The APG IV system of flowering plant classification is the fourth version of a modern, mostly molecular-based, system of plant taxonomy for flowering plants (angiosperms) being developed by the Angiosperm Phylogeny Group (APG). It was published in 2016, seven years after its predecessor the APG III system was published in 2009, and 18 years after the first APG system was published in 1998. In 2009, a linear arrangement of the system was published separately; the APG IV paper includes such an arrangement, cross-referenced to the 2009 one.
Compared to the APG III system, the APG IV system recognizes five new orders (Boraginales, Dilleniales, Icacinales, Metteniusales and Vahliales), along with some new families, making a total of 64 angiosperm orders and 416 families. In general, the authors describe their philosophy as "conservative", based on making changes from APG III only where "a well-supported need" has been demonstrated. This has sometimes resulted in placements that are not compatible with published studies, but where further research is needed before the classification can be changed.
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Detailed version
Key to symbols used:
<nowiki>*</nowiki> = the family has been added or its circumscription changed since the APG III system of 2009
â = the order has been added since the APG III system
- Canellales <small>Cronq.</small>
- Canellaceae <small>Mart., nom. cons.</small>
- Winteraceae <small>R.Br. ex Lindl., nom. cons.</small>
- Piperales <small>Bercht. & J.Presl</small>
- Saururaceae <small>Rich. ex T.Lestib., nom. cons.</small>
- Piperaceae <small>Giseke, nom. cons.</small>
- Aristolochiaceae <small>Juss., nom. cons.</small> * (including Asaraceae <small>Vent.</small>, Hydnoraceae <small>C.Agardh, nom. cons.</small>, Lactoridaceae <small>Engl., nom. cons.</small>)
- Magnoliales <small>Juss. ex Bercht. & J.Presl</small>
- Myristicaceae <small>R.Br., nom. cons.</small>
- Magnoliaceae <small>Juss., nom. cons.</small>
- Degeneriaceae <small>I.W.Bailey & A.C.Sm., nom. cons.</small>
- Himantandraceae <small>Diels, nom. cons.</small>
- Eupomatiaceae <small>Orb., nom. cons.</small>
- Annonaceae <small>Juss., nom. cons.</small>
- Laurales <small>Juss. ex Bercht. & J.Presl</small>
- Calycanthaceae <small>Lindl., nom. cons.</small>
- Siparunaceae <small>Schodde</small>
- Gomortegaceae <small>Reiche, nom. cons.</small>
- Atherospermataceae <small>R.Br.</small>
- Hernandiaceae <small>Blume, nom. cons.</small>
- Monimiaceae <small>Juss., nom. cons.</small>
- Lauraceae <small>Juss., nom. cons.</small>
Independent lineage: unplaced to more inclusive clade
- Acorales <small>Mart.</small>
- Acoraceae <small>Martinov</small>
- Alismatales <small>R.Br. ex Bercht. & J.Presl</small>
- Araceae <small>Juss., nom. cons.</small>
- Tofieldiaceae <small>Takht.</small>
- Alismataceae <small>Vent., nom. cons.</small>
- Butomaceae <small>Mirb., nom. cons.</small>
- Hydrocharitaceae <small>Juss., nom. cons.</small>
- Scheuchzeriaceae <small>F.Rudolphi, nom. cons.</small>
- Aponogetonaceae <small>Planch., nom. cons.</small>
- Juncaginaceae <small>Rich., nom. cons.</small> *
- Maundiaceae <small>Nakai</small> *
- Zosteraceae <small>Dumort., nom. cons.</small>
- Potamogetonaceae <small>Bercht. & J.Presl, nom. cons</small>
- Posidoniaceae <small>Vines, nom. cons.</small>
- Ruppiaceae <small>Horan., nom. cons.</small>
- Cymodoceaceae <small>Vines, nom. cons.</small>
- Petrosaviales <small>Takht.</small>
- Petrosaviaceae <small>Hutch., nom. cons.</small>
- Dioscoreales <small>Mart.</small>
- Nartheciaceae <small>Fr. ex Bjurzon</small>
- Burmanniaceae <small>Blume, nom. cons.</small> (not monophyletic.)
- Dioscoreaceae <small>R.Br., nom. cons.</small>
- Pandanales <small>R.Br. ex Bercht. & J.Presl</small>
- Triuridaceae <small>Gardner, nom. cons.</small>
- Velloziaceae <small>J.Agardh, nom. cons.</small>
- Stemonaceae <small>Caruel, nom. cons.</small>
- Cyclanthaceae <small>Poit. ex A.Rich., nom. cons.</small>
- Pandanaceae <small>R.Br., nom. cons.</small>
- Liliales <small>Perleb</small>
- Campynemataceae <small>Dumort.</small>
- Corsiaceae <small>Becc., nom. cons.</small>
- Melanthiaceae <small>Batsch ex Borkh., nom. cons.</small>
- Petermanniaceae <small>Hutch, nom. cons.</small>
- Alstroemeriaceae <small>Dumort., nom. cons.</small>
- Colchicaceae <small>DC., nom. cons.</small>
- Philesiaceae <small>Dumort., nom. cons.</small>
- Ripogonaceae <small>Conran & Clifford</small>
- Smilacaceae <small>Vent., nom. cons.</small>
- Liliaceae <small>Juss., nom. cons.</small>
- Asparagales <small>Link</small>
- Orchidaceae <small>Juss., nom. cons.</small>
- Boryaceae <small>M.W.Chase et al.</small>
- Blandfordiaceae <small>R.Dahlgren & Clifford</small>
- Asteliaceae <small>Dumort.</small>
- Lanariaceae <small>H.Huber ex R.Dahlgren</small>
- Hypoxidaceae <small>R.Br., nom. cons.</small>
- Doryanthaceae <small>R.Dahlgren & Clifford</small>
- Ixioliriaceae <small>Nakai</small> (as 'Ixiolirionaceae'; spelling corrected)
- Tecophilaeaceae <small>Leyb., nom. cons.</small>
- Iridaceae <small>Juss., nom. cons.</small>
- Xeronemataceae <small>M.W.Chase et al.</small>
- Asphodelaceae <small>Juss., nom. cons. prop.</small> (including Xanthorrhoeaceae <small>Dumort., nom. cons.</small>)
- Amaryllidaceae <small>J.St.-Hil., nom. cons.</small>
- Asparagaceae <small>Juss., nom. cons.</small>
- Arecales <small>Bromhead</small>
- Dasypogonaceae <small>Dumort.</small>
- Arecaceae <small>Bercht. & J.Presl, nom. cons.</small> (=Palmae <small>Juss., nom. cons.</small>)
- Commelinales <small>Mirb. ex Bercht. & J.Presl</small>
- Hanguanaceae <small>Airy Shaw</small>
- Commelinaceae <small>Mirb., nom. cons.</small>
- Philydraceae <small>Link, nom. cons.</small>
- Pontederiaceae <small>Kunth, nom. cons.</small>
- Haemodoraceae <small>R.Br., nom. cons.</small>
- Zingiberales <small>Griseb.</small>
- Strelitziaceae <small>Hutch., nom. cons.</small>
- Lowiaceae <small>Ridl., nom. cons.</small>
- Heliconiaceae <small>Vines</small>
- Musaceae <small>Juss., nom. cons.</small>
- Cannaceae <small>Juss., nom. cons.</small>
- Marantaceae <small>R.Br., nom. cons.</small>
- Costaceae <small>Nakai</small>
- Zingiberaceae <small>Martinov, nom. cons.</small>
- Poales <small>Small</small>
- Typhaceae <small>Juss., nom. cons.</small>
- Bromeliaceae <small>Juss., nom. cons.</small>
- Rapateaceae <small>Dumort., nom. cons.</small>
- Xyridaceae <small>C.Agardh, nom. cons.</small>
- Eriocaulaceae <small>Martinov, nom. cons.</small>
- Mayacaceae <small>Kunth, nom. cons.</small>
- Thurniaceae <small>Engl., nom. cons.</small>
- Juncaceae <small>Juss., nom. cons.</small>
- Cyperaceae <small>Juss., nom. cons.</small>
- Restionaceae <small>R.Br., nom. cons.</small> * (including Anarthriaceae <small>D.W.Cutler & Airy Shaw</small>, Centrolepidaceae <small>Endl., nom. cons.</small>)
- Flagellariaceae <small>Dumort., nom. cons.</small>
- Joinvilleaceae <small>Toml. & A.C.Sm.</small>
- Ecdeiocoleaceae <small>D.W.Cutler & Airy Shaw</small>
- Poaceae <small>Barnhart, nom. cons.</small> (= Gramineae <small>Juss., nom. cons.</small>)
Probable sister of eudicots
- Ranunculales <small>Juss. ex Bercht. & J.Presl</small>
- Eupteleaceae <small>K.Wilh., nom. cons.</small>
- Papaveraceae <small>Juss., nom. cons.</small>
- Circaeasteraceae <small>Hutch., nom. cons.</small>
- Lardizabalaceae <small>R.Br., nom. cons.</small>
- Menispermaceae <small>Juss., nom. cons.</small>
- Berberidaceae <small>Juss., nom. cons.</small>
- Ranunculaceae <small>Juss., nom. cons.</small>
- Proteales <small>Juss. ex Bercht. & J.Presl</small>
- Sabiaceae <small>Blume, nom. cons.</small>
- Nelumbonaceae <small>A.Rich., nom. cons.</small>
- Platanaceae <small>T.Lestib., nom. cons.</small>
- Proteaceae <small>Juss., nom. cons.</small>
- Trochodendrales <small>Takht. ex Cronq.</small>
- Trochodendraceae <small>Eichler, nom. cons.</small>
- Buxales <small>Takht. ex Reveal</small>
- Buxaceae <small>Dumort., nom. cons.</small> * (including Haptanthaceae <small>C.Nelson</small>)
- Saxifragales <small>Bercht. & J.Presl</small>
- Peridiscaceae <small>Kuhlm., nom. cons.</small>
- Paeoniaceae <small>Raf., nom. cons.</small>
- Altingiaceae <small>Lindl., nom. cons.</small>
- Hamamelidaceae <small>R.Br., nom. cons.</small>
- Cercidiphyllaceae <small>Engl., nom. cons.</small>
- Daphniphyllaceae <small>Müll.Arg., nom. cons.</small>
- Iteaceae <small>J.Agardh, nom. cons.</small>
- Grossulariaceae <small>DC., nom. cons.</small>
- Saxifragaceae <small>Juss., nom. cons.</small>
- Crassulaceae <small>J.St.-Hil., nom. cons.</small>
- Aphanopetalaceae <small>Doweld</small>
- Tetracarpaeaceae <small>Nakai</small>
- Penthoraceae <small>Rydb. ex Britton, nom. cons.</small>
- Haloragaceae <small>R.Br., nom. cons.</small>
- Cynomoriaceae <small>Endl. ex Lindl., nom. cons.</small>
- Vitales <small>Juss. ex Bercht. & J.Presl</small>
- Vitaceae <small>Juss., nom. cons.</small>
Fabids
- Zygophyllales <small>Link</small>
- Krameriaceae <small>Dumort., nom. cons.</small>
- Zygophyllaceae <small>R.Br., nom. cons.</small>
- Fabales <small>Bromhead</small>
- Quillajaceae <small>D.Don</small>
- Fabaceae <small>Lindl., nom. cons.</small> (= Leguminosae <small>Juss., nom. cons.</small>)
- Surianaceae <small>Arn., nom. cons.</small>
- Polygalaceae <small>Hoffmanns. & Link, nom. cons.</small>
- Rosales <small>Bercht. & J.Presl</small>
- Rosaceae <small>Juss., nom. cons.</small>
- Barbeyaceae <small>Rendle, nom. cons.</small>
- Dirachmaceae <small>Hutch.</small>
- Elaeagnaceae <small>Juss., nom. cons.</small>
- Rhamnaceae <small>Juss., nom. cons.</small>
- Ulmaceae <small>Mirb., nom. cons.</small>
- Cannabaceae <small>Martinov, nom. cons.</small>
- Moraceae <small>Gaudich., nom. cons.</small>
- Urticaceae <small>Juss., nom. cons.</small>
- Fagales <small>Engl.</small>
- Nothofagaceae <small>Kuprian.</small>
- Fagaceae <small>Dumort., nom. cons.</small>
- Myricaceae <small>Rich. ex Kunth, nom. cons.</small>
- Juglandaceae <small>DC. ex Perleb, nom. cons.</small>
- Casuarinaceae <small>R.Br., nom. cons.</small>
- Ticodendraceae <small>Gomez-Laur. & L.D.Gomez</small>
- Betulaceae <small>Gray, nom. cons.</small>
- Cucurbitales <small>Juss. ex Bercht. & J.Presl</small>
- Apodanthaceae <small>Tiegh. ex Takht.</small> *
- Anisophylleaceae <small>Ridl.</small>
- Corynocarpaceae <small>Engl., nom. cons.</small>
- Coriariaceae <small>DC., nom. cons.</small>
- Cucurbitaceae <small>Juss., nom. cons.</small>
- Tetramelaceae <small>Airy Shaw</small>
- Datiscaceae <small>Dumort., nom. cons.</small>
- Begoniaceae <small>C.Agardh, nom. cons.</small>
COM clade; placement uncertain
- Celastrales <small>Link</small>
- Lepidobotryaceae <small>J. Leonard, nom. cons.</small>
- Celastraceae <small>R.Br., nom. cons.</small>
- Oxalidales <small>Bercht. & J.Presl</small>
- Huaceae <small>A.Chev.</small>
- Connaraceae <small>R.Br., nom. cons.</small>
- Oxalidaceae <small>R.Br., nom. cons.</small>
- Cunoniaceae <small>R.Br., nom. cons.</small>
- Elaeocarpaceae <small>Juss., nom. cons.</small>
- Cephalotaceae <small>Dumort., nom. cons.</small>
- Brunelliaceae <small>Engl., nom. cons.</small>
- Malpighiales <small>Juss. ex Bercht. & J.Presl</small>
- Pandaceae <small>Engl. & Gilg, nom. cons.</small>
- Irvingiaceae <small>Exell & Mendonça, nom. cons.</small> * (including Allantospermum <small>Forman</small>)
- Ctenolophonaceae <small>Exell & Mendonça</small>
- Rhizophoraceae <small>Pers., nom. cons.</small>
- Erythroxylaceae <small>Kunth, nom. cons.</small>
- Ochnaceae <small>DC., nom. cons.</small>
- Bonnetiaceae <small>L.Beauvis. ex Nakai</small>
- Clusiaceae <small>Lindl., nom. cons.</small> (= Guttiferae <small>Juss., nom. cons.</small>)
- Calophyllaceae <small>J.Agardh</small>
- Podostemaceae <small>Rich. ex Kunth, nom. cons.</small>
- Hypericaceae <small>Juss., nom. cons.</small>
- Caryocaraceae <small>Voigt, nom. cons.</small>
- Lophopyxidaceae <small>H.Pfeiff.</small>
- Putranjivaceae <small>Meisn.</small>
- Centroplacaceae <small>Doweld & Reveal</small>
- Elatinaceae <small>Dumort., nom. cons.</small>
- Malpighiaceae <small>Juss., nom. cons.</small>
- Balanopaceae <small>Benth. & Hook.f., nom. cons.</small>
- Trigoniaceae <small>A.Juss., nom. cons.</small>
- Dichapetalaceae <small>Baill., nom. cons.</small>
- Euphroniaceae <small>Marc.-Berti</small>
- Chrysobalanaceae <small>R.Br., nom. cons.</small>
- Humiriaceae <small>A.Juss., nom. cons.</small>
- Achariaceae <small>Harms, nom. cons.</small>
- Violaceae <small>Batsch, nom. cons.</small>
- Goupiaceae <small>Miers</small>
- Passifloraceae <small>Juss. ex Roussel, nom. cons.</small>
- Lacistemataceae <small>Mart., nom. cons.</small>
- Salicaceae <small>Mirb., nom. cons.</small>
- Peraceae <small>Klotzsch</small> *
- Rafflesiaceae <small>Dumort., nom. cons.</small>
- Euphorbiaceae <small>Juss., nom. cons.</small> *
- Linaceae <small>DC. ex Perleb, nom. cons.</small>
- Ixonanthaceae <small>Planch. ex Miq., nom. cons.</small> *
- Picrodendraceae <small>Small, nom. cons.</small>
- Phyllanthaceae <small>Martinov, nom. cons.</small>
Malvids
- Geraniales <small>Juss. ex Bercht. & J.Presl</small>
- Geraniaceae <small>Juss., nom. cons.</small>
- Francoaceae <small>A.Juss., nom. cons.</small> * (including Bersamaceae <small>Doweld</small>, Greyiaceae <small>Hutch., nom. cons.</small>, Ledocarpaceae <small>Meyen</small>, Melianthaceae <small>Horan., nom. cons.</small>, Rhynchothecaceae <small>A.Juss.</small>, Vivianiaceae <small>Klotzsch, nom. cons.</small>)
- Myrtales <small>Juss. ex Bercht. & J.Presl</small>
- Combretaceae <small>R.Br., nom. cons.</small>
- Lythraceae <small>J.St.-Hil., nom. cons.</small>
- Onagraceae <small>Juss., nom. cons.</small>
- Vochysiaceae <small>A.St.-Hil., nom. cons.</small>
- Myrtaceae <small>Juss., nom. cons.</small>
- Melastomataceae <small>Juss., nom. cons.</small>
- Crypteroniaceae <small>A.DC., nom. cons.</small>
- Alzateaceae <small>S.A.Graham</small>
- Penaeaceae <small>Sweet ex Guill., nom. cons.</small>
- Crossosomatales <small>Takht. ex Reveal</small>
- Aphloiaceae <small>Takht.</small>
- Geissolomataceae <small>A.DC., nom. cons.</small>
- Strasburgeriaceae <small>Tiegh., nom. cons.</small>
- Staphyleaceae <small>Martinov, nom. cons.</small>
- Guamatelaceae <small>S.H.Oh & D.Potter</small>
- Stachyuraceae <small>J.Agardh, nom. cons.</small>
- Crossosomataceae <small>Engl., nom. cons.</small>
- Picramniales <small>Doweld</small>
- Picramniaceae <small>Fernando & Quinn</small>
- Huerteales <small>Doweld</small>
- Gerrardinaceae <small>M.H.Alford</small>
- Petenaeaceae <small>Christenh. et al.</small> *
- Tapisciaceae <small>Takht.</small>
- Dipentodontaceae <small>Merr., nom. cons.</small>
- Sapindales <small>Juss. ex Bercht. & J.Presl</small>
- Biebersteiniaceae <small>Schnizl.</small>
- Nitrariaceae <small>Lindl.</small>
- Kirkiaceae <small>Takht.</small>
- Burseraceae <small>Kunth, nom. cons.</small>
- Anacardiaceae <small>R.Br., nom. cons.</small>
- Sapindaceae <small>Juss., nom. cons.</small> (including Xanthocerataceae <small>Buerki et al.</small>, as 'Xanthoceraceae')
- Rutaceae <small>Juss., nom. cons.</small>
- Simaroubaceae <small>DC., nom. cons.</small>
- Meliaceae <small>Juss., nom. cons.</small>
- Malvales <small>Juss. ex Bercht. & J.Presl</small>
- Cytinaceae <small>A.Rich.</small>
- Muntingiaceae <small>C.Bayer et al.</small>
- Neuradaceae <small>Kostel., nom. cons.</small>
- Malvaceae <small>Juss., nom. cons.</small>
- Sphaerosepalaceae <small>Bullock</small>
- Thymelaeaceae <small>Juss., nom. cons.</small>
- Bixaceae <small>Kunth, nom. cons.</small>
- Cistaceae <small>Juss., nom. cons.</small> * (including Pakaraimaea <small>Maguire & P.S.Ashton</small>)
- Sarcolaenaceae <small>Caruel, nom. cons.</small>
- Dipterocarpaceae <small>Blume, nom. cons.</small> *
- Brassicales <small>Bromhead</small>
- Akaniaceae <small>Stapf, nom. cons.</small>
- Tropaeolaceae <small>Juss. ex DC., nom. cons.</small>
- Moringaceae <small>Martinov, nom. cons.</small>
- Caricaceae <small>Dumort., nom. cons.</small>
- Limnanthaceae <small>R.Br., nom. cons.</small>
- Setchellanthaceae <small>Iltis</small>
- Koeberliniaceae <small>Engl., nom. cons.</small>
- Bataceae <small>Mart. ex Perleb, nom. cons.</small>
- Salvadoraceae <small>Lindl., nom. cons.</small>
- Emblingiaceae <small>Airy Shaw</small>
- Tovariaceae <small>Pax, nom. cons.</small>
- Pentadiplandraceae <small>Hutch. & Dalziel</small>
- Gyrostemonaceae <small>A.Juss., nom. cons.</small>
- Resedaceae <small>Martinov, nom. cons.</small> * (including Borthwickiaceae <small>J.X.Su et al.</small>, Stixidaceae <small>Doweld</small> as 'Stixaceae', Forchhammeria <small>Liebm.</small>)
- Capparaceae <small>Juss., nom. cons.</small> *
- Cleomaceae <small>Bercht. & J.Presl</small>
- Brassicaceae <small>Burnett, nom. cons.</small> (= Cruciferae <small>Juss., nom. cons.</small>)
- Berberidopsidales <small>Doweld</small>
- Aextoxicaceae <small>Engl. & Gilg, nom. cons.</small>
- Berberidopsidaceae <small>Takht.</small>
- Santalales <small>R.Br. ex Bercht. & J.Presl</small>
- Olacaceae <small>R.Br., nom. cons.</small> (not monophyletic; including Aptandraceae <small>Miers</small>, Coulaceae <small>Tiegh.</small>, Erythropalaceae <small>Planch. ex Miq., nom. cons.</small>, Octoknemaceae <small>Soler. nom. cons.</small>, Strombosiaceae <small>Tiegh.</small>, Ximeniaceae <small>Horan.</small>)
- Opiliaceae <small>Valeton, nom. cons.</small>
- Balanophoraceae <small>Rich., nom. cons.</small>
- Santalaceae <small>R.Br., nom. cons.</small> (not monophyletic if Balanophoraceae are embedded; including Amphorogynaceae <small>Nickrent & Der</small>, Cervantesiaceae <small>Nickrent & Der</small>, Comandraceae <small>Nickrent & Der</small>, Nanodeaceae <small>Nickrent & Der</small>, Thesiaceae <small>Vest</small>, Viscaceae <small>Batsch</small>)
- Misodendraceae <small>J.Agardh, nom. cons.</small>
- Schoepfiaceae <small>Blume</small>
- Loranthaceae <small>Juss., nom. cons.</small>
- Caryophyllales <small>Juss. ex Bercht. & J.Presl</small>
- Frankeniaceae <small>Desv., nom. cons.</small>
- Tamaricaceae <small>Link, nom. cons.</small>
- Plumbaginaceae <small>Juss., nom. cons.</small>
- Polygonaceae <small>Juss., nom. cons.</small>
- Droseraceae <small>Salisb., nom. cons.</small>
- Nepenthaceae <small>Dumort, nom. cons.</small>
- Drosophyllaceae <small>Chrtek et al.</small>
- Dioncophyllaceae <small>Airy Shaw, nom. cons.</small>
- Ancistrocladaceae <small>Planch. ex Walp., nom. cons.</small>
- Rhabdodendraceae <small>Prance</small>
- Simmondsiaceae <small>Tiegh.</small>
- Physenaceae <small>Takht.</small>
- Asteropeiaceae <small>Takht. ex Reveal & Hoogland</small>
- Macarthuriaceae <small>Christenh.</small> *
- Microteaceae <small>Schäferhoff & Borsch</small> *
- Caryophyllaceae <small>Juss., nom. cons.</small>
- Achatocarpaceae <small>Heimerl, nom. cons.</small>
- Amaranthaceae <small>Juss., nom. cons.</small>
- Stegnospermataceae <small>Nakai</small>
- Limeaceae <small>Shipunov ex Reveal</small> *
- Lophiocarpaceae <small>Doweld & Reveal</small>
- Kewaceae <small>Christenh.</small> *
- Barbeuiaceae <small>Nakai</small>
- Gisekiaceae <small>Nakai</small>
- Aizoaceae <small>Martinov, nom. cons.</small>
- Phytolaccaceae <small>R.Br., nom. cons.</small> *
- Petiveriaceae <small>C.Agardh</small> * (including Rivinaceae <small>C.Agardh</small>)
- Sarcobataceae <small>Behnke</small>
- Nyctaginaceae <small>Juss., nom. cons.</small>
- Molluginaceae <small>Bartl., nom. cons.</small> *
- Montiaceae <small>Raf.</small>
- Didiereaceae <small>Radlk., nom. cons.</small>
- Basellaceae <small>Raf., nom. cons.</small>
- Halophytaceae <small>S.Soriano</small>
- Talinaceae <small>Doweld</small>
- Portulacaceae <small>Juss., nom. cons.</small>
- Anacampserotaceae <small>Eggli & Nyffeler</small>
- Cactaceae <small>Juss., nom. cons.</small>
- Cornales <small>Link</small>
- Nyssaceae <small>Juss. ex Dumort., nom. cons.</small>
- Hydrostachyaceae <small>Engl., nom. cons.</small>
- Hydrangeaceae <small>Dumort., nom. cons.</small>
- Loasaceae <small>Juss., nom. cons.</small>
- Curtisiaceae <small>Takht.</small>
- Grubbiaceae <small>Endl. ex Meisn., nom. cons.</small>
- Cornaceae <small>Bercht. & J.Presl, nom. cons.</small>
- Ericales <small>Bercht. & J.Presl</small>
- Balsaminaceae <small>A.Rich., nom. cons.</small>
- Marcgraviaceae <small>Bercht. & J.Presl, nom. cons.</small>
- Tetrameristaceae <small>Hutch.</small>
- Fouquieriaceae <small>DC., nom. cons.</small>
- Polemoniaceae <small>Juss., nom. cons.</small>
- Lecythidaceae <small>A.Rich., nom. cons.</small>
- Sladeniaceae <small>Airy Shaw</small>
- Pentaphylacaceae <small>Engl., nom. cons.</small>
- Sapotaceae <small>Juss., nom. cons.</small>
- Ebenaceae <small>Gürke, nom. cons.</small>
- Primulaceae <small>Batsch ex Borkh., nom. cons.</small>
- Theaceae <small>Mirb., nom. cons.</small>
- Symplocaceae <small>Desf., nom. cons.</small>
- Diapensiaceae <small>Lindl., nom. cons.</small>
- Styracaceae <small>DC. & Spreng., nom. cons.</small>
- Sarraceniaceae <small>Dumort., nom. cons.</small>
- Roridulaceae <small>Martinov, nom. cons.</small>
- Actinidiaceae <small>Gilg & Werderm., nom. cons.</small>
- Clethraceae <small>Klotzsch, nom. cons.</small>
- Cyrillaceae <small>Lindl., nom. cons.</small>
- Ericaceae <small>Juss., nom. cons.</small>
- Mitrastemonaceae <small>Makino, nom. cons.</small> (placement in order unclear)
Lamiids
- Icacinales <small>Tiegh.</small> âÂÂ
- Oncothecaceae <small>Kobuski ex Airy Shaw</small>
- Icacinaceae <small>Miers, nom. cons.</small> *
- Metteniusales <small>Takht.</small> âÂÂ
- Metteniusaceae <small>H.Karst. ex Schnizl.</small> *
- Garryales <small>Mart.</small>
- Eucommiaceae <small>Engl., nom. cons.</small>
- Garryaceae <small>Lindl., nom. cons.</small>
- Gentianales <small>Juss. ex Bercht. & J.Presl</small>
- Rubiaceae <small>Juss., nom. cons.</small>
- Gentianaceae <small>Juss., nom. cons.</small>
- Loganiaceae <small>R.Br. ex Mart., nom. cons.</small>
- Gelsemiaceae <small>L.Struwe & V.A.Albert</small> * (including Pteleocarpaceae <small>Brummitt</small>)
- Apocynaceae <small>Juss., nom. cons.</small>
- Boraginales <small>Juss. ex Bercht. & J.Presl</small> âÂÂ
- Boraginaceae <small>Juss., nom. cons.</small> (including Codonaceae <small>Weigend & Hilger</small>)
- Vahliales <small>Doweld</small> âÂÂ
- Vahliaceae <small>Dandy</small>
- Solanales <small>Juss. ex Bercht. & J.Presl</small>
- Convolvulaceae <small>Juss., nom. cons.</small>
- Solanaceae <small>Juss., nom. cons.</small>
- Montiniaceae <small>Nakai, nom. cons.</small>
- Sphenocleaceae <small>T.Baskerv., nom. cons.</small>
- Hydroleaceae <small>R.Br.</small>
- Lamiales <small>Bromhead</small>
- Plocospermataceae <small>Hutch.</small>
- Carlemanniaceae <small>Airy Shaw</small>
- Oleaceae <small>Hoffmanns. & Link, nom. cons.</small>
- Tetrachondraceae <small>Wettst.</small>
- Calceolariaceae <small>Olmstead</small>
- Gesneriaceae <small>Rich. & Juss., nom. cons.</small> * (the position of Peltanthera <small>Benth.</small> is problematic and here considered unplaced to family)
- Plantaginaceae <small>Juss., nom. cons.</small>
- Scrophulariaceae <small>Juss., nom. cons.</small>
- Stilbaceae <small>Kunth, nom. cons.</small>
- Linderniaceae <small>Borsch et al.</small>
- Byblidaceae <small>Domin, nom. cons.</small>
- Martyniaceae <small>Horan., nom. cons.</small>
- Pedaliaceae <small>R.Br., nom. cons.</small>
- Acanthaceae <small>Juss., nom. cons.</small>
- Bignoniaceae <small>Juss., nom. cons.</small>
- Lentibulariaceae <small>Rich., nom. cons.</small>
- Schlegeliaceae <small>Reveal</small>
- Thomandersiaceae <small>Sreem.</small>
- Verbenaceae <small>J.St.Hil., nom. cons.</small>
- Lamiaceae <small>Martinov, nom. cons.</small> (= Labiatae <small>Juss., nom. cons.</small>)
- Mazaceae <small>Reveal</small>
- Phrymaceae <small>Schauer, nom. cons.</small> *
- Paulowniaceae <small>Nakai</small>
- Orobanchaceae <small>Vent., nom. cons.</small> * (including Lindenbergiaceae <small>Doweld</small>, Rehmanniaceae <small>Reveal</small>)
Campanulids
- Aquifoliales <small>Senft</small>
- Stemonuraceae <small>KÃÂ¥rehed</small>
- Cardiopteridaceae <small>Blume, nom. cons.</small>
- Phyllonomaceae <small>Small</small>
- Helwingiaceae <small>Decne.</small>
- Aquifoliaceae <small>Bercht. & J.Presl, nom. cons.</small>
- Asterales <small>Link</small>
- Rousseaceae <small>DC.</small>
- Campanulaceae <small>Juss., nom. cons.</small>
- Pentaphragmataceae <small>J.Agardh, nom. cons.</small>
- Stylidiaceae <small>R.Br., nom. cons.</small>
- Alseuosmiaceae <small>Airy Shaw</small>
- Phellinaceae <small>Takht.</small>
- Argophyllaceae <small>Takht.</small>
- Menyanthaceae <small>Dumort., nom. cons.</small>
- Goodeniaceae <small>R.Br., nom. cons.</small>
- Calyceraceae <small>R.Br. ex Rich., nom. cons.</small>
- Asteraceae <small>Bercht. & J.Presl, nom. cons.</small> (= Compositae <small>Giseke, nom. cons.</small>)
- Escalloniales <small>Link</small>
- Escalloniaceae <small>R.Br. ex Dumort., nom. cons.</small>
- Bruniales <small>Dumort.</small>
- Columelliaceae <small>D.Don, nom. cons.</small>
- Bruniaceae <small>R.Br. ex DC., nom. cons.</small>
- Paracryphiales <small>Takht. ex Reveal</small>
- Paracryphiaceae <small>Airy Shaw</small>
- Dipsacales <small>Juss. ex Bercht. & J.Presl</small>
- Adoxaceae <small>E.Mey., nom. cons.</small> (= Viburnaceae <small>Raf., nom. cons. prop.</small>)
- Caprifoliaceae <small>Juss., nom. cons.</small>
- Apiales <small>Nakai</small>
- Pennantiaceae <small>J.Agardh</small>
- Torricelliaceae <small>Hu</small>
- Griseliniaceae <small>Takht., nom. cons. prop.</small>
- Pittosporaceae <small>R.Br., nom. cons.</small>
- Araliaceae <small>Juss., nom. cons.</small>
- Myodocarpaceae <small>Doweld</small>
- Apiaceae <small>Lindl., nom. cons.</small> (= Umbelliferae <small>Juss., nom. cons.</small>)
Phylogeny
Like the earlier APG systems, the APG IV revision is based on a phylogenetic tree for the angiosperms, as shown below.
References
Bibliography