagus rubidus <small>Orbigny, 1913</small>

Onthophagus rubrescens <small>Blanchard, 1845</small>

Onthophagus rubricatus <small>Orbigny, 1902</small>

Onthophagus rubrimaculatus <small>MacLeay, 1864</small>

Onthophagus rubripennis <small>Arrow, 1907</small>

Onthophagus rufescens <small>Bates, 1887</small>

Onthophagus ruficauda <small>Arrow, 1931</small>

Onthophagus rufipodex <small>Orbigny, 1913</small>

Onthophagus rufocastaneus <small>Fairmaire, 1887</small>

Onthophagus rufolimbatus <small>Orbigny, 1913</small>

Onthophagus rufonotatus <small>Orbigny, 1902</small>

Onthophagus rufosignatus <small>MacLeay, 1864</small>

Onthophagus rufovirens <small>Orbigny, 1904</small>

Onthophagus rugicollis <small>Harold, 1880</small>

Onthophagus rugidorsis <small>Orbigny, 1913</small>

Onthophagus rugosicollis <small>Gillet, 1925</small>

Onthophagus rugosipennis <small>Frey, 1973</small>

Onthophagus rugosissimus <small>Orbigny, 1913</small>

Onthophagus rupicapra <small>Waterhouse, 1894</small>

Onthophagus rutilans <small>Sharp, 1875</small>

Onthophagus rutriceps <small>Krikken & Huijbregts, 2012</small>

Onthophagus sabahensis <small>Ochi & Kon, 2006</small>

Onthophagus sahai <small>Biswas & Chatterjee, 1986</small>

Onthophagus saigonensis <small>Boucomont, 1924</small>

Onthophagus sakaeratensis <small>Masumoto, Hanboonsong & Ochi, 2002</small>

Onthophagus sakainoi <small>Masumoto, 1991</small>

Onthophagus salebrosus <small>MacLeay, 1888</small>

Onthophagus saleyeri <small>Lansberge, 1883</small>

Onthophagus salvadorensis <small>Zunino & Halffter, 1988</small>

Onthophagus salvazai <small>Paulian, 1978</small>

Onthophagus samoengus <small>Masumoto, Ochi & Hanboonsong, 2008</small>

Onthophagus sancristobalensis <small>Moctezuma & Halffter, 2020</small>

Onthophagus sanggona <small>Huijbregts & Krikken, 2009</small>

Onthophagus sangirensis <small>Boucomont, 1914</small>

Onthophagus sanguineus <small>Orbigny, 1902</small>

Onthophagus sanguinolentus <small>Orbigny, 1908</small>

Onthophagus sangwalus <small>Masumoto, Ochi & Hanboonsong, 2008</small>

Onthophagus sanpabloetlorum <small>Moctezuma & Halffter, 2020</small>

Onthophagus sansibaricus <small>Harold, 1878</small>

Onthophagus santamariensis <small>Moctezuma, Sánchez-Huerta & Halffter, 2020</small>

Onthophagus sarasinorum <small>Krikken & Huijbregts, 2011</small>

Onthophagus savanicola <small>Cambefort, 1984</small>

Onthophagus scaber <small>Roth, 1851</small>

Onthophagus scapularis <small>Orbigny, 1902</small>

Onthophagus sceptrifer <small>Boucomont, 1924</small>

Onthophagus schaefferi <small>Howden & Cartwright, 1963</small>

Onthophagus schawalleri <small>Scheuern, 1996</small>

Onthophagus schunckei <small>Paulian, 1936</small>

Onthophagus sciron <small>Balthasar, 1969</small>

Onthophagus sculptilis <small>Gerstaecker, 1871</small>

Onthophagus seabrai <small>Gomes Alves, 1944</small>

Onthophagus secundarius <small>Roth, 1851</small>

Onthophagus sellatulus <small>Orbigny, 1902</small>

Onthophagus sembeli <small>Krikken & Huijbregts, 2008</small>

Onthophagus semiaratus <small>Orbigny, 1902</small>

Onthophagus semiasper <small>Orbigny, 1902</small>

Onthophagus semicroceus <small>Orbigny, 1915</small>

Onthophagus semiflavus <small>Boheman, 1860</small>

Onthophagus semigraniger <small>Orbigny, 1905</small>

Onthophagus semigranosus <small>Lansberge, 1883</small>

Onthophagus semilaevis <small>Orbigny, 1913</small>

Onthophagus semimetallicus <small>Lea, 1923</small>

Onthophagus seminitens <small>Orbigny, 1902</small>

Onthophagus seminitidus <small>Orbigny, 1908</small>

Onthophagus semiopacus <small>Harold, 1869</small>

Onthophagus semipacificus <small>Ochi & Kon, 2006</small>

Onthophagus semivestitus <small>Paulian, 1937</small>

Onthophagus semivirescens <small>Orbigny, 1902</small>

Onthophagus semiviridis <small>Orbigny, 1904</small>

Onthophagus senegalensis <small>Orbigny, 1902</small>

Onthophagus senescens <small>Péringuey, 1908</small>

Onthophagus sepilokensis <small>Ochi & Kon, 2006</small>

Onthophagus seramicus <small>Huijbregts & Krikken, 2012</small>

Onthophagus serdangensis <small>Lansberge, 1886</small>

Onthophagus sericans <small>Frey, 1975</small>

Onthophagus sericeicollis <small>Boucomont, 1928</small>

Onthophagus serienotatus <small>Orbigny, 1902</small>

Onthophagus seseba <small>Krikken & Huijbregts, 2017</small>

Onthophagus setchan <small>Masumoto, 1984</small>

Onthophagus setosus <small>FÃ¥hraeus, 1857</small>

Onthophagus sexdentatus <small>Boucomont, 1919</small>

Onthophagus sexstriatus <small>Montrouzier, 1855</small>

Onthophagus sharpi <small>Harold, 1875</small>

Onthophagus sibela <small>Huijbregts & Krikken, 2012</small>

Onthophagus sibuyanus <small>Boucomont, 1924</small>

Onthophagus sidama <small>Gestro, 1895</small>

Onthophagus sidangeaus <small>Masumoto, Ochi & Higurashi, 2021</small>

Onthophagus signaticollis <small>Frey, 1970</small>

Onthophagus signifer <small>Harold, 1877</small>

Onthophagus sihkahonoi <small>Huijbregts & Krikken, 2009</small>

Onthophagus sikkimensis <small>Gillet, 1925</small>

Onthophagus simillimus <small>Orbigny, 1913</small>

Onthophagus simius <small>Reitter, 1892</small>

Onthophagus simoni <small>Orbigny, 1902</small>

Onthophagus simplex <small>Raffray, 1877</small>

Onthophagus simplicifrons <small>Reitter, 1892</small>

Onthophagus simulator <small>Orbigny, 1905</small>

Onthophagus singulariformis <small>Kohlmann & Solis, 2001</small>

Onthophagus sinicus <small>Hope, 1842</small>

Onthophagus sinuosicollis <small>Orbigny, 1905</small>

Onthophagus sinuosus <small>Orbigny, 1913</small>

Onthophagus sipilouensis <small>Cambefort, 1984</small>

Onthophagus sisyphoides <small>Krikken, 1977</small>

Onthophagus sjoestethi <small>Orbigny, 1904</small>

Onthophagus skelleyi <small>Sánchez-Huerta, Zunino & Halffter, 2018</small>

Onthophagus sloanei <small>Blackburn, 1903</small>

Onthophagus smeenki <small>Cambefort, 1980</small>

Onthophagus smetanai <small>Balthasar, 1952</small>

Onthophagus snoflaki <small>Balthasar, 1944</small>

Onthophagus socialis <small>Arrow, 1931</small>

Onthophagus solidus <small>Gillet, 1927</small>

Onthophagus solisi <small>Howden & Gill, 1993</small>

Onthophagus solomonensis <small>Krikken & Huijbregts, 2012</small>

Onthophagus somalicola <small>Balthasar, 1941</small>

Onthophagus songsokensis <small>Biswas & Chatterjee, 1985</small>

Onthophagus sophiae <small>Nicolas, 2006</small>

Onthophagus sopu <small>Krikken & Huijbregts, 2017</small>

Onthophagus sparsepunctatus <small>Frey, 1956</small>

Onthophagus sparsulus <small>Reitter, 1892</small>

Onthophagus spiculatus <small>Boucomont, 1914</small>

Onthophagus spinicornis <small>Gillet, 1930</small>

Onthophagus spinifex <small>(Fabricius, 1781)</small>

Onthophagus splendidoides <small>Josso, 2013</small>

Onthophagus splendidus <small>Boucomont, 1932</small>

Onthophagus spurcatus <small>Orbigny, 1905</small>

Onthophagus squalidus <small>Lea, 1923</small>

Onthophagus stanleyi <small>Moretto & Génier, 2010</small>

Onthophagus stehliki <small>Balthasar, 1966</small>

Onthophagus steinheili <small>Harold, 1880</small>

Onthophagus stellio <small>Erichson, 1843</small>

Onthophagus stenocerus <small>Harold, 1867</small>

Onthophagus sternalis <small>Arrow, 1931</small>

Onthophagus sternax <small>Balthasar, 1959</small>

Onthophagus stictus <small>Orbigny, 1913</small>

Onthophagus stigmosus <small>Orbigny, 1902</small>

Onthophagus stillatus <small>Orbigny, 1904</small>

Onthophagus stockwelli <small>Howden & Young, 1981</small>

Onthophagus stomachosus <small>Krell, 2009</small>

Onthophagus striatulus <small>(Palisot de Beauvois, 1809)</small>

Onthophagus strictestriatus <small>Orbigny, 1913</small>

Onthophagus stuhlmanni <small>Orbigny, 1908</small>

Onthophagus subaeneus <small>(Palisot de Beauvois, 1811)</small>

Onthophagus subalternans <small>Orbigny, 1902</small>

Onthophagus subcancer <small>Howden, 1973</small>

Onthophagus subcinctus <small>Orbigny, 1913</small>

Onthophagus subdivisus <small>Orbigny, 1908</small>

Onthophagus subhumeralis <small>Orbigny, 1902</small>

Onthophagus subnudus <small>Orbigny, 1902</small>

Onthophagus subocellatus <small>Orbigny, 1905</small>

Onthophagus subocelliger <small>Blackburn, 1903</small>

Onthophagus subopacus <small>Robinson, 1940</small>

Onthophagus subplanus <small>Orbigny, 1902</small>

Onthophagus subrugosus <small>Orbigny, 1905</small>

Onthophagus subsulcatus <small>Orbigny, 1908</small>

Onthophagus subtropicus <small>Howden & Cartwright, 1963</small>

Onthophagus subulifer <small>Orbigny, 1908</small>

Onthophagus suillus <small>Arrow, 1931</small>

Onthophagus sulawesiensis <small>Krikken & Huijbregts, 2011</small>

Onthophagus sulcatulus <small>Orbigny, 1907</small>

Onthophagus sulci <small>Balthasar, 1935</small>

Onthophagus sulcipennis <small>Orbigny, 1902</small>

Onthophagus sumbavensis <small>Boucomont, 1914</small>

Onthophagus sumptuosus <small>Paulian, 1937</small>

Onthophagus sundanensis <small>Lansberge, 1883</small>

Onthophagus surdus <small>Boucomont, 1925</small>

Onthophagus sutiliceps <small>Orbigny, 1902</small>

Onthophagus sydneyensis <small>Blackburn, 1903</small>

Onthophagus sylvestris <small>Walter & Cambefort, 1977</small>

Onthophagus sylvipapuanus <small>Krikken & Huijbregts, 2012</small>

Onthophagus symbioticus <small>(Arrow, 1920)</small>

Onthophagus synceri <small>Cambefort, 1984</small>

Onthophagus taayai <small>Masumoto, 1995</small>

Onthophagus tabellicornis <small>MacLeay, 1864</small>

Onthophagus tabellifer <small>Gillet, 1927</small>

Onthophagus tabidus <small>Balthasar, 1935</small>

Onthophagus taboranus <small>Orbigny, 1902</small>

Onthophagus tacanensis <small>Chamé-Vázquez & Sánchez-Hernández, 2022</small>

Onthophagus taiensis <small>Cambefort, 1984</small>

Onthophagus takaoi <small>Ochi & Kon, 2014</small>

Onthophagus talpa <small>FÃ¥hraeus, 1857</small>

Onthophagus tambing <small>Krikken & Huijbregts, 2011</small>

Onthophagus tamworthi <small>Blackburn, 1903</small>

Onthophagus tanganus <small>Orbigny, 1913</small>

Onthophagus tapirus <small>Sharp, 1877</small>

Onthophagus taprobanus <small>Arrow, 1931</small>

Onthophagus tarascus <small>Zunino & Halffter, 1988</small>

Onthophagus tarasovi <small>Krikken & Huijbregts, 2012</small>

Onthophagus tarsius <small>Arrow, 1941</small>

Onthophagus tatsienluensis <small>Balthasar, 1942</small>

Onthophagus tauroides <small>Gillet, 1930</small>

Onthophagus taurus <small>(Schreber, 1759)</small> - (Scarab beetle)

Onthophagus taxillus <small>Balthasar, 1969</small>

Onthophagus teitanicus <small>Orbigny, 1902</small>

Onthophagus telegonus <small>Balthasar, 1969</small>

Onthophagus telephus <small>Balthasar, 1969</small>

Onthophagus temporalis <small>Orbigny, 1902</small>

Onthophagus tenax <small>Balthasar, 1964</small>

Onthophagus tenebrosus <small>Harold, 1871</small>

Onthophagus tenuigraniger <small>Orbigny, 1913</small>

Onthophagus tenuistriatus <small>Orbigny, 1905</small>

Onthophagus terminatus <small>(Eschscholtz, 1822)</small>

Onthophagus terrara <small>Storey, 1977</small>

Onthophagus tersicollis <small>Müller, 1947</small>

Onthophagus tersipennis <small>Orbigny, 1902</small>

Onthophagus tersus <small>Orbigny, 1913</small>

Onthophagus tesseratus <small>Orbigny, 1908</small>

Onthophagus tessulatus <small>Harold, 1871</small>

Onthophagus testaceoviolaceus <small>Paulian, 1937</small>

Onthophagus tetricus <small>Harold, 1877</small>

Onthophagus thainuaensis <small>Masumoto, Ochi & Hanboonsong, 2008</small>

Onthophagus tharalithae <small>Karimbumkara & Priyadarsanan, 2016</small>

Onthophagus tholaayi <small>Masumoto, 1990</small>

Onthophagus thoreyi <small>Harold, 1868</small>

Onthophagus timorensis <small>Boucomont, 1914</small>

Onthophagus tiniocelloides <small>Boucomont, 1925</small>

Onthophagus tirapensis <small>Biswas & Chatterjee, 1985</small>

Onthophagus tnai <small>Nithya & Sabu, 2012</small>

Onthophagus togeman <small>Matthews, 1972</small>

Onthophagus tonsus <small>Orbigny, 1902</small>

Onthophagus tonywhitteni <small>Krikken & Huijbregts, 2011</small>

Onthophagus toopi <small>Monteith & Storey, 2013</small>

Onthophagus toraut <small>Krikken & Huijbregts, 2011</small>

Onthophagus totonacus <small>Moctezuma & Halffter, 2021</small>

Onthophagus totonicapamus <small>Bates, 1887</small>

Onthophagus toxopeus <small>Krikken & Huijbregts, 2012</small>

Onthophagus traginus <small>Frey, 1972</small>

Onthophagus transisthmius <small>Howden & Young, 1981</small>

Onthophagus transvestitus <small>Huijbregts & Krikken, 2009</small>

Onthophagus trapezicornis <small>Orbigny, 1902</small>

Onthophagus traversii <small>Orbigny, 1904</small>

Onthophagus trawalla <small>Storey & Weir, 1990</small>

Onthophagus tricariniger <small>Orbigny, 1902</small>

Onthophagus tricavicollis <small>Lea, 1923</small>

Onthophagus trichopygus <small>Orbigny, 1905</small>

Onthophagus tricolor <small>Boucomont, 1914</small>

Onthophagus tricorniger <small>Boheman, 1860</small>

Onthophagus tridenticeps <small>Orbigny, 1902</small>

Onthophagus trifidisetis <small>Orbigny, 1905</small>

Onthophagus triimpressus <small>Orbigny, 1905</small>

Onthophagus trinodosus <small>FÃ¥hraeus, 1857</small>

Onthophagus trinominatus <small>Goidanich, 1926</small>

Onthophagus tripartitus <small>Orbigny, 1902</small>

Onthophagus tripolitanus <small>Heyden, 1890</small>

Onthophagus triptolemus <small>Balthasar, 1969</small>

Onthophagus tristis <small>Harold, 1873</small>

Onthophagus tritinctus <small>Boucomont, 1914</small>

Onthophagus triundulatus <small>Balthasar, 1964</small>

Onthophagus truncaticornis <small>(Schaller, 1783)</small>

Onthophagus tschadensis <small>Balthasar, 1963</small>

Onthophagus tschoffeni <small>Orbigny, 1904</small>

Onthophagus tshuapae <small>Balthasar, 1964</small>

Onthophagus tsubakii <small>Ochi & Kon, 2009</small>

Onthophagus tuberculifrons <small>Harold, 1871</small> - (Scarab beetle)

Onthophagus tubericollis <small>Raffray, 1877</small>

Onthophagus tuckonie <small>Matthews, 1972</small>

Onthophagus tumami <small>Masumoto, Ochi & Hanboonsong, 2002</small>

Onthophagus tumidulus <small>Gerstaecker, 1871</small>

Onthophagus tungkamangensis <small>Masumoto, Ochi & Hanboonsong, 2007</small>

Onthophagus tungkamungensis <small>Masumoto, Ochi & Hanboonsong, 2008</small>

Onthophagus turfanicus <small>Balthasar, 1971</small>

Onthophagus turgidus <small>Kohlmann & Solís, 2012</small>

Onthophagus turneri <small>Boucomont, 1936</small>

Onthophagus turrbal <small>Matthews, 1972</small>

Onthophagus tuzetae <small>Walter & Cambefort, 1977</small>

Onthophagus tweedensis <small>Blackburn, 1903</small>

Onthophagus ubaidillahi <small>Krikken & Huijbregts, 2012</small>

Onthophagus uelensis <small>Frey, 1960</small>

Onthophagus ugoi <small>Moretto & Génier, 2020</small>

Onthophagus ukerewensis <small>Balthasar, 1937</small>

Onthophagus umbilicatus <small>Orbigny, 1908</small>

Onthophagus umbratus <small>Orbigny, 1902</small>

Onthophagus undulans <small>Bates, 1889</small>

Onthophagus unidentatus <small>Boucomont, 1927</small>

Onthophagus unifasciatus <small>(Schaller, 1783)</small>

Onthophagus ursinus <small>Orbigny, 1902</small>

Onthophagus ursus <small>Boucomont, 1926</small>

Onthophagus usambaricus <small>Paulian, 1937</small>

Onthophagus vaneyeni <small>Frey, 1960</small>

Onthophagus varianus <small>Lea, 1923</small>

Onthophagus variatus <small>Orbigny, 1902</small>

Onthophagus variegranosus <small>Orbigny, 1905</small>

Onthophagus variolaris <small>Lansberge, 1883</small>

Onthophagus variolicollis <small>Lea, 1923</small>

Onthophagus variolosus <small>Orbigny, 1902</small>

Onthophagus varius <small>Orbigny, 1913</small>

Onthophagus vassei <small>Orbigny, 1908</small>

Onthophagus vatovai <small>Müller, 1942</small>

Onthophagus velliger <small>Orbigny, 1907</small>

Onthophagus velutinus <small>Horn, 1875</small>

Onthophagus ventralis <small>Lansberge, 1883</small>

Onthophagus ventrosus <small>Orbigny, 1905</small>

Onthophagus veracruzensis <small>Delgado & Pensado, 1998</small>

Onthophagus vermiculatus <small>Frey, 1970</small>

Onthophagus verrucosus <small>Orbigny, 1902</small>

Onthophagus vesanus <small>Balthasar, 1967</small>

Onthophagus vespertilio <small>Howden, Cartwright & Hallfter, 1956</small>

Onthophagus vestitus <small>Orbigny, 1902</small>

Onthophagus vethi <small>Krikken, 1977</small>

Onthophagus vicinus <small>Harold, 1886</small>

Onthophagus victoriensis <small>Blackburn, 1903</small>

Onthophagus vigens <small>Péringuey, 1901</small>

Onthophagus vilis <small>Harold, 1877</small>

Onthophagus villanuevai <small>Delgado & Deloya, 1990</small>

Onthophagus villosus <small>MacLeay, 1888</small>

Onthophagus vinctoides <small>Frey, 1957</small>

Onthophagus vinctus <small>Erichson, 1843</small>

Onthophagus violaceotinctus <small>Gillet, 1925</small>

Onthophagus violaceoviridis <small>Paulian, 1937</small>

Onthophagus violetae <small>Zunino & Halffter, 1997</small>

Onthophagus virescens <small>Harold, 1867</small>

Onthophagus viridicatus <small>Orbigny, 1902</small>

Onthophagus viridichevrolati <small>Moctezuma & Halffter, 2020</small>

Onthophagus viridivinosus <small>Kohlmann & Solis, 2001</small>

Onthophagus viviensis <small>Orbigny, 1905</small>

Onthophagus vladimiri <small>Frey, 1957</small>

Onthophagus volsellatus <small>Boucomont, 1920</small>

Onthophagus vuattouxi <small>Cambefort, 1984</small>

Onthophagus vulpes <small>Harold, 1877</small>

Onthophagus vulpinaris <small>Schönfeldt, 1906</small>

Onthophagus vulpinus <small>Arrow, 1931</small>

Onthophagus vultuosus <small>Orbigny, 1902</small>

Onthophagus vultur <small>Arrow, 1931</small>

Onthophagus vylderi <small>Orbigny, 1913</small>

Onthophagus wagamen <small>Matthews, 1972</small>

Onthophagus wakelbura <small>Matthews, 1972</small>

Onthophagus walteri <small>MacLeay, 1887</small>

Onthophagus waminda <small>Matthews, 1972</small>

Onthophagus wanappe <small>Storey, 1977</small>

Onthophagus wangnamkhieoensis <small>Masumoto, Hanboonsong & Ochi, 2002</small>

Onthophagus wangnamkieous <small>Masumoto, Ochi & Hanboonsong, 2008</small>

Onthophagus waterloti <small>Orbigny, 1908</small>

Onthophagus waterstradti <small>Boucomont, 1914</small>

Onthophagus wayaua <small>Huijbregts & Krikken, 2012</small>

Onthophagus wensis <small>Josso & Prévost, 2006</small>

Onthophagus weringerong <small>Storey & Weir, 1990</small>

Onthophagus wiebesi <small>Krikken, 1977</small>

Onthophagus wigmungan <small>Matthews, 1972</small>

Onthophagus wilgi <small>Matthews, 1972</small>

Onthophagus willameorum <small>Walter & Cambefort, 1977</small>

Onthophagus williamsi <small>Storey & Weir, 1990</small>

Onthophagus witteianus <small>Krell, 2009</small>

Onthophagus wombalano <small>Matthews, 1972</small>

Onthophagus worooa <small>Storey & Weir, 1990</small>

Onthophagus xanthochlorus <small>Walter & Cambefort, 1977</small>

Onthophagus xanthomerus <small>Bates, 1887</small>

Onthophagus xanthopterus <small>Orbigny, 1908</small>

Onthophagus xanthopygus <small>Orbigny, 1910</small>

Onthophagus yackatoon <small>Storey & Weir, 1990</small>

Onthophagus yamaokai <small>Masumoto, Ochi & Hanboonsong, 2002</small>

Onthophagus yangi <small>Masumoto, Tsai & Ochi, 2006</small>

Onthophagus yanoi <small>Matsumura, 1938</small>

Onthophagus yaran <small>Storey & Weir, 1990</small>

Onthophagus yarrumba <small>Storey, 1977</small>

Onthophagus yasuhikoi <small>Masumoto, Ochi & Sakchoowong, 2012</small>

Onthophagus yescaensis <small>Moctezuma, Hernández & Sánchez-Huerta, 2022</small>

Onthophagus yeyeko <small>Matthews, 1972</small>

Onthophagus yifer <small>Krikken & Huijbregts, 2012</small>

Onthophagus yiryoront <small>Matthews, 1972</small>

Onthophagus yourula <small>Storey & Weir, 1990</small>

Onthophagus yucatanus <small>Delgado-Castillo, Peraza & Deloya, 2006</small>

Onthophagus yungaburra <small>Matthews, 1972</small>

Onthophagus yunkara <small>Matthews, 1972</small>

Onthophagus zairensis <small>Walter & Cambefort, 1977</small>

Onthophagus zambundaticeps <small>Josso, 2022</small>

Onthophagus zapotecus <small>Zunino & Halffter, 1988</small>

Onthophagus zebra <small>Arrow, 1931</small>

Onthophagus zebu <small>Boucomont, 1921</small>

Onthophagus zicsii <small>Balthasar, 1967</small>

Onthophagus zimmermanni <small>Balthasar, 1959</small>

Onthophagus zinovskyi <small>Qaryagdy, 1939</small>

Onthophagus zumpti <small>Frey, 1954</small>

Onthophagus zymoticus <small>Moxey, 1963</small>

†Onthophagus bisontinus <small>Heer, 1862</small>

†Onthophagus crassus <small>Heer, 1862</small>

†Onthophagus everestae <small>Pierce, 1946</small>

†Onthophagus luteus <small>Oustalet, 1874</small>

†Onthophagus ovatulus <small>Heer, 1847</small>

†Onthophagus pilauco <small>Tello, Verdú, Rossini & Zunino, 2021</small>

†Onthophagus prodromus <small>Heer, 1862</small>

†Onthophagus spitsbergeniensis <small>Krell, 2010</small>

†Onthophagus statzi <small>Krell, 1990</small>

†Onthophagus urusheeri <small>Krell, 2000</small>

Subgenus Palaeonthophagus <small>Zunino, 1979</small>

Onthophagus aerarius <small>Reitter, 1892</small>

Onthophagus afghanus <small>Petrovitz, 1961</small>

Onthophagus akinini <small>Koenig, 1889</small>

Onthophagus albarracinus <small>Baraud, 1979</small>

Onthophagus aleppensis <small>Redtenbacher, 1843</small>

Onthophagus amirus <small>Kabakov, 1982</small>

Onthophagus anatolicus <small>Petrovitz, 1962</small>

Onthophagus andalusicus <small>Waltl, 1835</small>

Onthophagus angorensis <small>Petrovitz, 1963</small>

Onthophagus arnoldii <small>Kabakov, 1982</small>

Onthophagus baraudi <small>Nicolas, 1964</small>

Onthophagus basipustulatus <small>Heyden, 1889</small>

Onthophagus bonsae <small>Zunino, 1976</small>

Onthophagus bytinskii <small>Balthasar, 1960</small>

Onthophagus carpanetoi <small>Pittino, 1982</small>

Onthophagus clitellifer <small>Reitter, 1894</small>

Onthophagus coenobita <small>(Herbst, 1783)</small>

Onthophagus conspersus <small>Reitter, 1892</small>

Onthophagus cruciatus <small>Ménétriés, 1832</small>

Onthophagus dacatrai <small>Pittino, 2004</small>

Onthophagus dellacasai <small>Pittino & Mariani, 1981</small>

Onthophagus dorsosignatus <small>Orbigny, 1898</small>

Onthophagus eulaminicornis <small>Pittino, 2006</small>

Onthophagus excisus <small>Reiche & Saulcy, 1856</small>

Onthophagus excubitor <small>Ziani & Gudenzi, 2006</small>

Onthophagus finschi <small>Harold, 1877</small>

Onthophagus fissicornis <small>(Steven, 1809)</small>

Onthophagus fissinasus <small>Fairmaire, 1895</small>

Onthophagus flagrans <small>Reitter, 1892</small>

Onthophagus formaneki <small>Reitter, 1897</small>

Onthophagus fortigibber <small>Reitter, 1909</small>

Onthophagus fracticornis <small>(Preyssler, 1790)</small>

Onthophagus furciceps <small>Marseul, 1869</small>

Onthophagus gibbulus <small>(Pallas, 1781)</small>

Onthophagus glasunowi <small>Koshantschikov, 1894</small>

Onthophagus gorodinskii <small>Kabakov, 2008</small>

Onthophagus grossepunctatus <small>Reitter, 1905</small>

Onthophagus hermonensis <small>Baraud, 1982</small>

Onthophagus hissariensis <small>Kabakov, 2006</small>

Onthophagus isikdagensis <small>Pittino, 2004</small>

Onthophagus jiupengensis <small>Masumoto, Lan & Kiuchi, 2017</small>

Onthophagus joannae <small>Goljan, 1953</small>

Onthophagus kindermanni <small>Harold, 1877</small>

Onthophagus kolenatii <small>Reitter, 1892</small>

Onthophagus koryoensis <small>Kim, 1985</small>

Onthophagus laticornis <small>Gebler, 1823</small>

Onthophagus latigena <small>Orbigny, 1897</small>

Onthophagus lemur <small>(Fabricius, 1781)</small>

Onthophagus lemuroides <small>Orbigny, 1898</small>

Onthophagus leucostigma <small>(Steven, 1812)</small>

Onthophagus lucidus <small>(Illiger, 1800)</small>

Onthophagus marginalis <small>(Gebler, 1817)</small>

Onthophagus massai <small>Baraud, 1975</small>

Onthophagus medius <small>(Kugelann, 1792)</small>

Onthophagus medvedevi <small>Kabakov, 1982</small>

Onthophagus merdarius <small>Chevrolat, 1865</small>

Onthophagus muelleri <small>Novak, 1921</small>

Onthophagus nebulosus <small>Reiche, 1864</small>

Onthophagus necessarius <small>Reitter, 1892</small>

Onthophagus nikolajevi <small>Kabakov, 2006</small>

Onthophagus nuchicornis <small>(Linnaeus, 1758)</small> - (Scarab beetle)

Onthophagus nurestanicus <small>Kabakov, 1982</small>

Onthophagus ocellatopunctatus <small>Waterhouse, 1875</small>

Onthophagus olsoufieffi <small>Boucomont, 1924</small>

Onthophagus opacicollis <small>Reitter, 1892</small>

Onthophagus osellai <small>Pittino, 1982</small>

Onthophagus ovatus <small>(Linnaeus, 1767)</small>

Onthophagus panici <small>Petrovitz, 1964</small>

Onthophagus parmatus <small>Reitter, 1892</small>

Onthophagus petrovitzianus <small>Pittino, 2006</small>

Onthophagus pljushtchi <small>Ivanova, 2012</small>

Onthophagus ponticus <small>Harold, 1883</small>

Onthophagus pseudocaccobius <small>Reitter, 1889</small>

Onthophagus psychopompus <small>Ziani & Garakhloo, 2010</small>

Onthophagus pygargus <small>Motschulsky, 1845</small>

Onthophagus quadrinodus <small>Reitter, 1896</small>

Onthophagus rachelis <small>Martín-Piera, 1985</small>

Onthophagus rakovici <small>Pittino, 2004</small>

Onthophagus rodentium <small>Pittino, 2004</small>

Onthophagus roessneri <small>Ziani, 2016</small>

Onthophagus ruficapillus <small>Brullé, 1832</small>

Onthophagus rufimanus <small>Kabakov, 1982</small>

Onthophagus sacharovskii <small>Olsoufieff, 1918</small>

Onthophagus samai <small>Ziani, 2011</small>

Onthophagus scabriusculus <small>Harold, 1873</small>

Onthophagus semicornis <small>(Panzer, 1798)</small>

Onthophagus sericatus <small>Reitter, 1892</small>

Onthophagus shapovalovi <small>Gusakov, 2012</small>

Onthophagus silus <small>Balthasar, 1960</small>

Onthophagus similis <small>(Scriba, 1790)</small>

Onthophagus speculifer <small>Solsky, 1876</small>

Onthophagus strabo <small>Reitter, 1892</small>

Onthophagus stylocerus <small>Graëlls, 1851</small>

Onthophagus suermelii <small>Petrovitz, 1963</small>

Onthophagus sutleinensis <small>Splichal, 1910</small>

Onthophagus suturellus <small>Brullé, 1832</small>

Onthophagus tesquorum <small>Semenov & Medvedev, 1927</small>

Onthophagus trigibber <small>Reitter, 1892</small>

Onthophagus trispinus <small>Reitter, 1892</small>

Onthophagus truchmenus <small>Kolenati, 1846</small>

Onthophagus vacca <small>(Linnaeus, 1767)</small>

Onthophagus verticicornis <small>(Laicharting, 1781)</small>

Onthophagus viridis <small>Ménétriés, 1832</small>

Onthophagus vitulus <small>(Fabricius, 1776)</small>

Onthophagus vlasovi <small>Medvedev, 1958</small>

Onthophagus zuninoi <small>Martín-Piera, 1985</small>

Onthophagus zuvandi <small>Qaryagdy, 1939</small>

Subgenus Paraphanaeomorphus <small>Balthasar, 1959</small>

Onthophagus acuticornis <small>Endrödi, 1973</small>

Onthophagus argyropygus <small>Gillet, 1927</small>

Onthophagus barbieri <small>Paulian, 1978</small>

Onthophagus bifasciatus <small>(Fabricius, 1781)</small>

Onthophagus carinensis <small>Boucomont, 1914</small>

Onthophagus comottoi <small>Lansberge, 1885</small>

Onthophagus comottoides <small>Kabakov, 1998</small>

Onthophagus cyaneus <small>Kabakov, 2006</small>

Onthophagus frugivorus <small>Arrow, 1931</small>

Onthophagus hiranoyasutoshii <small>Ochi & Kon, 2018</small>

Onthophagus insignicollis <small>Frey, 1954</small>

Onthophagus itoianus <small>Masumoto, Ochi & Higurashi, 2022</small>

Onthophagus janushevi <small>Kabakov, 1983</small>

Onthophagus jeannelianus <small>Paulian, 1945</small>

Onthophagus miichunus <small>Masumoto, Ochi & Higurashi, 2020</small>

Onthophagus morimotoi <small>Masumoto, Ochi & Higurashi, 2021</small>

Onthophagus napolovi <small>Kabakov, 1998</small>

Onthophagus phanaeicollis <small>Lansberge, 1883</small>

Onthophagus phanaeides <small>Frey, 1956</small>

Onthophagus phanaeiformis <small>Boucomont, 1914</small>

Onthophagus pseudojavanus <small>Paulian, 1931</small>

Onthophagus punneeae <small>Masumoto, 1989</small>

Onthophagus quadricolor <small>Kabakov, 1983</small>

Onthophagus saksphus <small>Masumoto, Ochi & Higurashi, 2021</small>

Onthophagus semipiceus <small>Kabakov, 1998</small>

Onthophagus shunichii <small>Ochi, Kon & Masumoto, 2021</small>

Onthophagus sobrius <small>Balthasar, 1959</small>

Onthophagus strnadi <small>Kabakov, 1998</small>

Onthophagus trituber <small>(Wiedemann, 1823)</small>

Onthophagus vaulogeri <small>Boucomont, 1923</small>

Onthophagus yujii <small>Masumoto, Ochi & Sakchoowong, 2012</small>

Subgenus Parentius <small>Zunino, 1979</small>

Onthophagus atricapillus <small>Orbigny, 1908</small>

Onthophagus emarginatus <small>Mulsant, 1842</small>

Onthophagus hispanicus <small>Baraud, 1963</small>

Onthophagus kabakovi <small>Martín-Piera, 1985</small>

Onthophagus lilliputanus <small>Lansberge, 1883</small>

Onthophagus nigellus <small>(Illiger, 1803)</small>

Onthophagus punctator <small>Reitter, 1892</small>

Onthophagus punctatus <small>(Illiger, 1803)</small>

Onthophagus zagrosicus <small>(Kabakov, 2006)</small>

Subgenus Phanaeomorphus <small>Balthasar, 1935</small>

Onthophagus ater <small>Waterhouse, 1875</small>

Onthophagus bakeri <small>Boucomont, 1919</small>

Onthophagus batanensis <small>Ochi, Kawahara & Cabras, 2021</small>

Onthophagus cernyi <small>Balthasar, 1935</small>

Onthophagus dorsofasciatus <small>Fairmaire, 1893</small>

Onthophagus fodiens <small>Waterhouse, 1875</small>

Onthophagus gagates <small>Hope, 1831</small>

Onthophagus gagatoides <small>Kabakov, 2006</small>

Onthophagus ginyunensis <small>Všetečka, 1942</small>

Onthophagus hingstoni <small>Arrow, 1931</small>

Onthophagus mindanaoensis <small>Frey, 1971</small>

Onthophagus potanini <small>Kabakov, 1979</small>

Onthophagus procurvus <small>Balthasar, 1935</small>

Onthophagus pseudojaponicus <small>Balthasar, 1941</small>

Onthophagus schaefernai <small>Balthasar, 1935</small>

Onthophagus strandi <small>Balthasar, 1935</small>

Onthophagus sycophanta <small>Fairmaire, 1887</small>

Onthophagus taiwanus <small>Nomura, 1973</small>

Onthophagus zavreli <small>Balthasar, 1935</small>

Onthophagus zetteli <small>Masumoto, Ochi & Hanboonsong, 2002</small>

Subgenus Pseudophanaeomorphus <small>Ochi, 2007</small>

Onthophagus bangueyensis <small>Boucomont, 1914</small>

Onthophagus baramtagal <small>Ochi, Kon & Barclay, 2016</small>

Onthophagus borneotagal <small>Ochi, Kon & Barclay, 2016</small>

Onthophagus chandrai <small>Ochi, 2007</small>

Onthophagus cupreotagal <small>Ochi, Kon & Barclay, 2016</small>

Onthophagus hiroyukii <small>Ochi, 2007</small>

Onthophagus johkii <small>Ochi & Kon, 1994</small>

Onthophagus koni <small>Ochi, 2007</small>

Onthophagus maryatiae <small>Ochi & Kon, 2005</small>

Onthophagus mentaveiensis <small>Boucomont, 1914</small>

Onthophagus mulutagal <small>Ochi, Kon & Barclay, 2016</small>

Onthophagus palawantagal <small>Ochi, Kon & Barclay, 2016</small>

Onthophagus parachandrai <small>Ochi, Kon & Tsubaki, 2009</small>

Onthophagus pasotagal <small>Ochi, Kon & Barclay, 2016</small>

Onthophagus quasijohkii <small>Ochi & Kon, 2005</small>

Onthophagus quasitagal <small>Ochi & Kon, 2005</small>

Onthophagus ranongjohkii <small>Ochi, Kon & Barclay, 2016</small>

Onthophagus sugihartoi <small>Ochi, 2007</small>

Onthophagus sulawesijohkii <small>Ochi, Kon & Barclay, 2016</small>

Onthophagus tagal <small>Boucomont, 1924</small>

Subgenus Serrophorus <small>Balthasar, 1935</small>

Onthophagus atropolitus <small>Orbigny, 1902</small>

Onthophagus axanicus <small>Ochi, Kon & Pham, 2021</small>

Onthophagus baolocensis <small>Ochi & Kon, 2015</small>

Onthophagus laevis <small>Harold, 1880</small>

Onthophagus mulleri <small>Lansberge, 1883</small>

Onthophagus oblongus <small>Zhang, 1997</small>

Onthophagus sagittarius <small>(Fabricius, 1775)</small> - (Sri Lankan Dung Beetle)

Onthophagus senex <small>Boucomont, 1914</small>

Onthophagus seniculus <small>(Fabricius, 1781)</small>

Onthophagus thailaevis <small>Masumoto, Ochi & Hanboonsong, 2008</small>

Subgenus Sinonthophagus <small>Kabakov, 2006</small>

Onthophagus angkhanensis <small>Masumoto, Ochi & Hanboonsong, 2013</small>

Onthophagus angustatus <small>Boucomont, 1914</small>

Onthophagus katayamai <small>Masumoto, Ochi & Sakchoowong, 2012</small>

Onthophagus nampatensis <small>Tarasov & Kabakov, 2010</small>

Onthophagus nigropubens <small>Orbigny, 1908</small>

Onthophagus poeophagus <small>Kabakov, 1979</small>

Onthophagus productus <small>Arrow, 1907</small>

Onthophagus rugulosus <small>Harold, 1886</small>

Onthophagus sumatranus <small>Lansberge, 1883</small>

Subgenus Strandius <small>Balthasar, 1935</small>

Onthophagus changshouensis <small>Zhang, 1997</small>

Onthophagus gibbicollis <small>Lansberge, 1885</small>

Onthophagus japonicus <small>Harold, 1875</small>

Onthophagus kuraruanus <small>Matsumura, 1938</small>

Onthophagus lenzii <small>Harold, 1875</small>

Onthophagus oshimanus <small>Nakane, 1960</small>

Onthophagus subansiriensis <small>Biswas, 1979</small>

Onthophagus tonkineus <small>Gillet, 1921</small>

Onthophagus yakuinsulanus <small>Nakane, 1984</small>

Subgenus Sunenaga <small>Ochi, 2003</small>

Onthophagus anguliceps <small>Boucomont, 1914</small>

Onthophagus avocetta <small>Arrow, 1933</small>

Onthophagus avocettoides <small>(Kabakov, 1983)</small>

Onthophagus blumei <small>Lansberge, 1883</small>

Onthophagus cameloides <small>Orbigny, 1900</small>

Onthophagus digitatus <small>Arrow, 1931</small>

Onthophagus kaengkrachangus <small>Masumoto, Ochi & Hanboonsong, 2008</small>

Onthophagus maesalongensis <small>Masumoto, Ochi & Hanboonsong, 2013</small>

Onthophagus mindanaensis <small>Boucomont, 1914</small>

Onthophagus ribbei <small>Boucomont, 1914</small>

Onthophagus streltsovi <small>Tarasov & Kabakov, 2010</small>

Onthophagus wallacei <small>Harold, 1871</small>

Subgenus Trichonthophagus <small>Zunino, 1979</small>

Onthophagus corniculatus <small>Reiche, 1850</small>

Onthophagus dynastoides <small>Arrow, 1931</small>

Onthophagus fumatus <small>Orbigny, 1913</small>

Onthophagus hirtus <small>(Illiger, 1803)</small>

Onthophagus juvencus <small>Klug, 1835</small>

Onthophagus loveni <small>Gillet, 1928</small>

Onthophagus maki <small>(Illiger, 1803)</small>

Onthophagus mediofuscatus <small>Orbigny, 1908</small>

Onthophagus producticollis <small>Orbigny, 1908</small>

Onthophagus quadricuspis <small>Orbigny, 1908</small>

Onthophagus sprecherae <small>Moretto, 2013</small>

Onthophagus tarandus <small>(Fabricius, 1792)</small>

Onthophagus triacanthus <small>Castelnau, 1840</small>

Onthophagus verticalis <small>FÃ¥hraeus, 1857</small>

Onthophagus aries <small>Kirby, 1825</small>

References