Lithobius is a large genus of centipedes in the family Lithobiidae, commonly called stone centipedes, common centipedes or brown centipedes.
Description
Most Lithobius species are typical representatives of the family Lithobiidae. They are about long and brownish in colour. The adult's body has 18 segments, and 15 pairs of legs. The special characteristics include the dispersed openings of coxal glands of the last pair of legs.
Behaviour and ecology
Stone centipedes are found under stones or bark, in soil and decaying matter. Some are common in gardens. Lithobius forficatus is the most abundant centipede species in Europe. Like other centipedes, they are more active at night. They feed on insects and other small invertebrates. The eggs are deposited singly in soil. The lifespan can be over 3 years.
Taxonomy
The genus Lithobius was erected in 1814 by William Elford Leach, in an article published in David Brewster's Edinburgh Encyclopædia. The name derives from Ancient Greek (lÃÂthos), meaning "stone", and (bÃÂos), meaning "life". Leach did not designate a type species, and none was selected until Pierre André Latreille chose Scolopendra forficata <small>Linnaeus, 1758</small> (now Lithobius forficatus). Lithobius is the name-giving genus for the family Lithobiidae.
Lithobius is one of about 100 genera or subgenera in the subfamily Lithobiinae. It contains over 500 species and numerous subspecies. The classification of species inside the genus is a matter of discussion too. Some authors divide the genus into subgenera Eulithobius, Lithobius, Monotarsobius, Neolithobius, Pleurolithobius, Pseudolithobius, Sigibius, Thracolithobius and Troglolithobius.
Species
The genus Lithobius includes the following species:
- Lithobius acherontis <small>Verhoeff, 1900</small>
- Lithobius acipayamus <small>Chamberlin, 1952</small>
- Lithobius adherens <small>Chamberlin, 1952</small>
- Lithobius aidonensis <small>Verhoeff, 1943</small>
- Lithobius allotiphlus <small>(Silvestri, 1908)</small>
- Lithobius alpicosiensis <small>Latzel, 1887</small>
- Lithobius alpinus <small>Koch, 1862 non Matic & Darabantu, 1971</small>
- Lithobius altus <small>Matic & Darabantu, 1971</small>
- Lithobius ameles <small>Chamberlin, 1952</small>
- Lithobius angolianus <small>Chamberlin, 1952</small>
- Lithobius anisanus <small>Verhoeff, 1937</small>
- Lithobius ankarensis <small>Verhoeff, 1944</small>
- Lithobius anodus <small>Latzel, 1880</small>
- Lithobius antipai <small>Matic, 1969</small>
- Lithobius aostanus <small>Verhoeff, 1934</small>
- Lithobius apfelbecki <small>Verhoeff, 1900</small>
- Lithobius ardesiacus <small>Fedrizzi, 1877</small>
- Lithobius argaeensis <small>Attems, 1905</small>
- Lithobius argus <small>Newport, 1844</small>
- Lithobius armatus <small>Selivanov, 1878</small>
- Lithobius asper <small>Muralevitch, 1926</small>
- Lithobius athesinus <small>Verhoeff, 1937</small>
- Lithobius atticus <small>Verhoeff, 1901</small>
- Lithobius audax <small>Meinert, 1872</small>
- Lithobius bartsiokasi <small>Matic & Stavropoulos, 1990</small>
- Lithobius bebekensis <small>(Verhoeff, 1944)</small>
- Lithobius berkeleyensis <small>Verhoeff, 1937</small>
- Lithobius beroni <small>Negrea, 1965</small>
- Lithobius biarmatus <small>Matic, 1970</small>
- Lithobius bicolor <small>Tömösvary, 1879</small>
- Lithobius bicuspidatus <small>(Matic, 1957)</small>
- Lithobius bifidus <small>Matic, 1973</small>
- Lithobius binaghii <small>Manfredi, 1937</small>
- Lithobius biporus <small>Silvestri, 1894</small>
- Lithobius boettgeri <small>Verhoeff, 1925</small>
- Lithobius bolognai <small>Zapparoli, 1991</small>
- Lithobius bonensis <small>Meinert, 1872</small>
- Lithobius bosporanus <small>(Verhoeff, 1941)</small>
- Lithobius brachycephalus <small>Fanzago, 1880</small>
- Lithobius brevicornis <small>Daday, 1889</small>
- Lithobius buakheriacus <small>Zapparoli, 1985</small>
- Lithobius bulgaricus <small>(Verhoeff, 1925)</small>
- Lithobius bullatus <small>Eason, 1993</small>
- Lithobius burzenlandicus <small>(Verhoeff, 1931)</small>
- Lithobius buxtoni <small>Brölemann, 1924</small>
- Lithobius calabrensis <small>Fedrizzi, 1878</small>
- Lithobius calamatanus <small>(Verhoeff, 1899)</small>
- Lithobius canariensis <small>Eason, 1992</small>
- Lithobius cantabrigensis <small>Meinert</small>
- Lithobius capreae <small>Verhoeff, 1943</small>
- Lithobius carmenae <small>Matic, 1968</small>
- Lithobius cassinensis <small>Verhoeff, 1925</small>
- Lithobius castaneus <small>Newport, 1844</small>
- Lithobius cavernicola <small>Fanzago, 1877</small>
- Lithobius cerberulus <small>Verhoeff, 1941</small>
- Lithobius cerii <small>Verhoeff, 1937</small>
- Lithobius chalusensis <small>Matic, 1969</small>
- Lithobius chikerensis <small>Verhoeff, 1936</small>
- Lithobius cinnamomenus <small>Koch, 1862</small>
- Lithobius clarki <small>Eason, 1975</small>
- Lithobius communis <small>Koch, 1844</small>
- Lithobius consimilis <small>Eason, 1992</small>
- Lithobius corcyraeus <small>Verhoeff, 1899</small>
- Lithobius coriaceus <small>Koch, 1862</small>
- Lithobius corrigendus <small>Dobroruka, 1988</small>
- Lithobius crassipes <small>Koch, 1862</small>
- Lithobius creticus <small>Dobroruka, 1977</small>
- Lithobius crissolensis <small>Verhoeff</small>
- Lithobius croaticus <small>Matic & Teodoreanu, 1967</small>
- Lithobius cryptobius <small>Silvestri, 1897</small>
- Lithobius curtipes <small>Koch, 1847</small>
- Lithobius curtirostris <small>Eisen & Stuxberg, 1868</small>
- Lithobius dacicus <small>(Matic, 1959)</small>
- Lithobius dadayi <small>Tömösvary, 1880</small>
- Lithobius dahlii <small>Verhoeff, 1925</small>
- Lithobius degerboelae <small>Eason, 1981 or 1986</small>
- Lithobius demavendicus <small>Matic, 1969</small>
- Lithobius depressus <small>Fanzago, 1880</small>
- Lithobius diana <small>Verhoeff, 1901</small>
- Lithobius dieuzeidei <small>Brölemann, 1931</small>
- Lithobius discolor <small>Verhoeff, 1937</small>
- Lithobius doderoi <small>Silvestri, 1908</small>
- Lithobius dolinophilus <small>Verhoeff, 1937</small>
- Lithobius dollfusi <small>Verhoeff, 1931</small>
- Lithobius domogledicus <small>(Matic, 1961)</small>
- Lithobius doriae <small>Pocock, 1890</small>
- Lithobius drescoi <small>Demange, 1958</small>
- Lithobius dudichi <small>Loksa, 1947</small>
- Lithobius easoni <small>Matic, 1969</small>
- Lithobius elbanus <small>Verhoeff, 1931</small>
- Lithobius elbursensis <small>Matic, 1969</small>
- Lithobius electrinus <small>(Verhoeff, 1937)</small>
- Lithobius electron <small>Verhoeff, 1928</small>
- Lithobius electus <small>Silvestri, 1935</small>
- Lithobius elegans <small>Selivanov, 1931</small>
- Lithobius enghofi <small>Eason, 1986</small>
- Lithobius ercijasius <small>Verhoeff, 1943</small>
- Lithobius erdschiasius <small>Verhoeff, 1943</small>
- Lithobius errantus <small>Chamberlin, 1952</small>
- Lithobius erythrocephalus <small>C.L.Koch, 1847</small>
- Lithobius eucuemis <small>Stuxberg</small>
- Lithobius evae <small>Dobroruka, 1958</small>
- Lithobius evasus <small>(Chamberlin, 1952)</small>
- Lithobius excellens <small>Silvestri, 1894</small>
- Lithobius exiguus <small>Meinert, 1886</small>
- Lithobius eximius <small>Meinert, 1872</small>
- Lithobius falteronensis <small>Manfredi, 1936</small>
- Lithobius fangensis <small>Eason, 1986</small>
- Lithobius fanzagoi <small>Fedrizzi, 1877</small>
- Lithobius feae
- Lithobius femorosulcatus <small>Eason, 1986</small>
- Lithobius festivus <small>Koch, 1862</small>
- Lithobius finitimus <small>Fanzago, 1878</small>
- Lithobius flavus <small>Meinert, 1872</small>
- Lithobius forficatus <small>(Linnaeus, 1758)</small>
- Lithobius fossor <small>Koch, 1862</small>
- Lithobius franzi <small>Attems, 1949</small>
- Lithobius galatheae <small>Meinert, 1886</small>
- Lithobius geyeri <small>Verhoeff, 1935</small>
- Lithobius glabratus <small>Koch, 1847</small>
- Lithobius glacialis <small>Verhoeff, 1937</small>
- Lithobius gracilipes <small>Meinert, 1872</small>
- Lithobius gracilis <small>Meinert, 1872</small>
- Lithobius grandiporosus <small>Verhoeff, 1937</small>
- Lithobius granulatus <small>Koch, 1862</small>
- Lithobius gridellii <small>Manfredi, 1955</small>
- Lithobius grossidens <small>Meinert, 1872</small>
- Lithobius grossipes <small>Koch, 1847</small>
- Lithobius haarlovi <small>Eason, 1986</small>
- Lithobius hardwickei <small>Newport, 1844</small>
- Lithobius hatayensis <small>Verhoeff, 1943</small>
- Lithobius hawaiiensis <small>Silvestri, 1904</small>
- Lithobius helvolus <small>Attems, 1951</small>
- Lithobius herzegowinensis <small>Verhoeff, 1900</small>
- Lithobius hispanicus <small>Meinert, 1872</small>
- Lithobius hopanus <small>(Chamberlin, 1952)</small>
- Lithobius hortensis <small>Koch, 1862</small>
- Lithobius hummelii <small>Verhoeff, 1933</small>
- Lithobius ilgazensis <small>Matic, 1983</small>
- Lithobius immutabilis <small>Koch, 1862</small>
- Lithobius imperialis <small>Meinert, 1872</small>
- Lithobius inaequidens <small>Fedrizzi, 1877</small>
- Lithobius incertus <small>Matic, 1966</small>
- Lithobius indicus <small>Eason, 1981</small>
- Lithobius inexpectatus <small>Matic, 1962</small>
- Lithobius infossus <small>Silvestri, 1894</small>
- Lithobius inopinatus <small>Matic, 1970</small>
- Lithobius inquirendus <small>Attems, 1951</small>
- Lithobius insignis <small>Meinert, 1872</small>
- Lithobius ionicus <small>(Silvestri, 1896)</small>
- Lithobius iranicus <small>Attems, 1951</small>
- Lithobius italicus <small>Matic, 1957</small>
- Lithobius javanicus <small>Pocock, 1894</small>
- Lithobius kansuanus <small>Verhoeff, 1933</small>
- Lithobius karamani <small>Verhoeff, 1937</small>
- Lithobius kastamouensis <small>Matic, 1983</small>
- Lithobius koenigi <small>Verhoeff, 1891</small>
- Lithobius koreanus <small>Verhoeff, 1938</small>
- Lithobius kosswigi <small>Chamberlin, 1952</small>
- Lithobius kurdistanus <small>Verhoeff, 1944</small>
- Lithobius laccatus <small>Attems, 1951</small>
- Lithobius laevilabrum <small>Leach, 1815</small>
- Lithobius lagrecai <small>Matic, 1962</small>
- Lithobius lanzai <small>Matic, 1961</small>
- Lithobius lapadensis <small>Verhoeff, 1900</small>
- Lithobius latebricola <small>Meinert, 1872</small>
- Lithobius latzellii <small>Meinert</small>
- Lithobius lindbergi <small>Chamberlin, 1956</small>
- Lithobius litoralis <small>Koch, 1867</small>
- Lithobius lobifer <small>(Chamberlin, 1952)</small>
- Lithobius loeiensis <small>Eason, 1986</small>
- Lithobius lorioli <small>Demange, 1962</small>
- Lithobius lubricus <small>Koch, 1862</small>
- Lithobius lundii <small>Meinert, 1886</small>
- Lithobius luteus <small>Loksa, 1947</small>
- Lithobius macrocentrus <small>Attems, 1949</small>
- Lithobius magnus <small>(Trotzina, 1894)</small>
- Lithobius magurensis <small>Dobroruka, 1971</small>
- Lithobius malaccanus <small>Verhoeff, 1937</small>
- Lithobius malayicus <small>Verhoeff, 1937</small>
- Lithobius marginatus <small>Fedrizzi, 1877</small>
- Lithobius martensi <small>Eason, 1989</small>
- Lithobius matici <small>Prunescu, 1966</small>
- Lithobius matulicii <small>Verhoeff, 1899</small>
- Lithobius maximovici <small>Folkmanova, 1946</small>
- Lithobius mediolus <small>Chamberlin, 1952</small>
- Lithobius megalaporus <small>(Stuxberg, 1875)</small>
- Lithobius meifengensis <small>Chao, Lee, Chang, 2018</small>
- Lithobius melanocephalus <small>Koch, 1863</small>
- Lithobius memorabilis <small>Attems, 1951</small>
- Lithobius meridionalis <small>Fedrizzi, 1877</small>
- Lithobius micropodus <small>Matic, 1983</small>
- Lithobius microps <small>Meinert, 1868</small>
- Lithobius minellii <small>Matic & Darabantu, 1971</small>
- Lithobius minimus <small>Koch, 1862</small>
- Lithobius minutus <small>Koch, 1847</small>
- Lithobius moananus <small>(Chamberlin, 1926)</small>
- Lithobius moellensis <small>Verhoeff, 1940</small>
- Lithobius molleri <small>Verhoeff, 1893</small>
- Lithobius molophai <small>Restivo de Miranda, 1978</small>
- Lithobius mongolicus <small>Verhoeff, 1933</small>
- Lithobius montellicus <small>Fanzago, 1874</small>
- Lithobius mordax <small>Koch, 1862</small>
- Lithobius multidentatus <small>Newport</small>
- Lithobius muscorum <small>Koch, 1862</small>
- Lithobius nigrocullis <small>Folkmanova, 1928</small>
- Lithobius nocellensis <small>Verhoeff, 1943</small>
- Lithobius nodulipes <small>Latzel, 1880</small>
- Lithobius nudicornis <small>Gervais, 1837</small>
- Lithobius oblongus <small>Sseliwanoff, 1881</small>
- Lithobius obscurus <small>Meinert, 1872</small>
- Lithobius occultus <small>Silvestri, 1894</small>
- Lithobius ocraceus <small>Fedrizzi, 1878</small>
- Lithobius octops <small>Menge, 1851</small>
- Lithobius oellatus <small>Verhoeff, 1925</small>
- Lithobius oligoporus <small>Latzel, 1885</small>
- Lithobius oraniensis <small>Matic, 1967</small>
- Lithobius orientis <small>(Chamberlin, 1952)</small>
- Lithobius orotavae <small>Latzel, 1895</small>
- Lithobius pachypus <small>Verhoeff, 1925</small>
- Lithobius paghmanensis <small>Eason, 1986</small>
- Lithobius paidamus <small>Chamberlin, 1952</small>
- Lithobius palmarum <small>Verhoeff, 1934</small>
- Lithobius palnis <small>(Eason, 1973)</small>
- Lithobius palustris <small>Selivanov, 1878</small>
- Lithobius pantokratoris <small>Attems, 1903</small>
- Lithobius paradisiacus <small>Matic & Darabantu, 1971</small>
- Lithobius parisiensis <small>Koch, 1862</small>
- Lithobius parvicornis <small>(Porat, 1893)</small>
- Lithobius parvolus <small>Fedrizzi, 1877</small>
- Lithobius patriarchalis <small>(Berlese, 1894)</small>
- Lithobius paurus <small>Chamberlin, 1952</small>
- Lithobius pedemontanus <small>Matic & Darabantu, 1971</small>
- Lithobius peggauensis <small>Verhoeff, 1937</small>
- Lithobius peregrinus <small>Latzel, 1880</small>
- Lithobius persicus <small>Pocock, 1899</small>
- Lithobius piceiavus <small>Verhoeff, 1901</small>
- Lithobius pilatoi <small>Matic, 1968</small>
- Lithobius pleonops <small>Menge, 1851</small>
- Lithobius plesius <small>(Chamberlin, 1952)</small>
- Lithobius plumbeus <small>Manfredi, 1948</small>
- Lithobius politicus <small>Chamberlin, 1952</small>
- Lithobius polyodontus <small>Attems, 1951</small>
- Lithobius ponzianus <small>Matic & Darabantu, 1969</small>
- Lithobius portchinski <small>Sseliwanoff, 1881</small>
- Lithobius postspoliatus <small>Verhoeff, 1942</small>
- Lithobius potanini <small>Selivanov, 1881</small>
- Lithobius pseudoagilis <small>Dobroruka, 1965</small>
- Lithobius pubescens <small>Koch, 1867</small>
- Lithobius pulcher <small>Meinert, 1872</small>
- Lithobius purkynei <small>Dobroruka, 1957</small>
- Lithobius purpureus <small>Chamberlin, 1901</small>
- Lithobius pustulatus <small>Matic, 1964</small>
- Lithobius pygmaeus <small>Latzel, 1880</small>
- Lithobius quadridentatus <small>Menge, 1851</small>
- Lithobius rapax <small>Meinert, 1872</small>
- Lithobius radjai <small>ÃÂtienne Iorio & Ivan Kos, 2021</small>
- Lithobius rapitus <small>Chamberlin, 1952</small>
- Lithobius reiseri <small>Verhoeff, 1900</small>
- Lithobius remyi <small>Jawlowski, 1923</small>
- Lithobius rhaeticus <small>Meinert, 1872</small>
- Lithobius rhiknus <small>Attems, 1951</small>
- Lithobius riggioi <small>Matic, 1968</small>
- Lithobius rilaicus <small>Verhoeff, 1937</small>
- Lithobius ruffoi <small>Matic, 1966</small>
- Lithobius rupivagus <small>Verhoeff, 1937</small>
- Lithobius saalachiensis <small>Verhoeff, 1937</small>
- Lithobius salernitanus <small>Manfredi, 1956</small>
- Lithobius salicis <small>Verhoeff, 1925</small>
- Lithobius sardous <small>Silvestri, 1897</small>
- Lithobius sbordonii <small>Matic, 1967</small>
- Lithobius scabrior <small>Chamberlin, 1920</small>
- Lithobius schawalleri <small>Eason, 1989</small>
- Lithobius scotophilus <small>Latzel, 1887</small>
- Lithobius sculpturatus <small>(Pocock, 1901)</small>
- Lithobius scutigeroides <small>Verhoeff, 1892</small>
- Lithobius semperi <small>Haase, 1887</small>
- Lithobius separatus <small>Verhoeff, 1943</small>
- Lithobius serbicus <small>Matic, 1979</small>
- Lithobius sibillinicus <small>Matic, 1966</small>
- Lithobius sibiricus <small>(Gerstfeldt, 1858)</small>
- Lithobius sicheli <small>Matic, 1986</small>
- Lithobius silvivagus <small>Verhoeff, 1925</small>
- Lithobius simrothi <small>Verhoeff, 1937</small>
- Lithobius sitianus <small>Chamberlin, 1956</small>
- Lithobius sloanei <small>Koch, 1862</small>
- Lithobius socius <small>Chamberlin, 1901</small>
- Lithobius songi <small>Pei et al., 2011</small>
- Lithobius sordidus <small>Koch, 1862</small>
- Lithobius speluncarum <small>Fanzago, 1877</small>
- Lithobius sphinx <small>(Verhoeff, 1941)</small>
- Lithobius stammeri <small>Verhoeff, 1939</small>
- Lithobius subtilis <small>Latzel, 1880</small>
- Lithobius suevicus <small>Meinert, 1863</small>
- Lithobius sulcatus <small>Koch, 1862</small>
- Lithobius sumatranus <small>Silvestri, 1895</small>
- Lithobius svenhedini <small>Verhoeff, 1933</small>
- Lithobius tahirensis <small>Matic, 1983</small>
- Lithobius targionii <small>Fanzago, 1874</small>
- Lithobius tatricus <small>Dobroruka, 1958</small>
- Lithobius tauricus <small>Selivanov, 1878</small>
- Lithobius temnensis <small>Verhoeff, 1943</small>
- Lithobius tenuicornis <small>Verhoeff, 1937</small>
- Lithobius tenuinguis <small>Eason, 1980</small>
- Lithobius tetraspinus <small>Pei, Lu, Liu, Hou, Ma, 2018</small>
- Lithobius terreus <small>Fedrizzi, 1877</small>
- Lithobius thetidis <small>Karsch, 1880</small>
- Lithobius tibiosetosus <small>Eason, 1986</small>
- Lithobius tibiotenuis <small>Eason, 1989</small>
- Lithobius tidissimus <small>(Chamberlin, 1952)</small>
- Lithobius tiphlus <small>(Latzel, 1886)</small>
- Lithobius transmarinus <small>Koch, 1862</small>
- Lithobius trebinjanus <small>(Verhoeff, 1900)</small>
- Lithobius tricuspis <small>Meinert, 1872</small>
- Lithobius trilineatus <small>Koch, 1862</small>
- Lithobius trinacrius <small>Verhoeff, 1925</small>
- Lithobius turritanus <small>Fanzago, 1881</small>
- Lithobius tweedii <small>Verhoeff, 1937</small>
- Lithobius tylopus <small>Latzel, 1882</small>
- Lithobius typhlus <small>Latzel, 1886</small>
- Lithobius ulterior <small>(Chamberlin, 1952)</small>
- Lithobius uludagensis <small>Matic, 1983</small>
- Lithobius variegatus <small>Leach, 1814</small>
- Lithobius varioporus <small>Pei et al., 2020</small>
- Lithobius varius <small>Koch, 1863</small>
- Lithobius velox <small>Koch, 1862</small>
- Lithobius venator <small>Koch, 1862</small>
- Lithobius veronensis <small>Matic & Darabantu, 1971</small>
- Lithobius viduus <small>Attems</small>
- Lithobius vinciguerrae <small>Silvestri, 1895</small>
- Lithobius vindelicius <small>Verhoeff, 1935</small>
- Lithobius vinosus <small>(Fanzago, 1874)</small>
- Lithobius violaceus <small>Fedrizzi, 1877</small>
- Lithobius visicae <small>Ribarov, 1987</small>
- Lithobius vorax <small>Meinert, 1872</small>
- Lithobius vosseleri <small>Verhoeff</small>
- Lithobius vulgaris <small>Leach, 1817</small>
- Lithobius walachius <small>Verhoeff, 1901</small>
- Lithobius weberi <small>Pocock, 1894</small>
- Lithobius werneri <small>Attems, 1902</small>
- Lithobius zachii <small>Restivo de Miranda, 1978</small>
- Lithobius zeylanus <small>(Chamberlin, 1952)</small>
References
External links