Geomitridae is a taxonomic family of small to medium-sized air-breathing land snails, terrestrial pulmonate gastropod mollusks in the superfamily Helicoidea.
Anatomy
The family is characterized by the presence of a free right ommatophoral retractor (passing outside the peni-oviducal angle) and a double stimulatory apparatus. A free right ommatophoral retractor has been linked to adaptation to xeric habitats.
Taxonomy
The family Geomitridae is subdivided in the following subfamilies (according to Razkin et al., 2015). Genera with the family Geomitridae include:
Family Geomitridae<br>
- Not belonging to a subfamily:
- Keraea <small>Gude, 1911</small>
- Peridotitea <small>Torres Alba, Holyoak, D. T., Holyoak, G. A., Vázquez Toro & Ripoll, 2018</small>
- Subfamily Geomitrinae
- tribe Cochlicellini <small>Schileyko, 1972</small>
- Cochlicella <small>Férussac, 1821</small>
- Monilearia <small>Mousson, 1872</small>
- Obelus <small>W. Hartmann, 1842</small>
- Ripkeniella <small>Hutterer & E. Gittenberger, 1998</small>
- tribe Geomitrini <small>C. Boettger, 1909 - synonym: Ochthephilinae Zilch, 1960 (n.a.)</small>
- Actinella <small>R. T. Lowe, 1852</small>
- Callina <small>R. T. Lowe, 1855</small>
- Caseolus <small>R. T. Lowe, 1852</small>
- Discula <small>R. T. Lowe, 1852</small>
- Disculella <small>Pilsbry, 1895</small>
- Domunculifex <small>Brozzo, De Mattia, Harl & Neiber, 2020</small>
- Geomitra <small>Swainson, 1840</small>
- Helicomela <small>R. T. Lowe, 1855</small>
- Heterostoma <small>W. Hartmann, 1843</small>: synonym of Steenbergia <small>Mandahl-Barth, 1950</small> (invalid: junior homonym of Heterostoma <small>de Filippi, 1837</small> [Platyhelminthes])
- Hystricella <small>R. T. Lowe, 1855</small>
- Lemniscia <small>R. T. Lowe, 1855</small>
- Moreletina <small>de Frias Martins, 2002</small>
- Plebecula <small>R. T. Lowe, 1852</small>
- Pseudocampylaea <small>L. Pfeiffer, 1877</small>
- Serratorotula <small>Groh & Hemmen, 1986</small>
- Spirorbula <small>R. T. Lowe, 1852</small>
- Steenbergia <small>Mandahl-Barth, 1950</small>
- Testudodiscula <small>Brozzo, De Mattia, Harl & Neiber, 2020</small>
- Wollastonaria <small>De Mattia, Neiber & Groh, 2018</small>
- tribe Ponentinini <small>Razkin et al., 2015</small>
- Ponentina <small>P. Hesse, 1921</small>
- subfamily Helicellinae
- tribe Cernuellini <small>Schileyko, 1991</small>
- Alteniella <small>Clerx & Gittenberger, 1977</small>
- Cernuella <small>Schlüter, 1838</small>
- Microxeromagna <small>Ortiz de Zárate López, 1950</small>
- Xeroplana <small>Monterosato, 1892</small>
- Xerosecta <small>Monterosato, 1892</small>
- tribe Helicellini <small>Ihering, 1909 - synonym: Jacostidae Pilsbry, 1948 (inv.)</small>
- Backeljaia <small>Chueca et al., 2018</small>
- Candidula <small>Kobelt, 1871</small>
- Helicella <small>Férussac, 1821</small>
- Orexana <small>Chueca et al., 2018</small>
- Xerogyra <small>Monterosato, 1892</small>
- Xeroleuca <small>Kobelt, 1877</small>
- Xeroplexa <small>Monterosato, 1892</small>
- Xerotricha <small>Monterosato, 1892</small>
- Zarateana <small>Chueca et al., 2018</small>
- tribe Helicopsini <small>H. Nordsieck, 1987</small>
- Helicopsis <small>Fitzinger, 1833</small>
- Pseudoxerophila <small>Westerlund, 1879</small>
- Xerolenta <small>Monterosato, 1892</small>
- Xeromunda <small>Monterosato, 1892</small>
- Xeropicta <small>Monterosato, 1892</small>
- tribe Plentuisini <small>Razkin et al., 2015</small>
- Plentuisa <small>Puente & Prieto, 1992</small>
- tribe Trochoideini <small>H. Nordsieck, 1987</small>
- Trochoidea <small>T. Brown, 1827</small>
- Xerocrassa <small>Monterosato, 1892</small>
References
- Bouchet P., Rocroi J.P., Hausdorf B., Kaim A., Kano Y., Nützel A., Parkhaev P., Schrödl M. & Strong E.E. (2017). Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia. 61(1-2): 1-526