Cochliopidae

Cochliopidae is a family of small freshwater snails with gills and an operculum, aquatic gastropod mollusks.

Paludestrina <small>d'Orbigny, 1840</small> is an archaic synonym, and has been placed on the Official Index of Rejected and Invalid Names by ICZN Opinion 2202.

This family is in the superfamily Truncatelloidea and in the order Littorinimorpha (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005).

Description

Cochliopidae snails are characterized by sharp, elongated spire-like structures, and can be found in rivers or brackish water.

2005 taxonomy

The family Cochliopidae consists of 3 subfamilies (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005):

• Cochliopinae <small>Tryon, 1866</small> - synonyms: Mexithaumatinae <small>D. W. Taylor, 1966</small>, Paludiscalinae <small>D. W. Taylor, 1966</small>
• Littoridininae <small>Thiele, 1928</small>
• Semisalsinae <small>Guiusti & Pezzoli, 1980</small> - synonym: Heleobiini <small>Bernasconi, 1991</small>

Genera

Liu et al. (2001) have recognized 34 genera with more than 260 species within the subfamily Cochliopinae.

Strong et al. (2008) have recognized 246 freshwater species within Cochliopidae.

Genera within the family Cochliopidae include:

subfamily Cochliopinae

• Aphaostracon <small>F. G. Thompson, 1968</small>
• Coahuilix <small>Taylor, 1966</small>
• Cochliopa <small>Stimpson, 1865</small> - type genus of the family Cochliopidae
• Cochliopina <small>Morrison, 1946</small>
• Emmericiella <small>Pilsbry, 1909</small>
• Lobogenes <small>Pilsbry & Bequaert, 1927</small>
• Mexithauma <small>D. W. Taylor, 1966</small>
• Nanivitrea <small>Thiele, 1927</small>
• Subcochliopa <small>J. P. E. Morrison, 1946</small>
• Tepalcatia <small>F. G. Thompson & Hershler, 2002</small>

subfamily Littoridininae

• Antrobia <small>Hubricht, 1971</small> - with the only species Antrobia culveri Hubricht, 1971 - Tumbling Creek cavesnail
• Littoridina <small>Souleyet, 1852</small>
• Pseudocaspia <small>Starobogatov, 1972</small>
• Pseudotryonia <small>Hershler, 2001</small>
• Pseudotryonia adamantina
• Pseudotryonia alamosae
• Pseudotryonia brevissima
• Pseudotryonia grahamae
• Pseudotryonia mica Hershler, Liu & Landye, 2011
• Pseudotryonia pasajae Hershler, Liu & Landye, 2011

subfamily Semisalsinae

• Heleobia <small>Stimpson, 1865</small>
• Heleobops <small>Thompson, 1968</small>
• Paludiscala <small>D. W. Taylor, 1966</small>
• Semisalsa <small>Radoman, 1974</small>

subfamily ? (either Cochliopinae or Littoridininae)

• Aroapyrgus <small>H. B. Baker, 1931</small>
• Balconorbis <small>Hershler & Longley, 1986</small>
• † Carinulorbis <small>Yen, 1949</small>
• Chorrobius <small>Hershler, Liu & Landye, 2011</small>
• Dyris <small>Conrad, 1871</small> - it has extant species and also 26 species in Miocene Pebas Formation
• Eremopyprgus <small>Hershler, 1999</small>
• † Feliconcha <small>Wesselingh, Anderson & Kadolsky, 2006</small> - two species from Miocene of the Pebas Formation
• Feliconcha feliconcha <small>Wesselingh, Anderson & Kadolsky, 2006</small>
• Feliconcha reticulata <small>Wesselingh, Anderson & Kadolsky, 2006</small>
• (probably extant) Glabertryonia Wesselingh, Anderson & Kadolsky, 2006 - three species
• † Glabertryonia glabra <small>Wesselingh, Anderson & Kadolsky, 2006</small> - from Miocene of the Pebas Formation
• † Glabertryonia sp. 1 - from Pliocene of the Las Piedras Formation
• (probably extant) Glabertryonia sp. 2 - from Holocene of Surinam, probably extant
• Juturnia <small>Hershler, Liu & Stockwell, 2002</small>
• Lithococcus <small>Pilsbry, 1911</small>
• Mesobia <small>F. G. Thompson & Hershler, 1991</small>
• Mexipyrgus <small>Taylor, 1966</small>
• Minckleyella <small>Hershler, Liu & Landye, 2011</small> - with the only species Minckleyella balnearis Hershler, Liu & Landye, 2011
• Onobops <small>Thompson, 1968</small>
• Pyrgophorus <small>Ancey, 1888</small>
• Sioliella <small>Haas, 1949</small>
• Spurwinkia <small>Davis, Mazurkiewicz & Mandracchia, 1982</small>
• Texadina <small>Abbott & Ladd, 1951</small>
• Thalassobia <small>Bourguignat in Mabille, 1877</small>

Cladogram

A cladogram based on sequences of mitochondrial cytochrome-c oxidase I (COI) genes showing phylogenic relations of species within Cochliopidae:

References

Further reading

• Hershler R., Davis C. L., Kitting C. L & Liu H.-P. (2007). "Discovery of introduced and cryptogenic cochliopid gastropods in the San Francisco Estuary, California". Journal of Molluscan Studies 73(4): 323-332. PDF.
• Hershler R. & Thompson F. G. (1992). "A Review of the Aquatic Gastropod Subfamily Cochliopinae (Prosobranchia: Hydrobiidae)". Malacological Review, Supplement 5: 1-140. pages 16–18.
• Wilke T., Haase M., Hershler R., Liu H.-P., Misof B. & Ponder W. (2013) Pushing short DNA fragments to the limit: Phylogenetic relationships of ‘hydrobioid’ gastropods (Caenogastropoda: Rissooidea). Molecular Phylogenetics and Evolution, 66: 715-736

External links

• Cochliopidae on WoRMS