Syllidae, commonly known as the necklace worms, is a family of small to medium-sized polychaete worms. Syllids are distinguished from other polychaetes by the presence of a muscular region of the anterior digestive tract known as the proventricle.
Syllid worms range in size from to . Most syllids are benthic organisms that transition to a pelagic epitoke for reproduction. They are found in all regions of the ocean, from the intertidal zone to the deep sea, and are especially abundant in shallow water.
They are found in a range of habitats, moving actively on rock and sandy substrates, hiding in crevices and among seaweeds, and climbing on sponges, corals, hydrozoans, seagrasses and mangroves. They are generalist feeders. A young Syllid was one of the first worms to be found with pollen from seagrass in its stomach, making it a possible pollinator.
The proventricle, Syllid worm's most distinctive anatomical feature, allows the worm to feed by sucking due to its pumping action. It also plays a role in hormone production, and thus the worm's sexual development. The proventricle is composed of strirated muscle cells with the longest known sacromeres among animals. The proventricle is usually visible through the body wall.
Syllis ramosa was the first polychaete discovered to have a branching body plan. Later, two species of Ramisyllis were discovered to have a branching body plan.
Subfamilies
- Anoplosyllinae
- Streptosyllis
- Streptosyllis bidentata <small>(Southern, 1914)</small>
- Streptosyllis websteri <small>(Southern, 1914)</small>
- Syllides
- Syllides benedicti <small>(Banse, 1971)</small>
- Syllides longocirrata <small>(ÃÂrsted, 1845)</small>
- Eusyllinae
- Clavisyllis
- Clavisyllis alternata <small>Knox, 1957</small>
- Clavisyllis tenjini <small>Cejp, Jimi & Aguado, 2023</small>
- Clavisyllis yongei <small>Watson, 2009</small>
- Eusyllis
- Eusyllis assimilis <small>(Marenzeller, 1875)</small>
- Eusyllis blomstrandi <small>(Malmgren, 1867)</small>
- Eusyllis lamelligera <small>(Marion & Bobretzky, 1875)</small>
- Nudisyllis
- Nudisyllis divaricata <small>(Keferstein, 1862)</small>
- Nudisyllis pulligera <small>(Krohn, 1852)</small>
- Odontosyllis
- Odontosyllis ctenostoma <small>(Claparède, 1868)</small>
- Odontosyllis enopla <small>(Verrill, 1900)</small>
- Odontosyllis fulgurans <small>(Audouin & Milne-Edwards, 1833)</small>
- Odontosyllis gibba <small>(Claparède, 1863)</small>
- Odontosyllis phosphorea <small>(Moore, 1909)</small>
- Opisthodonta
- Opisthodonta longocirrata <small>(Saint-Joseph, 1886)</small>
- Pionosyllis
- Pionosyllis compacta <small>(Malmgren, 1867)</small>
- Pionosyllis serratisetosa <small>(López, San MartÃÂn & Jiménez, 1997)</small>
- Synmerosyllis
- Synmerosyllis lamelligera <small>(Saint-Joseph, 1886)</small>
- Autolytinae
- Epigamia
- Epigamia alexandri <small>(Malmgren, 1867)</small>
- Myrianida
- Myrianida brachycephala <small>(Marenzeller, 1874)</small>
- Myrianida edwarsi <small>(Saint-Joseph, 1886)</small>
- Myrianida inermis <small>(Saint-Joseph, 1886)</small>
- Myrianida langerhansi <small>(Gidholm, 1967)</small>
- Myrianida pinnigera <small>(Montagu, 1808)</small>
- Myrianida prolifera <small>(O.F. Müller, 1788)</small>
- Myrianida quinquedecimdentata <small>(Langerhans, 1884)</small>
- Myrianida rubropunctata <small>(Grube, 1860)</small>
- Proceraea
- Proceraea aurantiaca <small>(Claparède, 1868)</small>
- Proceraea cornuta <small>(Agassiz, 1862)</small>
- Proceraea picta <small>(Ehlers, 1864)</small>
- Proceraea prismatica <small>(O.F. Müller, 1776)</small>
- Procerastea halleziana <small>(Malaquin, 1893)</small>
- Procerastea nematodes <small>(Langerhans, 1884)</small>
- Exogoninae
- Brania
- Brania pusilla <small>(Dujardin, 1851)</small>
- Erinaceusyllis
- Erinaceusyllis cirripapillata <small>(San MartÃÂn, 2005)</small>
- Erinaceusyllis erinaceus <small>(Claparède, 1863)</small>
- Erinaceusyllis ettiennei <small>(San MartÃÂn, 2005)</small>
- Erinaceusyllis hartmannschroederae <small>(San MartÃÂn, 2005)</small>
- Erinaceusyllis kathrynae <small>(San MartÃÂn, 2005)</small>
- Exogone
- Exogone dispar <small>(Webster, 1879)</small>
- Exogone naidina <small>(ÃÂrsted, 1845)</small>
- Exogone verugera <small>(Claparède, 1868)</small>
- Exogonita <small>Hartman & Fauchald, 1971</small>
- Parapionosyllis <small>Fauvel, 1923</small>
- Parexogone
- Parexogone longicirris <small>(Webster & Benedict, 1887)</small>
- Parexogone hebes <small>(Webster & Benedict, 1884)</small>
- Prosphaerosyllis
- Prosphaerosyllis tetralix <small>(Eliason, 1920)</small>
- Salvatoria <small>McIntosh, 1885</small>
- Salvatoria clavata <small>(Claparède, 1863)</small>
- Salvatoria limbata <small>(Claparède, 1868)</small>
- Salvatoria swedmarki <small>(Gidholm, 1962)</small>
- Sphaerosyllis
- Sphaerosyllis bardukaciculata <small>(San MartÃÂn, 2005)</small>
- Sphaerosyllis bulbosa <small>(Southern, 1914)</small>
- Sphaerosyllis goorabantennata <small>(San MartÃÂn, 2005)</small>
- Sphaerosyllis georgeharrisoni <small>(San MartÃÂn, 2005)</small>
- Sphaerosyllis hystrix <small>(Claparède, 1863)</small>
- Sphaerosyllis levantina<small>(Faulwetter et al., 2011)</small>
- Sphaerosyllis pirifera <small>(Claparède, 1868)</small>
- Sphaerosyllis taylori <small>(Perkins, 1980)</small>
- Sphaerosyllis voluntariorum <small>(San MartÃÂn, 2005)</small>
- Syllinae
- Syllis
- Syllis aciculigrossa <small>(San MartÃÂn, 1990)</small>
- Syllis adamantea <small>(Treadwell, 1914)</small>
- Syllis albae <small>(ÃÂlvarez-Campos & Verdes, 2017)</small>
- Syllis albanyensis <small>(Hartmann-Schröder, 1984)</small>
- Syllis alosae <small>(San MartÃÂn, 1992)</small>
- Syllis alternata <small>(Moore, 1908)</small>
- Syllis amica <small>(Quatrefages, 1866)</small>
- Syllis amicarmillaris <small>(Simon, San MartÃÂn & Robinson, 2014)</small>
- Syllis anoculata <small>(Hartmann-Schröder, 1962)</small>
- Syllis antoniae <small>(Salcedo Oropeza, San MartÃÂn & SolÃÂs-Weiss, 2012)</small>
- Syllis armillaris <small>(O.F. Müller, 1776)</small>
- Syllis augeneri <small>(Haswell, 1920)</small>
- Syllis barbata <small>(San MartÃÂn, 1992)</small>
- Syllis bella <small>(Chamberlin, 1919)</small>
- Syllis beneliahuae <small>(Campoy & Alquézar, 1982)</small>
- Syllis benguellana <small>(Day, 1963)</small>
- Syllis boggemanni <small>(San MartÃÂn, ÃÂlvarez-Campos & Hutchings, 2017)</small>
- Syllis botosaneanui <small>(Hartmann-Schröder, 1973)</small>
- Syllis brasiliensis <small>(McIntosh, 1885)</small>
- Syllis breviarticulata <small>(Grube, 1857)</small>
- Syllis brevicirrata <small>(McIntosh, 1908)</small>
- Syllis brevicirris <small>(Hansen, 1882)</small>
- Syllis broomensis <small>(Hartmann-Schröder, 1979)</small>
- Syllis caeca <small>(Monro, 1933)</small>
- Syllis castroviejoi <small>(Capa, San MartÃÂn & López, 2001)</small>
- Syllis cerina <small>(Grube, 1878)</small>
- Syllis cirrita <small>(Lee & Rho, 1994)</small>
- Syllis columbretensis <small>(Campoy, 1982)</small>
- Syllis compacta <small>(Gravier, 1900)</small>
- Syllis corallicola <small>(Verrill, 1900)</small>
- Syllis cornuta <small>(Rathke, 1843)</small>
- Syllis crassicirrata <small>(Treadwell, 1925)</small>
- Syllis cruzi <small>(Núñez & San MartÃÂn, 1991)</small>
- Syllis curticirris <small>(Monro, 1937)</small>
- Syllis danieli <small>(San MartÃÂn, 1992)</small>
- Syllis deleoni <small>(Salcedo Oropeza, San MartÃÂn & SolÃÂs-Weiss, 2012)</small>
- Syllis edensis <small>(Hartmann-Schröder, 1989)</small>
- Syllis elongata <small>(Johnson, 1901)</small>
- Syllis ergeni <small>(ÃÂinar, 2005)</small>
- Syllis erikae <small>(Hartmann-Schröder, 1981)</small>
- Syllis fasciata <small>(Malmgren, 1867)</small>
- Syllis ferrani <small>(Alós & San MartÃÂn, 1987)</small>
- Syllis filidentata <small>(Hartmann-Schröder, 1962)</small>
- Syllis garciai <small>(Campoy, 1982)</small>
- Syllis gerlachi <small>(Hartmann-Schröder, 1960)</small>
- Syllis gerundensis <small>(Alós & Campoy, 1981)</small>
- Syllis glandulata <small>(Nogueira & San MartÃÂn, 2002)</small>
- Syllis glarearia <small>(Westheide, 1974)</small>
- Syllis golfonovensis <small>(Hartmann-Schröder, 1962)</small>
- Syllis gracilis <small>(Grube, 1840)</small>
- Syllis guidae <small>(Nogueira & Yunda-Guarin, 2008)</small>
- Syllis heterochaeta <small>(Moore, 1909)</small>
- Syllis hyalina <small>(Grube, 1863)</small>
- Syllis hyllebergi <small>(Licher, 1999)</small>
- Syllis joaoi <small>(San MartÃÂn, ÃÂlvarez-Campos & Hutchings, 2017)</small>
- Syllis jorgei <small>(San MartÃÂn & López, 2000)</small>
- Syllis kabilica <small>(Ben-Eliahu, 1977)</small>
- Syllis karlae <small>(San MartÃÂn, ÃÂlvarez-Campos & Hutchings, 2017)</small>
- Syllis kas <small>(Lucas, Sikorski & San MartÃÂn, 2018)</small>
- Syllis komodoensis <small>(Aguado, San MartÃÂn & ten Hove, 2008)</small>
- Syllis krohnii <small>(Ehlers, 1864)</small>
- Syllis lagunae <small>(Tovar-Hernández, Hernández-Alcántara & SolÃÂs-Weiss, 2008)</small>
- Syllis latifrons <small>(Grube, 1857)</small>
- Syllis licheri <small>(Ravara, San MartÃÂn & Moreira, 2004)</small>
- Syllis limbata <small>(Grube, 1880)</small>
- Syllis longesegmentata <small>(Grube, 1857)</small>
- Syllis lunaris <small>(Imajima, 1966)</small>
- Syllis lutea <small>(Hartmann-Schröder, 1960)</small>
- Syllis luteoides <small>(Hartmann-Schröder, 1962)</small>
- Syllis macroceras <small>(Grube, 1857)</small>
- Syllis macrodentata <small>(Hartmann-Schröder, 1982)</small>
- Syllis magdalena <small>(Wesenberg-Lund, 1962)</small>
- Syllis magnapalpa <small>(Hartmann-Schröder, 1965)</small>
- Syllis marceloi <small>(San MartÃÂn, ÃÂlvarez-Campos & Hutchings, 2017)</small>
- Syllis marugani <small>(Aguado, San MartÃÂn & Nishi, 2006)</small>
- Syllis maryae <small>(San MartÃÂn, 1992)</small>
- Syllis mauretanica <small>(Licher, 1999)</small>
- Syllis mayeri <small>(Musco & Giangrande, 2005)</small>
- Syllis mercedesae <small>(Lucas, San MartÃÂn & Parapar, 2012)</small>
- Syllis mexicana <small>(Rioja, 1960)</small>
- Syllis microoculata <small>(Hartmann-Schröder, 1965)</small>
- Syllis monilaris <small>(Savigny in Lamarck, 1818)</small>
- Syllis mytilorum <small>(Studer, 1889)</small>
- Syllis nigra <small>(Augener, 1925)</small>
- Syllis nigrescens <small>(Grube, 1878)</small>
- Syllis nigricirris <small>(Grube, 1863)</small>
- Syllis nigriscens <small>(Grube, 1878)</small>
- Syllis nigropunctata <small>(Haswell, 1886)</small>
- Syllis notocera <small>(Ehlers, 1905)</small>
- Syllis nuchalis <small>(Hartmann-Schröder, 1960)</small>
- Syllis obscura <small>(Grube, 1857)</small>
- Syllis onkylochaeta <small>(Hartmann-Schröder, 1991)</small>
- Syllis ortizi <small>(San MartÃÂn, 1992)</small>
- Syllis parapari <small>(San MartÃÂn & López, 2000)</small>
- Syllis parturiens <small>(Haswell, 1920)</small>
- Syllis pectinans <small>(Haswell, 1920)</small>
- Syllis picta <small>(Kinberg, 1866)</small>
- Syllis pigmentata <small>(Chamberlin, 1919)</small>
- Syllis pilosa <small>(Aguado, San MartÃÂn & Nishi, 2008)</small>
- Syllis pontxioi <small>(San MartÃÂn & López, 2000)</small>
- Syllis profunda <small>(Cognetti, 1955)</small>
- Syllis prolifera <small>(Krohn, 1852)</small>
- Syllis prolixa <small>(Ehlers, 1901)</small>
- Syllis pseudoarmillaris <small>(Nogueira & San MartÃÂn, 2002)</small>
- Syllis pulvinata <small>(Langerhans, 1881)</small>
- Syllis quadrifasciata <small>(Fischli, 1900)</small>
- Syllis quaternaria <small>(Moore, 1906)</small>
- Syllis ramosa <small>(McIntosh, 1879)</small>
- Syllis riojai <small>(San MartÃÂn, 1990)</small>
- Syllis robertianae <small>(McIntosh, 1885)</small>
- Syllis rosea <small>(Langerhans, 1879)</small>
- Syllis rubicunda <small>(Aguado, San MartÃÂn & Nishi, 2008)</small>
- Syllis rudolphi <small>(Delle Chiaje, 1841)</small>
- Syllis schulzi <small>(Hartmann-Schröder, 1960)</small>
- Syllis sclerolaema <small>(Ehlers, 1901)</small>
- Syllis setoensis <small>(Imajima, 1966)</small>
- Syllis sol <small>(San MartÃÂn, 2004)</small>
- Syllis stenura <small>(Blanchard in Gay, 1849)</small>
- Syllis tamarae <small>(ÃÂlvarez-Campos & Verdes, 2017)</small>
- Syllis tiedemanni <small>(Delle Chiaje, 1841)</small>
- Syllis torquata <small>(Marion & Bobretzky, 1875)</small>
- Syllis tripantu <small>(ÃÂlvarez-Campos & Verdes, 2017)</small>
- Syllis truncata <small>(Haswell, 1920)</small>
- Syllis tyrrhena <small>(Licher & Kuper, 1998)</small>
- Syllis umbricolor <small>(Grube, 1878)</small>
- Syllis unzima <small>(Simon, San MartÃÂn & Robinson, 2014)</small>
- Syllis valida <small>(Grube, 1857)</small>
- Syllis variegata <small>(Grube, 1860)</small>
- Syllis villenai <small>(Aguado, San MartÃÂn & ten Hove, 2008)</small>
- Syllis violacea <small>(Grube, 1870)</small>
- Syllis vittata <small>(Grube, 1840)</small>
- Syllis vivipara <small>(Krohn, 1869)</small>
- Syllis warrnamboolensis <small>(Hartmann-Schröder, 1987)</small>
- Syllis westheidei <small>(San MartÃÂn, 1984)</small>
- Syllis yallingupensis <small>(Hartmann-Schröder, 1982)</small>
- Syllis ypsiloides <small>(Aguado, San MartÃÂn & ten Hove, 2008)</small>
- Ramisyllis
- Ramisyllis multicaudata <small>(Glasby, Schroeder & Aguado, 2012)</small>
- Ramisyllis kingghidorahi <small>(M. Teresa Aguado, et al. 2022)</small>
References