Bulimulidae

Bulimulidae is a taxonomic family of medium-sized to large, air-breathing, tropical and sub-tropical land snails, terrestrial pulmonate gastropod molluscs in the superfamily Orthalicoidea.

Distribution

Distribution of species in the family Bulimulidae includes Ecuador (9 genera) and other South American countries. Some species also occur in North America.

Fossil record

The family's oldest fossil record dates from the late Cretaceous of Brazil (ItaboraÃÂ Basin).

Anatomy

Members of this family have a haploid chromosome number between 26 and 30 (according to the values in this table).

Shells of species within Bostrycinae have a smooth protoconch and the genital organs feature a relatively long penis sheath (ca. 1/4Ã¢ÂÂ1/6 total phallus length).

Taxonomy

Previously, the members of the Orthalicidae were also included in this family, as the subfamily Orthalicinae, and the taxa listed here were placed in their own subfamily, the Bulimulinae.

2005 taxonomy

Bulimulinae was placed in the family Orthalicidae according to the taxonomy of the Gastropoda (Bouchet & Rocroi, 2005).

The subfamily Bulimulinae included 3 tribes:

Tribe Bulimulini Tryon, 1867 Ã¢ÂÂ synonyms: Bulimidae Guilding, 1828 (inv.); Berendtiinae P. Fischer & Crosse, 1872; Bothiembryontidae Iredale, 1937
Tribe Odontostomini Pilsbry & Vanatta, 1898 Ã¢ÂÂ synonym: Tomogeridae Jousseaume, 1877
Tribe Simpulopsini Schileyko, 1999

2010 taxonomy

Breure et al. (2010) moved Bothriembryon to Placostylidae (since 2012 as Bothriembryontidae), elevated Bulimulinae to Bulimulidae and Odontostomini to Odontostomidae.

2012 taxonomy

Breure & Romero (2012) confirmed previous results from 2010, additionally they elevated Simpulopsini to Simpulopsidae. There are three subfamilies within Bulimulidae:

subfamily Bulimulinae Tryon, 1867
subfamily Peltellinae Gray, 1855
subfamily Bostrycinae Breure, 2012

Genera

Genera in the family Bulimulidae include:

subfamily Bulimulinae Tryon, 1887: (synonym: Berendtiinae P. Fischer & Crosse, 1872)

Anctus E. von Martens, 1860
Auris Spix, 1827
Berendtia Crosse & P. Fischer, 1869
Bocourtia Rochebrune, 1882
Bulimulus Leach, 1814 Ã¢ÂÂ type genus of the subfamily Bulimulinae
Cochlorina Jan, 1830
Graptostracus Pilsbry, 1939
Kora Simone, 2012
Lopesianus Weyrauch, 1958
Naesiotus Albers, 1850
Neopetraeus E. von Martens, 1885
Newboldius Pilsbry, 1932
Otostomus H. Beck, 1837
Oxychona MÃÂ¶rch, 1852
Protoglyptus Pilsbry, 1897
Pseudoxychona Pilsbry, 1930
Rabdotus Albers, 1850
Sanniostracus Salvador, F. S. Silva & Cavallari, 2023
Scutalus Albers, 1850
Spartocentrum Dall, 1895
Sphaeroconcha Breure, 1978
Stenostylus Pilsbry, 1898
Suniellus Breure, 1978

subfamily Peltellinae

Antidrymaeus Germain, 1907
Drymaeus Albers, 1850
Peltella Gray, 1855 Ã¢ÂÂ type genus of the subfamily Peltellinae
Mesembrinus Albers, 1850

subfamily Bostrycinae

Bostryx Troschel, 1847 sensu lato- type genus of the subfamily Bostrycinae

Unassigned within Bulimulidae

Bulimus BruguiÃÂ¨re, 1789 (temporary name)
Itaborahia Maury, 1935
Ã¢ÂÂ Oreoconus D. W. Taylor in McKenna et al., 1962
Ã¢ÂÂ Palaeobulimulus Parodiz, 1949
Stapafurdius Simone, 2021
Ã¢ÂÂ Tocobaga Auffenberg, Slapcinsky & Portell, 2015

Synonyms

Adzharia P. Hesse, 1933: synonym of Bulimulus Leach, 1814 (junior synonym)
Antidrymaeus Germain, 1907: synonym of Drymaeus (Mesembrinus) Albers, 1850 represented as Drymaeus Albers, 1850
Atahualpa Strebel, 1910: synonym of Thaumastus E. von Martens, 1860
Ataxellus Dall, 1912: synonym of Bostryx (Geopyrgus) Pilsbry, 1896 represented asBostryx Troschel, 1847
Ataxus Albers, 1850: synonym of Bostryx Troschel, 1847
Bilamelliferus Weyrauch, 1958: synonym of Bostryx Troschel, 1847 (junior synonymy)
Diaphanomormus Weyrauch, 1964: synonym of Drymaeus (Mesembrinus) Albers, 1850 represented as Drymaeus Albers, 1850
Elatibostryx Weyrauch, 1958: synonym of Bostryx Troschel, 1847
Floreziellus Weyrauch, 1967: synonym of Bostryx Troschel, 1847 (junior synonymy)
Geoceras Pilsbry, 1896: synonym of Bostryx (Geopyrgus) Pilsbry, 1896 represented as Bostryx Troschel, 1847
Globulinus Crosse & P. Fischer, 1875: synonym of Rabdotus Albers, 1850
Goniognathmus Crosse & P. Fischer, 1875: synonym of Drymaeus Albers, 1850 (junior synonym)
Hamadryas Albers, 1850: synonym of Drymaeus Albers, 1850
Hannarabdotus Emerson & Jacobson, 1964: synonym of Rabdotus Albers, 1850
Kionoptyx Haas, 1966: synonym of Bostryx Troschel, 1847 (junior synonymy)
Kuschelenia Hylton Scott, 1951: synonym of Bocourtia (Kuschelenia) Hylton Scott, 1951 represented as Bocourtia Rochebrune, 1882
Leptodrymaeus Pilsbry, 1946: synonym of Drymaeus (Mesembrinus) Albers, 1850 represented as Drymaeus Albers, 1850
Leptomerus Albers, 1850: synonym of Bulimulus Leach, 1814
Leptomormus Weyrauch, 1958: synonym of Drymaeus (Mesembrinus) Albers, 1850 represented as Drymaeus Albers, 1850
Liostracus [sic]: synonym of Leiostracus Albers, 1850 (misspelling of original genus, Leiostracus Albers, 1850)
Loboa Ihering, 1917: synonym of Bulimulus Leach, 1814
Mormus E. von Martens, 1860: synonym of Drymaeus Albers, 1850
Multifasciatus Weyrauch, 1958: synonym of Bostryx (Geopyrgus) Pilsbry, 1896 represented as Bostryx Troschel, 1847
Naesiotellus Weyrauch, 1967: synonym of Bostryx Troschel, 1847 (junior synonym)
Navicula Spix, 1827: synonym of Cochlorina Jan, 1830
Nesiotes E. von Martens, 1860: synonym of Naesiotus Albers, 1850
Obstrussus Parodiz, 1946: synonym of Naesiotus Albers, 1850
Olinodia <big>Dall, 1920</big>: synonym of Naesiotus (Reclasta) Dall, 1920 represented as Naesiotus Albers, 1850
Orphaicus Schaufuss, 1869: synonym of Thaumastus E. von Martens, 1860
Orphnus Albers, 1850: synonym of Thaumastus E. von Martens, 1860
Orthotomium Crosse & P. Fischer, 1875: synonym of Rabdotus Albers, 1850
Pachyotus H. Beck, 1837: synonym of Auris Spix, 1827
Pachytholus Strebel, 1909: synonym of Thaumastus E. von Martens, 1860
Pampasinus Weyrauch, 1958: synonym of Bostryx (Platybostryx) Pilsbry, 1896: synonym ofBostryx Troschel, 1847
Peronaeus Albers, 1850: synonym of Bostryx Troschel, 1847
Phenacotaxus Dall, 1912: synonym of Bostryx (Geopyrgus) Pilsbry, 1896 represented as Bostryx Troschel, 1847
Plecochilus Agassiz, 1846: synonym of Plekocheilus Guilding, 1827 (unjustified emendation)
Pseudoperonaeus Weyrauch, 1958: synonym of Bostryx (Geopyrgus) Pilsbry, 1896 represented as Bostryx Troschel, 1847
Pseudorhodea Dall, 1895: synonym of Rabdotus (Plicolumna) J.G. Cooper, 1895 represented as Rabdotus Albers, 1850
Pyrgus Albers, 1850: synonym of Bostryx (Geopyrgus) Pilsbry, 1896 represented as Bostryx Troschel, 1847
Rhabdotus [sic]: synonym of Rabdotus Albers, 1850 (incorrect subsequent spelling)
Rhaphiellus L. Pfeiffer, 1856: synonym of Naesiotus Albers, 1850
Rimatula Parodiz, 1946: synonym of Protoglyptus Pilsbry, 1897
Saeronia Dall, 1920: synonym of Naesiotus Albers, 1850
Semiclausaria L. Pfeiffer, 1856: synonym of Drymaeus Albers, 1850 (junior synonym)
Siphalomphix Rafinesque, 1833: synonym of Bulimulus Leach, 1814
Sonorina Pilsbry, 1896: synonym of Rabdotus (Leptobyrsus) Crosse & P. Fischer, 1875 represented as Rabdotus Albers, 1850
Spiroscutalus Pilsbry, 1932: synonym of Scutalus Albers, 1850
Stenostoma Spix, 1827: synonym of Anctus E. von Martens, 1860
Tatutor Jousseaume, 1887: synonym of Thaumastus E. von Martens, 1860
Teneritia J. Mabille, 1898: synonym of Spartocentrum Dall, 1895
Tholus Strebel, 1909: synonym of Thaumastus E. von Martens, 1860
Xenothauma Fulton 1896: synonym of Scutalus Albers, 1850
Zaplagius Pilsbry, 1896: synonym of Cochlorina Jan, 1830

References

This article incorporates CC-BY-3.0 text from the reference

Bouchet P., Rocroi J.P., Hausdorf B., Kaim A., Kano Y., NÃÂ¼tzel A., Parkhaev P., SchrÃÂ¶dl M. & Strong E.E. (2017). Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia. 61(1-2): 1-526