Bythinella is a genus of very small freshwater snails, an aquatic gastropod mollusk in the family Bythinellidae (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005).
These gonochoristic snails are small animals, with a shell length of about 2âÂÂ4 mm. Amnicola is a close relative, containing North American snails with essentially similar appearance and ecology. The scientific name means "small Bithynia"; the latter snail genus is somewhat similar at a casual glance (though much larger) but not at all closely related among the Rissooidea.
Distribution
The present genus occurs in springs, and sometimes caves or groundwater, from Catalonia across central Europe to the Aegean Region of Turkey. In and around France a particularly high diversity of these snails is found. Due to the limited range and special and easily destroyed habitat, these snails are liable to become endangered species.
Species
More than 80 species and subspecies are recognized at present. However, it is not at all clear that all of those recognized as good species are actually that distinct, and some in fact seem to be local morphs instead of valid taxa. Some undescribed species are also known to exist.
- Bythinella abbreviata <small>Michaud, 1831</small>
- Bythinella alexpeteri <small>Glöer & Hirschfelder, 2020</small>
- â Bythinella alta <small>Deshayes, 1862</small>
- Bythinella amira <small>Glöer & Hirschfelder, 2020</small>
- Bythinella anatolica <small>Yñldñrñm, Kebapçñ & Bahadñr Koca, 2015</small>
- Bythinella andorrensis <small>Paladilhe, 1875</small>
- Bythinella aneliae <small>Georgiev & Stoycheva, 2011</small>
- Bythinella angelitae <small>Haase, Wilke & Mildner, 2007</small>
- Bythinella angelovi <small>Glöer & Georgiev, 2011</small>
- Bythinella angusta <small>Clessin, 1911</small>
- Bythinella anianensis <small>Paladilhe, 1870</small> - accepted as: B. cebennensis
- â Bythinella antiguensis <small>A. P. Brown & Pilsbry, 1914</small>
- â Bythinella archaea <small>Bourguignat, 1869</small>
- â Bythinella atomus <small>Brongniart, 1810</small>
- Bythinella atypikos <small>A. Reischütz, P. L. Reischütz & W. Fischer, 2008</small>
- â Bythinella auriculata <small>SzÃ
Âts, 1953</small>
- Bythinella austriaca <small>Frauenfeld, 1856</small>
- Bythinella badensis <small>Boeters, 1981</small>: accepted as Bythinella reyniesii
- Bythinella baidashnikovi <small>Sitnikova, Starobogatov & V. Anistratenko, 1992</small>
- Bythinella batalleri <small>Bofill, 1925</small>
- Bythinella baudoni <small>(Paladilhe, 1874)</small>
- Bythinella bavarica <small>Clessin, 1877</small>
- â Bythinella bearnensis <small>Cossmann & Peyrot, 1918</small>
- Bythinella beckmanni <small>A. Reischütz, P. L. Reischütz & W. Fischer, 2008</small>
- Bythinella bertrandi <small>R. Bernasconi, 2000</small>
- Bythinella bicarinata <small>(Des Moulins, 1827)</small>
- Bythinella blidariensis <small>Glöer, 2013</small>
- Bythinella bouldouyrensis <small>Girardi, 2015</small>
- Bythinella bouloti <small>Girardi, Bichain & Wienin, 2002</small>
- Bythinella calimanica <small>Falniowski, Szarowska & Sirbu, 2009</small>
- Bythinella calquierensis <small>Girardi & Boeters, 2015</small>
- Bythinella carcasonis <small>Boeters & Falkner, 2008</small>
- Bythinella carinulata <small>(Drouët, 1867)</small>: accepted as Bythinella viridis <small>(Poiret, 1801)</small>
- Bythinella cebennensis <small>(Dupuy, 1851)</small>
- Bythinella charpentieri <small>(Roth, 1855)</small>
- â Bythinella cirsophora <small>Cossmann, 1888</small>
- Bythinella collingi <small>Boeters, 2009</small>
- Bythinella companyoi <small>(Paladilhe, 1870)</small>
- Bythinella conica <small>Clessin, 1910</small>
- â Bythinella contemta <small>Brusina, 1897</small>
- Bythinella cretensis <small>Schütt, 1980</small>
- Bythinella curta <small>Clessin, 1911</small>
- â Bythinella cyclothyra <small>(Boettger, 1869)</small>
- Bythinella cylindracea <small>(Paladilhe, 1869)</small>
- Bythinella cylindrica <small>(Frauenfeld, 1857)</small>
- Bythinella dacica <small>Grossu, 1946</small>
- â Bythinella dalmatica <small>Brusina, 1897</small>
- â Bythinella damintunensis <small>Youluo, 1978</small>
- Bythinella darrieuxii <small>(de Folin & Bérillon, 1877)</small>
- Bythinella dedovi <small>Glöer & Georgiev, 2011</small>
- Bythinella deidamiae <small>Glöer & Porfyris, 2022</small>
- â Bythinella deshayesiana <small>(Bourguignat, 1869)</small>
- â Bythinella desnoyersi <small>(Bourguignat, 1869)</small>
- Bythinella dierckingi <small>Glöer & Georgiev, 2011</small>
- Bythinella dispersa <small>Radoman, 1976</small>
- Bythinella drimica <small>Radoman, 1976</small>
- Bythinella dromensis <small>Boeters & Falkner, 2008</small>
- â Bythinella dumenisliana <small>(Bourguignat, 1869)</small>
- â Bythinella edwardsiana <small>(Bourguignat, 1869)</small>
- Bythinella ehrmanni <small>Pax, 1938</small>
- Bythinella elenae <small>Glöer & Georgiev, 2011</small>
- Bythinella eleousae <small>Glöer & Hirschfelder, 2020</small>
- Bythinella ellinikae <small>Glöer & Hirschfelder, 2020</small>
- â Bythinella elongata <small>Roshka, 1973</small>
- Bythinella espanoli <small>Bech, 1980</small>
- Bythinella etrusca <small>(Paladilhe, 1867)</small>
- â Bythinella eugenii <small>(Deshayes, 1862)</small>
- Bythinella eurystoma <small>(Paladilhe, 1870)</small>
- Bythinella eutrepha <small>(Paladilhe, 1867)</small>
- â Bythinella expulsa <small>(Deshayes, 1862)</small>
- â Bythinella eysdenensis <small>ÃÂ. Vincent, 1930</small>
- Bythinella falniowskii <small>Glöer, 2013</small>
- Bythinella feheri <small>Glöer, 2013</small>
- Bythinella ferussina <small>(Des Moulins, 1827)</small>
- Bythinella friderici <small>Boeters & Falkner, 2008</small>
- Bythinella fuscata <small>Brancsik, 1889</small>
- Bythinella galerae <small>Girardi, Bichain & Wienin, 2002</small>: accepted as Bythinella bouloti <small>H. Girardi, Bichain & Wienin, 2002</small>
- Bythinella geisserti <small>Boeters & Falkner, 2003</small>
- Bythinella georgievi <small>Glöer, 2013</small>
- Bythinella gibbosa <small>(Moquin-Tandon, 1856)</small>
- Bythinella gloeeri <small>Georgiev, 2009</small>
- Bythinella gokceadaensis <small>D. A. OdabaÃ
Âñ, 2019</small>
- Bythinella golemoensis <small>Glöer & Mrkvicka, 2015</small>
- â Bythinella gracillima <small>SzÃ
Âts, 1953</small>
- Bythinella gregoi <small>Glöer & ErÃ
Âss, 2015</small>
- Bythinella grossui <small>Falniowski, Szarowska & Sirbu, 2009</small>
- Bythinella guranensis <small>(Paladilhe, 1870)</small>: accepted as Bythinella rubiginosa <small>(Boubée, 1833)</small>
- â Bythinella hanjiangensis <small>W. Yü, 1977</small>
- Bythinella hansboetersi <small>Glöer & PeÃ
¡iÃÂ, 2006</small>
- Bythinella hasanspahici <small>Mulaomerovià& Glöer, 2022</small>
- Bythinella heydeni <small>Clessin, 1879</small>
- Bythinella heynemanniana <small>Hazay, 1881</small>
- Bythinella hungarica <small>Hazay, 1880</small>
- â Bythinella intermedia <small>(Melleville, 1843)</small>
- Bythinella istanbulensis <small>Yñldñrñm, Kebapçñ & Yüce, 2015</small>
- Bythinella istoka <small>Glöer & PeÃ
¡iÃÂ, 2014</small>
- Bythinella izvorica <small>Glöer & Georgiev, 2011</small>
- Bythinella jobae <small>(Bourguignat, 1862)</small>
- Bythinella jodevidtsi <small>Bichain & Bertrand, 2022</small>
- Bythinella joinvillensis <small>(Bourguignat, 1869)</small>
- Bythinella jourdei <small>R. Bernasconi, 2000</small>
- Bythinella jozefgregoi <small>Glöer & Reuselaars, 2020</small>
- Bythinella kambosensis <small>Glöer & Hirschfelder, 2020</small>
- Bythinella kapelana <small>Radoman, 1976</small>
- Bythinella kastaliae <small>Glöer & Hirschfelder, 2020</small>
- Bythinella kastanolongosensis <small>Glöer & PeÃ
¡iÃÂ, 2020</small>
- Bythinella kazdaghensis <small>OdabaÃ
Âi & Georgiev, 2014</small>
- Bythinella kleptuzica <small>Glöer & Georgiev, 2011</small>
- Bythinella klimaensis <small>Glöer & Reuselaars, 2020</small>
- Bythinella konstadinensis <small>Glöer & Reuselaars, 2020</small>
- Bythinella kosensis <small>Schütt, 1980</small>
- Bythinella kosmosi <small>Clessin, 1910</small>
- Bythinella kwanti <small>Glöer & Reuselaars, 2020</small>
- Bythinella kyriaki <small>Glöer & Reuselaars, 2020</small>
- Bythinella lancevelei <small>Locard, 1894</small> accepted as Bythinella viridis <small>(Poiret, 1801)</small>
- â Bythinella lartetiana <small>(Bourguignat, 1869)</small>
- â Bythinella lecongensis <small>W. Yü & X.-Q. Zhang, 1982</small>
- Bythinella liandinaensis <small>Glöer & Reuselaars, 2020</small>
- Bythinella ligurica <small>(Paladilhe, 1867)</small>
- Bythinella limnopsis <small>Letourneux & Bourguignat, 1887</small>
- Bythinella longula <small>Brancsik, 1889</small>
- Bythinella lunzensis <small>Boeters, 2008</small>
- Bythinella luteola <small>Radoman, 1976</small>
- Bythinella magdalenae <small>Yñldñrñm, Kebapçñ & Bahadñr Koca, 2015</small>
- Bythinella magna <small>Radoman, 1976</small>
- Bythinella major <small>(Pascal, 1873)</small>
- Bythinella margritae <small>Glöer & Georgiev, 2011</small>
- Bythinella marianramosae <small>Falniowski, JaszczyÃ
Âska & Hofman, 2023</small>
- Bythinella marici <small>Glöer & PeÃ
¡iÃÂ, 2014</small>
- Bythinella markovi <small>Glöer & Georgiev, 2009</small>
- Bythinella mauritanica <small>Letourneux & Bourguignat, 1887</small>
- â Bythinella megarensis <small>Bukowski, 1896</small>
- Bythinella melovskii <small>Glöer & Slavevska-StamenkoviÃÂ, 2015</small>
- Bythinella metarubra <small>Falniowski, 1987</small>
- Bythinella micherdzinskii <small>Falniowski, 1980</small>
- Bythinella microcochlia <small>Letourneux & Bourguignat, 1887</small>
- â Bythinella minuata <small>(Deshayes, 1862)</small>
- Bythinella molcsanyi <small>H. Wagner, 1941</small>
- Bythinella moulinsii <small>(Dupuy, 1849)</small>
- Bythinella muranyii <small>Glöer & ErÃ
Âss, 2015</small>
- Bythinella navacellensis <small>Prié & Bichain, 2009</small>
- Bythinella nonveilleri <small>Glöer, 2008</small>
- Bythinella normalis <small>Courty, 1907</small>
- Bythinella nothites <small>(Westerlund, 1886)</small>
- Bythinella occasiuncula <small>Boeters & Falkner, 2001</small>
- Bythinella olymbosensis <small>Glöer & Reuselaars, 2020</small>
- Bythinella opaca <small>(M. von Gallenstein, 1848)</small>
- Bythinella padana <small>R. Bernasconi, 1989</small>
- Bythinella padiraci <small>Locard, 1903</small>
- Bythinella pannonica <small>(Frauenfeld, 1865)</small>
- â Bythinella parkinsoni <small>(J. Morris, 1854)</small>
- Bythinella parvula <small>Locard, 1893</small>: accepted as Bythinella reyniesii <small>(Dupuy, 1851)</small>
- Bythinella perilongata <small>Altimira, 1959</small>
- Bythinella perivoliensis <small>Glöer & Reuselaars, 2020</small>
- Bythinella pesici <small>Glöer & Reuselaars, 2020</small>
- Bythinella pesterica <small>Glöer, 2008</small>
- Bythinella petrosensis <small>Glöer, 2008</small>
- Bythinella pujolensis <small>R. Bernasconi, 2000</small>: accepted as Bythinella bicarinata <small>(Des Moulins, 1827)</small>
- â Bythinella pulcherrima <small> SzÃ
Âts, 1953</small>
- â Bythinella pulchra <small>(Deshayes, 1862)</small>
- Bythinella punica <small>Letourneux & Bourguignat, 1887</small>
- â Bythinella pupina <small>(Deshayes, 1862)</small>
- Bythinella pupoides <small>(Paladilhe, 1869)</small>
- Bythinella pyrenaica <small>(Bourguignat, 1861)</small>
- Bythinella rachonica <small>Georgiev & Glöer, 2020</small>
- Bythinella radomani <small>Falniowski, Szarowska & Sirbu, 2009</small>
- Bythinella ravnogorica <small>Glöer & Georgiev, 2009</small>
- Bythinella reischuetzi <small>Glöer & Georgiev, 2020</small>
- Bythinella rethymnonensis <small>Glöer & Hirschfelder, 2020</small>
- Bythinella reuselaarsi <small>Glöer & PeÃ
¡iÃÂ, 2020</small>
- Bythinella reyniesii <small>(Dupuy, 1851)</small>
- â Bythinella rhodanica <small>Roman, 1912</small>
- Bythinella rhodopensis <small>Glöer & Georgiev, 2011</small>
- Bythinella righaensis <small>Glöer & Reuselaars, 2020</small>
- Bythinella rilaensis <small>Georgiev & Glöer, 2013</small>
- â Bythinella rimulifera <small>Stache, 1889</small>
- Bythinella robiciana
- Bythinella robusta <small>Z.-B. Kang, 1986</small>
- Bythinella rolani <small>Boeters, 2019</small>
- Bythinella rondelaudi <small>R. Bernasconi, 2000</small>
- â Bythinella rothi <small>(Brusina, 1884)</small>
- Bythinella roubionensis <small>Caziot, 1910</small>
- Bythinella rouchi <small>Boeters & Falkner, 2008</small>
- Bythinella rouveyrolensis <small>Girardi, 2015</small>
- Bythinella rubiginosa <small>(Boubee, 1833)</small>
- Bythinella reyniesii <small>(Dupuy, 1851)</small>
- Bythinella samecana <small>Clessin, 1911</small>
- â Bythinella scalaris <small>(SlavÃÂk, 1869)</small>
- Bythinella schmidtii <small>(Küster, 1852)</small>: accepted as Bythinella opaca <small>(M. von Gallenstein, 1848)</small>
- â Bythinella scitula <small>Brusina, 1892</small>
- Bythinella serborientalis <small>Radoman, 1976</small>
- Bythinella servainiana (<small>Paladilhe, 1870)</small>
- Bythinella sijei <small>Mulaomerovià& Glöer, 2022</small>
- â Bythinella sinensis <small>W. Yü & H.-Z. Pan, 1982</small>
- Bythinella sirbui <small>Glöer, 2013</small>
- Bythinella sitiensis <small>Glöer & Georgiev, 2011</small>
- Bythinella slaveyae <small>Glöer & Georgiev, 2011</small>
- Bythinella smolyanica <small>Glöer & Georgiev, 2011</small>
- Bythinella solidula <small>Brancsik, 1889</small>
- Bythinella sordida <small>(Küster, 1853)</small>
- â Bythinella sphaeroidalis <small>Cossmann, 1888</small>
- Bythinella srednogorica <small>Glöer & Georgiev, 2009</small>
- Bythinella steffeki <small>Grego & Glöer, 2019</small>
- Bythinella stoychevae <small>Georgiev, 2011</small>
- Bythinella subovata <small>(Paladilhe, 1876)</small>
- Bythinella syngenes <small>Preston, 1914</small>
- Bythinella syntriculus <small>Boeters & Falkner, 2008</small>
- Bythinella szarowskae <small>Glöer, 2013</small>
- Bythinella tabakovaensis <small>Mulaomerovià& Glöer, 2023</small>
- Bythinella tajoensis <small>Boeters, 2019</small>
- Bythinella taraensis <small>Glöer & PeÃ
¡iÃÂ, 2010</small>
- Bythinella taygetensis <small>Glöer & Hirschfelder, 2020</small>
- Bythinella tejedoi <small>Boeters, 2019</small>
- Bythinella temelkovi <small>Georgiev & Glöer, 2014</small>
- Bythinella thermophila <small>Glöer, Varga & Mrkvicka, 2015</small>
- Bythinella tornensis <small>Hazay, 1881</small>
- Bythinella troyana <small>R. Bernasconi, 2000</small>
- Bythinella truncata <small>Courty, 1907</small>
- Bythinella tumidula <small>Clessin, 1910</small>
- Bythinella turca <small>(Radoman, 1976)</small>
- Bythinella turriculata <small>(Paladilhe, 1869)</small>
- Bythinella ullaensis <small>Boeters & Falkner, 2008</small>
- Bythinella utriculus <small>(Paladilhe, 1874)</small>
- Bythinella valkanovi <small>Glöer & Georgiev, 2011</small>
- Bythinella vesontiana <small>R. Bernasconi, 1989</small>
- Bythinella vidinovae <small>Dedov, Taseva & Georgiev, 2021</small>
- Bythinella vimperei <small>R. Bernasconi, 2000</small>
- Bythinella viridis <small>(Poiret, 1801)</small>
- Bythinella viseuiana <small>Falniowski, Szarowska & Sirbu, 2009</small>
- â Bythinella vitrellaeformis <small>Lörenthey, 1902</small>
- Bythinella walensae <small>Falniowski, Hofman & Rysiewska, 2016</small>
- Bythinella walkeri <small>Glöer & Georgiev, 2009</small>
- Bythinella wawrzineki <small>R. Bernasconi, 2002</small>
- Bythinella westerlundi <small>Glöer, 2022</small>
- Bythinella wilkei <small>Yñldñrñm, Kebapçñ & Bahadñr Koca, 2015</small>
- Bythinella yerlii <small>Gürlek, 2017</small>
- Bythinella zyvionteki <small>Falniowski, 1987</small>
<big>Relocated species:</big>
- â Bythinella chinensis <small>Y.-Y. Liu & W.-Z. Zhang, 1979</small> accepted as â Chencuia chinensis <small>Y.-Y. Liu & W.-Z. Zhang, 1979</small> '
- Bythinella cvijici <small>PavloviÃÂ, 1933</small> accepted as Prososthenia cvijici <small>PavloviÃÂ, 1933</small>
- â Bythinella falloti <small>Degrange-Touzin, 1892</small> accepted as â Stenothyroides falloti <small>(Degrange-Touzin, 1892)</small>
- Bythinella nipponica <small>S. Mori, 1937</small> accepted as Moria nipponica <small>(S. Mori, 1937)</small>
- Bythinella wantanensis <small>Z.-B. Kang, 1983</small> accepted as Erhaia wantanensis <small>(Z.-B. Kang, 1983)</small>
Footnotes
References
- (2007): A gleam in the dark: Phylogenetic species delimitation in the confusing spring-snail genus Bythinella Moquin-Tandon, 1856 (Gastropoda: Rissooidea: Amnicolidae). Mol. Phylogenet. Evol. 45(3): 927âÂÂ941. <small></small> PDF fulltext