Hemidactylus (from Ancient Greek ἡüù- (hÃÂmi-), meaning 'half', and ôìúÃÂÃÂ
ûÿà(dáktulos), meaning 'finger') is a genus of the common gecko family, Gekkonidae. It has 199 described species, newfound ones being described every few years. These geckos are found in all the tropical regions of the world, extending into the subtropical parts of Africa and Europe. They excel in colonizing oceanic islands by rafting on flotsam, and are for example found across most of Polynesia. In some archipelagoes, cryptic species complexes are found. Geckos like to live in and out of houses. They have been introduced to many areas around the world.
This species is closely related to the genus Gehyra, which belongs to the same family in Gekkonidae.
The species are typically known as house geckos, due to their readiness to adapt to and coexist with humans, and can be easily encountered in human habitations.
Taxonomy
This genus was originally established by Lorenz Oken in 1817 for the species at that time known as Hemidactylus tuberculosus, and now described as the tropical house gecko (Hemidactylus mabouia). The species name in turn comes from the Greek words hemi "half" and dáktylos "fingers" because its toes have split or "half" lamella underneath them.
Evolution
The origin of the genus Hemidactylus is still unclear as the higher level phylogeny is not well resolved. Moreover, much of the diversity in this group still remains to be discovered.
Feces
A house gecko will usually confine its excretions to one area of a house. This is sometimes considered a nuisance by home owners, and may stain certain surfaces. The feces are approximately five (5) millimeters in length, two (2) millimeters wide, and dark brown (almost black) in color.
Description
The dorsal scale pattern is either uniform or heterogeneous. The pupil of the eye is vertical. Males have pre-anal or femoral pores. Each finger or toe has a slender distal clawed joint, angularly bent and rising from within the extremity of the dilated portion.
The fingers and toes are free, or more or less webbed, and dilated; underneath they bear a straight line running down their lamellae, which is in a pattern resembling a paripinnate compound leaf. This leads to their other and more ambiguous common name, "leaf-toed geckos", used mainly for species from South Asia and its surroundings to prevent confusion with the many "leaf-toed" Gekkota not in Hemidactylus.
Some members of the genus, such as H. platyurus, are able to run quadrupedally across water by a partially surface tension-dependent mechanism distinct from the bipedal gait of basilisks.
Communication
Like many other gecko species, species in the genus Hemidactylus are able to communicate with distinct vocalizations. Depending on the species, their vocalizations range from quiet clicks to short squeaks and chirps. For example, the Asian common house gecko (Hemidactylus frenatus) is notable for its distinctive chirping.
Species
- Hemidactylus aaronbaueri <small>, 2008</small> â Aaron Bauer's house gecko
- Hemidactylus acanthopholis <small> & , 2014</small>
- Hemidactylus achaemenidicus
- Hemidactylus adensis <small> et al., 2014</small>
- Hemidactylus aemulus <small>, & , 2022</small>
- Hemidactylus afarensis â Afar gecko
- Hemidactylus agrius <small>Vanzolini, 1978</small> â country leaf-toed gecko
- Hemidactylus albituberculatus <small>, 2012</small>
- Hemidactylus albivertebralis <small>J.-F. Trape & , 2012</small>
- Hemidactylus albofasciatus <small> & , 1963</small> â white-striped viper gecko
- Hemidactylus albopunctatus <small>Loveridge, 1947</small> â white-spotted gecko, white-spotted leaf-toed gecko
- Hemidactylus alfarraji <small>Ã
 mid, , , & , 2016</small>
- Hemidactylus alkiyumii <small>Carranza & Arnold, 2012</small>
- Hemidactylus almakhwah <small>Ã
 mÃÂd, Uvizl, Shobrak, , Salim, R. Algethami, A. Algethami, Alanazi, Alsubaie, , Nováková, & Carranza, 2022</small>
- Hemidactylus angulatus <small>Hallowell, 1854</small>
- Hemidactylus ansorgii <small>Boulenger, 1901</small> â Nigerian leaf-toed gecko
- Hemidactylus aporus <small>Boulenger, 1906</small> â Annobon leaf-toed gecko
- Hemidactylus aquilonius <small> & , 2007</small>
- Hemidactylus arnoldi <small>Lanza, 1978</small> â Arnold's leaf-toed gecko
- Hemidactylus asirensis <small>Ã
 mid, Shobrak, Wilms, Joger & Carranza, 2016</small>
- Hemidactylus awashensis <small>Ã
 mÃÂd et al., 2014</small>
- Hemidactylus barbierii <small>, & , 2007</small>
- Hemidactylus barbouri <small>Loveridge, 1942</small> â BarbourâÂÂs leaf-toed gecko
- Hemidactylus barodanus <small>Boulenger, 1901</small> â enigmatic gecko
- Hemidactylus bavazzanoi <small>Lanza, 1978</small> â Somali banded gecko, BavazzanoâÂÂs gecko
- Hemidactylus bayonii <small>Bocage, 1893</small> â Barboza's leaf-toed gecko
- Hemidactylus benguellensis <small>Bocage, 1893</small> â Benguela house gecko
- Hemidactylus beninensis <small>, , & , 2006</small>
- Hemidactylus biokoensis <small>, & , 2014</small> â Bioko leaf-toed gecko
- Hemidactylus boavistensis <small>Boulenger, 1906</small> â Boa Vista leaf-toed gecko
- Hemidactylus bouvieri <small>(Bocourt, 1870)</small> â Bouvier's leaf-toed gecko, Cape Verde leaf-toed gecko
- Hemidactylus bowringii <small>(Gray, 1845)</small> â Oriental leaf-toed gecko, Bowring's gecko, Sikkimese dark-spotted gecko, Asian smooth gecko
- Hemidactylus brasilianus <small>Amaral, 1935</small> â Amaral's Brazilian gecko
- Hemidactylus brookii <small>Gray, 1845</small> â Brooke's house gecko, spotted house gecko
- Hemidactylus carivoensis <small>, , Iglesias, , , & , 2021</small>
- Hemidactylus chikhaldaraensis <small>, Bauer, Giri & , 2019</small> â Chikhaldara brookiish gecko
- Hemidactylus chipkali <small>Mirza & , 2017</small> â Central Indian leaf-toed gecko
- Hemidactylus cinganji <small>Lobón-Rovira, Conradie, Iglesias, R. Ernst, L. VerÃÂssimo, Baptista & Pinto, 2021</small>
- Hemidactylus citernii <small>Boulenger, 1912</small> â speedy leaf-toed gecko
- Hemidactylus coalescens <small>Wagner, Leaché & Fujita, 2014</small>
- Hemidactylus craspedotus <small>Mocquard, 1890</small> â Mocquard's house gecko, frilled gecko, frilled house gecko
- Hemidactylus curlei <small>Parker, 1942</small> â Parker's gecko, northern leaf-toed gecko
- Hemidactylus dawudazraqi <small>Moravec, , , , & , 2011</small>
- Hemidactylus depressus <small>Gray, 1842</small> â Sri Lankan leaf-nosed gecko, Kandyan gecko
- Hemidactylus dracaenacolus <small>Rösler & , 1999</small>
- Hemidactylus easai <small>, , , , & , 2022</small>
- Hemidactylus echinus <small>O'Shaughnessy, 1875</small> â hedgehog leaf-toed gecko
- Hemidactylus endophis <small>Carranza & Arnold, 2012</small>
- Hemidactylus eniangii <small>Wagner, Leaché & Fujita, 2014</small>
- Hemidactylus farasani <small>Ã
 mÃÂd, Uvizl, Shobrak, Busais, Salim, R. Algethami, A. Algethami, Alanazi, Alsubaie, Rovatsos, Nováková, Mazuch & Carranza, 2022</small> â Farasan gecko
- Hemidactylus fasciatus <small>Gray, 1842</small> â banded leaf-toed gecko
- Hemidactylus faustus <small>Lobón-Rovira, Conradie, Iglesias, R. Ernst, L. VerÃÂssimo, Baptista & Pinto, 2021</small>
- Hemidactylus festivus <small>Carranza & Arnold, 2012</small>
- Hemidactylus flavicauda <small>, Giri, & Agarwal, 2020</small> â Mahabubnagar yellow-tailed brookiish gecko
- Hemidactylus flaviviridis <small>Rüppell, 1835</small> â yellow-bellied gecko, northern house gecko
- Hemidactylus forbesii <small>Boulenger, 1899</small> â Socotra leaf-toed gecko
- Hemidactylus foudaii <small>, 2003</small>
- Hemidactylus fragilis <small>Calabresi, 1915</small>
- Hemidactylus frenatus <small>Schlegel, 1836</small> â common house gecko, Asian house gecko, Pacific house gecko
- Hemidactylus funaiolii <small>Lanza, 1978</small> â Archer's post gecko, Kenya leaf-toed gecko
- Hemidactylus garnotii <small>A.M.C. Duméril & Bibron, 1836</small> â Indo-Pacific gecko, Garnot's house gecko, fox gecko, Assam greyish-brown gecko
- Hemidactylus giganteus <small>Stoliczka, 1871</small> â giant leaf-toed gecko, giant southern tree gecko, giant gecko
- Hemidactylus gleadowi <small>Murray, 1884</small> â Gleadow's house gecko
- Hemidactylus gracilis <small>Blanford, 1870</small> â graceful leaf-toed gecko
- Hemidactylus gramineus <small>, Bauer, , Agarwal & , 2021</small>
- Hemidactylus granchii <small>Lanza, 1978</small> â Granchi's leaf-toed gecko
- Hemidactylus graniticolus <small>Agarwal, Giri & Bauer, 2011</small>
- Hemidactylus granosus <small>Heyden, 1827</small>
- Hemidactylus granti <small>Boulenger, 1899</small> â Grant's leaf-toed gecko
- Hemidactylus greeffii <small>Bocage, 1886</small> â Greeff's gecko, Greeff's giant gecko
- Hemidactylus gujaratensis <small>Giri, Bauer, & , 2009</small> â Gujarat gecko
- Hemidactylus hajarensis <small>Carranza & Arnold, 2012</small>
- Hemidactylus hegdei <small>Pal & Mirza, 2022</small>
- Hemidactylus hemchandrai <small> & , 2015</small>
- Hemidactylus homoeolepis <small>Blanford, 1881</small> â Arabian leaf-toed gecko
- Hemidactylus hunae <small>Deraniyagala, 1937</small> â spotted giant gecko
- Hemidactylus imbricatus <small>Bauer et al., 2008</small> â carrot-tailed viper gecko
- Hemidactylus inexpectatus <small>Carranza & Arnold, 2012</small>
- Hemidactylus inintellectus <small>Sindaco, Ziliani, Razzetti, , , 2009</small> â Socotran rock gecko
- Hemidactylus isolepis <small>Boulenger, 1895</small> â scaly leaf-toed gecko, uniform-scaled gecko
- Hemidactylus ituriensis <small>Schmidt, 1919</small> â Ituri leaf-toed gecko
- Hemidactylus jubensis <small>Boulenger, 1895</small> â Ethiopian gecko, Mrioen leaf-toed gecko
- Hemidactylus jumailiae <small>Busais & Joger, 2011</small>
- Hemidactylus kamdemtohami <small>Bauer & Pauwels, 2002</small> â Kamdem Toham's gecko
- Hemidactylus kangerensis <small>Mirza, & , 2017</small>
- Hemidactylus karenorum <small>(Theobald, 1868)</small> â Burmese spotted gecko, Burmese leaf-toed gecko
- Hemidactylus kimbulae <small>, , , & , 2021</small>
- Hemidactylus klauberi <small>, 1948</small>
- Hemidactylus kolliensis <small>Agarwal, Bauer, Giri & Khandekar, 2019</small> â Kolli rock gecko
- Hemidactylus kundaensis <small> & J.-F. Trape, 2012</small>
- Hemidactylus kushmorensis <small>Murray, 1884</small> â Kushmore house gecko
- Hemidactylus kyaboboensis <small>Wagner, Leaché & Fujita, 2014</small>
- Hemidactylus laevis <small>Boulenger, 1901</small> â common leaf-toed gecko
- Hemidactylus lamaensis <small>, & Böhme, 2010</small>
- Hemidactylus lankae <small>Deraniyagala, 1953</small> â termite hill gecko, Sri Lankan leaf-toed gecko
- Hemidactylus lanzai
- Hemidactylus laticaudatus <small>L.G. Andersson, 1910</small> â Andersson's leaf-toed gecko
- Hemidactylus lavadeserticus <small>Moravec & Bohme, 1997</small> â Syrian house gecko
- Hemidactylus lemurinus <small>Arnold, 1980</small> â Dhofar leaf-toed gecko, Oman ghost leaf-toed gecko
- Hemidactylus leschenaultii <small>A.M.C. Duméril & Bibron, 1836</small> â Leschenault's leaf-toed gecko
- Hemidactylus longicephalus <small>Bocage, 1873</small>
- Hemidactylus lopezjuradoi <small>Arnold, , , & Carranza, 2008</small> (formerly in H. bouvieri)
- Hemidactylus luqueorum <small>Carranza & Arnold, 2012</small>
- Hemidactylus mabouia <small>(, 1818)</small> â tropical house gecko, Afro-American house gecko, cosmopolitan house gecko
- Hemidactylus macropholis <small>Boulenger, 1896</small> â BoulengerâÂÂs gecko, large-scaled leaf-toed gecko
- Hemidactylus maculatus <small>A.M.C. Duméril & Bibron, 1836</small> â spotted leaf-toed gecko, giant spotted gecko
- Hemidactylus mahonyi <small>, , Kumar, , & , 2022</small>
- Hemidactylus makolowodei <small>Bauer, , Chirio, & , 2006</small>
- Hemidactylus malcolmsmithi <small>(, 1949)</small> â Malcolm's bow-fingered gecko, Smith's bent-toed gecko
- Hemidactylus mandebensis <small>Ã
 mÃÂd et al., 2014</small>
- Hemidactylus masirahensis <small>Carranza & Arnold, 2012</small>
- Hemidactylus matschiei <small>(Tornier, 1901)</small> â Togo leaf-toed gecko
- Hemidactylus megalops <small>Parker, 1932</small> â Parker's leaf-toed gecko
- Hemidactylus mercatorius <small>Gray, 1842</small>
- Hemidactylus mindiae <small>S. Baha El Din, 2005</small> â Mount Sinai gecko
- Hemidactylus minutus <small>Vasconcelos & Carranza, 2014</small>
- Hemidactylus modestus <small>(Günther, 1894)</small> â moderate leaf-toed gecko, Tana River gecko
- Hemidactylus montanus <small>Busais & Joger, 2011</small> â mountain leaf-toed gecko
- Hemidactylus mrimaensis <small> & Bauer, 2014</small> â Kaya gecko
- Hemidactylus muriceus <small>W. Peters, 1870</small> â Guinea leaf-toed gecko
- Hemidactylus murrayi <small>, 1887</small> â Murray's house gecko
- Hemidactylus multisulcatus â Madurai rock gecko
- Hemidactylus newtoni <small>, 1897</small> â Newton's leaf-toed gecko
- Hemidactylus nicolauensis <small>Vasconcelos, G. Köhler, & , 2020</small>
- Hemidactylus nzingae â Queen NzingaâÂÂs tropical gecko
- Hemidactylus ophiolepis <small>Boulenger, 1903</small> â snake-scaled leaf-toed gecko
- Hemidactylus ophiolepoides <small>Lanza, 1978</small> â Lanza's leaf-toed gecko
- Hemidactylus oxyrhinus <small>Boulenger, 1899</small> â sharp-nosed leaf-toed gecko
- Hemidactylus paaragowli <small>, , & Pal, 2018</small> â Travancore rock gecko
- Hemidactylus paivae â PaivaâÂÂs gecko
- Hemidactylus pakkamalaiensis
- Hemidactylus palaichthus <small>Kluge, 1969</small> â Antilles leaf-toed gecko
- Hemidactylus parvimaculatus <small>Deraniyagala, 1953</small> â spotted house gecko
- Hemidactylus paucifasciatus
- Hemidactylus pauciporosus <small>Lanza, 1978</small>
- Hemidactylus paucituberculatus <small>Carranza & Arnold, 2012</small>
- Hemidactylus persicus <small>J. Anderson, 1872</small> â Persian leaf-toed gecko, Persian gecko
- Hemidactylus pfindaensis <small>Lobón-Rovira, Conradie, Iglesias, R. Ernst, L. VerÃÂssimo, Baptista & Pinto, 2021</small>
- Hemidactylus pieresii <small>Kelaart, 1852</small>
- Hemidactylus platycephalus <small>W. Peters, 1854</small> â tree gecko, flat-headed leaf-toed gecko, baobab gecko
- Hemidactylus platyurus <small>(Schneider, 1792)</small> â flat-tailed house gecko, frilled house gecko, Asian house gecko
- Hemidactylus porbandarensis <small>, 1981</small>
- Hemidactylus prashadi <small>M.A. Smith, 1935</small> â Bombay leaf-toed gecko, Prashad's gecko
- Hemidactylus principensis <small>, & , 2012</small>
- Hemidactylus pseudomuriceus <small> & Böhme, 2003</small>
- Hemidactylus pseudoromeshkanicus
- Hemidactylus puccionii <small>Calabresi, 1927</small> â Zanzibar leaf-toed gecko, Somali plain gecko
- Hemidactylus pumilio <small>Boulenger, 1899</small> â pygmy leaf-toed gecko
- Hemidactylus quartziticolus
- Hemidactylus raya <small>Kumar, A. Srinivasulu & C. Srinivasulu, 2022</small>
- Hemidactylus reticulatus <small>Beddome, 1870</small> â reticulated leaf-toed gecko
- Hemidactylus richardsonii <small>Gray, 1845</small> â Richardson's leaf-toed gecko
- Hemidactylus rishivalleyensis <small> Agarwal, Thackeray & Khandekar, 2020</small> â Rishi Valley rock gecko
- Hemidactylus robustus <small>Heyden, 1827</small> â Heyden's gecko
- Hemidactylus romeshkanicus <small>Torki, 2011</small>
- Hemidactylus ruspolii <small>Boulenger, 1896</small> â farm leaf-toed gecko, Ruspoli's gecko, turnip-tailed black and yellow gecko
- Hemidactylus saba <small>Busais & Joger, 2011</small>
- Hemidactylus sahgali <small>Mirza, , R. Patil, & , 2018</small> â SahgalâÂÂs termite hill gecko
- Hemidactylus sankariensis <small>Agarwal, Bauer, Giri & Khandekar, 2019</small> â Sankari brookiish gecko
- Hemidactylus sassanidianus
- Hemidactylus sataraensis <small>Giri & Bauer, 2008</small> â Satara gecko
- Hemidactylus saxicolus <small>Kumar, A. Srinivasulu & C. Srinivasulu, 2022</small>
- Hemidactylus scabriceps <small>(Annandale, 1906)</small> â scaly gecko
- Hemidactylus shihraensis <small>Busais & Joger, 2011</small>
- Hemidactylus sinaitus <small>Boulenger, 1885</small> â Red Sea gecko, Sinai leaf-toed gecko
- Hemidactylus sirumalaiensis <small>Khandekar, Thackeray, Pawar & Agarwal, 2020</small>
- Hemidactylus siva <small>C. Srinivasulu, A. Srinivasulu & Kumar, 2018</small> â Hampi rock gecko
- Hemidactylus smithi <small>Boulenger, 1895</small> â Smith's leaf-toed gecko
- Hemidactylus somalicus <small>Parker, 1932</small> â Northern Somali leaf-toed gecko
- Hemidactylus squamulatus <small>Tornier, 1896</small> â Tornier's leaf-toed gecko, Nyika gecko
- Hemidactylus srikanthani <small>Adhikari, Achyuthan, Kumar, Khot, Shreeram & Ganesh, 2022</small>
- Hemidactylus stejnegeri <small> & , 1989</small> â Stejneger's leaf-toed gecko
- Hemidactylus subtriedrus <small>Jerdon, 1854</small> â Jerdon's gecko, Madras blotched gecko
- Hemidactylus sushilduttai <small>Giri, Bauer, Mohapatra, C. Srinivasulu & Agarwal, 2017</small> â Dutta Mahendragiri gecko
- Hemidactylus tamhiniensis <small>Khandekar, Thackeray & Agarwal, 2021</small> â Tamhini giant rock gecko, basalt giant rock gecko
- Hemidactylus tanganicus <small>Loveridge, 1929</small> â Tanzanian leaf-toed gecko, Tanzanian diamond gecko, Dutumi gecko
- Hemidactylus tasmani <small>Hewitt, 1932</small> â Tasmanian leaf-toed gecko
- Hemidactylus taylori <small>Parker, 1932</small>
- Hemidactylus tenkatei <small>Lidth de Jeude, 1895</small>
- Hemidactylus thayene <small>McMahan & Zug, 2007</small>
- Hemidactylus treutleri <small>, 2009</small> â Treutler's gecko
- Hemidactylus triedrus <small>(Daudin, 1802)</small> â termite hill gecko, Dakota's leaf-toed gecko, blotched house gecko
- Hemidactylus tropidolepis <small>Mocquard, 1888</small> â Mocquard's leaf-toed gecko, Ogaden gecko
- Hemidactylus turcicus <small>Linnaeus, 1758</small> â Mediterranean house gecko, Turkish gecko
- Hemidactylus ulii <small>Ã
 mÃÂd, Moravec, Kratochvil, Gvozdik, , Busais, Wilms, Shobrak & Carranza, 2013</small>
- Hemidactylus vanam <small>, Lajmi & Giri, 2018 </small> â Megamalai rock gecko
- Hemidactylus varadgirii <small>Chaitanya, Agarwal, Lajmi & Khandekar, 2019</small> â GiriâÂÂs brookiish gecko, Amboli brookiish gecko
- Hemidactylus vernayi <small>CerÃÂaco, Agarwal, & Bauer, 2020</small> â VernayâÂÂs tropical gecko
- Hemidactylus vietnamensis <small>Darevsky, & , 1984</small> â Vietnam leaf-toed gecko, Vietnam house gecko
- Hemidactylus vijayraghavani <small>Mirza, 2018</small>
- Hemidactylus whitakeri <small>Mirza, Gowande, R. Patil, Ambekar & Patel, 2018</small> â WhitakerâÂÂs termite hill gecko
- Hemidactylus xericolus <small>Lajmi, Giri, Singh & Agarwal, 2020</small> â Nalgonda yellow-tailed brookiish gecko
- Hemidactylus yajurvedi <small>, Bauer, Lajmi, Agarwal & Giri, 2015</small> â Kanker rock gecko
- Hemidactylus yerburyi <small>J. Anderson, 1895</small> â YerburyâÂÂs gecko, YerburiâÂÂs leaf-toed gecko
- Hemidactylus aravalliensis
- Hemidactylus kalinga
- Hemidactylus amarasinghei
- Hemidactylus gubanensis
- Hemidactylus huluul
Phylogeny
The following phylogeny is from Pyron, et al. (2013), and includes 47 Hemidactylus species. Hemidactylus is a sister group of Cyrtodactylus.
References
External links