Vachellia

Vachellia is a genus of flowering plants in the legume family, Fabaceae, commonly known as thorn trees or acacias. It belongs to the subfamily Mimosoideae. Its species were considered members of genus Acacia until 2009. Vachellia can be distinguished from other acacias by its capitate inflorescences and spinescent stipules. Before discovery of the New World, Europeans in the Mediterranean region were familiar with several species of Vachellia, which they knew as sources of medicine, and had names for them that they inherited from the Greeks and Romans.

The wide-ranging genus occurs in a variety of open, tropical to subtropical habitats, and is locally dominant. In parts of Africa, Vachellia species are shaped progressively by grazing animals of increasing size and height, such as gazelle, gerenuk, and giraffe. The genus in Africa has thus developed thorns in defence against such herbivory.

Nomenclature

By 2005, taxonomists had decided that Acacia sensu lato should be split into at least five separate genera. The ICN dictated that under these circumstances, the name of Acacia should remain with the original type, which was Acacia nilotica. However, that year the General Committee of the IBC decided that Acacia should be given a new type (Acacia penninervis) so that the ~920 species of Australian acacias would not need to be renamed Racosperma. This decision was opposed by 54.9% or 247 representatives at its 2005 congress, while 45.1% or 203 votes were cast in favor. However, since a 60% vote was required to override the committee, the decision was carried, and a nom. cons. propositum was listed in Appendix III (p. 286). The 2011 congress voted 373 to 172 to uphold the 2005 decision, which means that the name Acacia and a new type follow the majority of the species in Acacia sensu lato, rather than this genus. However, some members of the botanical community remain unconvinced, and the use of Acacia in the scientific literature continues to exceed the use of the new generic names.

Description

The members of Vachellia are trees or shrubs, sometimes climbing, and are always armed. Younger plants, especially, are armed with spines which are modified stipules, situated near the leaf bases. Some (cf. V. tortilis, ', V. luederitzii and V. reficiens) are also armed with paired, recurved prickles (in addition to the spines). The leaves are alternate and bipinnately arranged, and their pinnae are usually opposite. The racemose inflorescences usually grow from the leaf axils. The yellow or creamy white flowers are produced in spherical heads, or seldom in elongate spikes, which is the general rule in the related genus Senegalia. The flowers are typically bisexual with numerous stamens, but unisexual flowers have been noted in V. nilotica (cf. Sinha, 1971). The calyx and corolla are usually 4 to 5-lobed. Glands are usually present on the rachis and the upper side of the petiole. The seed pod may be straight, curved or curled, and either dehiscent or indehiscent.

Species list

Of the 163 species currently assigned to Vachellia, 52 are native to the Americas, 83 to Africa, Madagascar and the Mascarene Islands, 32 to Asia and 9 to Australia and the Pacific Islands. Vachellia comprises the following species:

Vachellia abyssinica (Hochst. ex. Benth.) Kyal. & Boatwr.Ã¢ÂÂflat top acacia

subsp. abyssinica (Hochst. ex. Benth.) Kyal. & Boatwr.

subsp. calophylla (Brenan) Kyal. & Boatwr.

Vachellia acuifera (Benth.) Seigler & EbingerÃ¢ÂÂBahama acacia, cassip, pork-and-doughboy, (Bahamas) rosewood

Vachellia albicortata (Burkart) Seigler & Ebinger

Vachellia allenii (D. H. Janzen) Seigler & EbingerÃ¢ÂÂAllen acacia

Vachellia amythethophylla (Steud. ex A.Rich.) Kyal. & Boatwr.

Vachellia ancistroclada (Brenan) Kyal. & Boatwr.

Vachellia anegadensis (Britton) Seigler & EbingerÃ¢ÂÂAnegada acacia, blackbrush-wattle, pokemeboy

Vachellia antunesii (Harms) Kyal. & Boatwr.

Vachellia arenaria (Schinz) Kyal. & Boatwr.Ã¢ÂÂsand acacia

Vachellia aroma (Gillies ex Hook. & Arn.) Seigler & Ebinger

var. aroma Gillies ex Hook. & Arn.

var. huarango RuÃÂz & J.Macbr.

Vachellia astringens (Gillies in Hook. et Arn.) Speg.

Vachellia baessleri Clarke, Siegler & Ebinger

Vachellia barahonensis (Urb. & Ekman) Seigler & Ebinger

Vachellia bavazzanoi (Pichi-Sermolli) Kyal. & Boatwr.

Vachellia belairioides (Urb.) Seigler & EbingerÃ¢ÂÂBellair acacia

Vachellia bellula (Drake) Boatwr.

Vachellia biaciculata (S. Watson) Seigler & Ebinger

Vachellia bidwillii (Benth.) KodelaÃ¢ÂÂcorkwood wattle, dogwood. "'Waneu', of the aboriginals of Central Queensland; 'Yadthor', of those of the Cloncurry River, Northern Queensland."

Vachellia bilimekii (J. Macbr.) Seigler & Ebinger

Vachellia bolei (R.P. Subhedar) Ragupathy, Seigler, Ebinger & Maslin

Vachellia borleae (Burtt Davy) Kyal. & Boatwr.Ã¢ÂÂsticky acacia, named for the collector Jeanne M. Borle.

Vachellia brandegeana (I. M. Johnst.) Seigler & EbingerÃ¢ÂÂBaja California acacia

Vachellia bravoensis (Isely) Seigler & Ebinger

Vachellia bricchettiana (Chiov.) Kyal. & Boatwr.

Vachellia bucheri (Marie-VictorÃÂn) Seigler & EbingerÃ¢ÂÂBucher acacia

Vachellia bullockii (Brenan) Kyal. & Boatwr.

var. bullockii (Brenan) Kyal. & Boatwr.

var. induta (Brenan) Kyal. & Boatwr.

Vachellia burttii (Bak. f.) Kyal. & Boatwr.

Vachellia bussei (Harms ex SjÃÂ¶stedt) Kyal. & Boatwr.

Vachellia californica (Brandegee) Seigler & Ebinger

Vachellia campechiana (Mill.) Seigler & EbingerÃ¢ÂÂboatthorn acacia, spoon-thorn acacia

f. campechiana (Mill.) Seigler & Ebinger

f. houghii (Britton & Rose) Seigler & Ebinger

Vachellia caurina (Barneby & Zanoni) Seigler & Ebinger

Vachellia caven (Molina) Seigler & Ebinger

var. caven (Molina) Seigler & Ebinger

var. dehiscens (Ciald.) Seigler & Ebinger

var. microcarpa (Speg.) Seigler & Ebinger

var. stenocarpa (Speg.) Seigler & Ebinger

Vachellia cernua (Thulin & Hassan) Kyal. & Boatwr.

Vachellia chiapensis (Saff.) Seigler & EbingerÃ¢ÂÂChiapas acacia

Vachellia choriophylla (Benth.) Seigler & EbingerÃ¢ÂÂcinnecord acacia, Florida acacia, (Bahamas) cinnecord

Vachellia clarksoniana (Pedley) Kodela

Vachellia collinsii (Saff.) Seigler & EbingerÃ¢ÂÂCollins acacia

Vachellia constricta (Benth.) Seigler & EbingerÃ¢ÂÂWhitethorn acacia, Mescat acacia

Vachellia cookii (Saff.) Seigler & EbingerÃ¢ÂÂCook acacia, cockspur acacia

Vachellia cornigera (L.) Seigler & EbingerÃ¢ÂÂbullhorn wattle, bull's-horn acacia, bull-horn thorn, oxhorn acacia

Vachellia cucuyo (Barneby & Zanoni) Seigler & EbingerÃ¢ÂÂCucuyo acacia

Vachellia curvifructa (Burkart) Seigler & Ebinger

Vachellia daemon (Ekman & Urb.) Seigler & EbingerÃ¢ÂÂCamagÃÂ¼ey acacia

Vachellia davyi (N.E.Br.) Kyal. & Boatwr.Ã¢ÂÂcorky-bark acacia

Vachellia ditricha (Pedley) Kodela

Vachellia dolichocephala (Harms) Kyal. & Boatwr.

Vachellia douglasica (Pedley) Kodela

Vachellia drepanolobium (Harms ex SjÃÂ¶stedt) P.J.H. HurterÃ¢ÂÂwhistling thorn

Vachellia dyeri (P.P.Swartz) Kyal. & Boatwr.

Vachellia eburnea (L.f.) P. Hurter & Mabb.

Vachellia ebutsiniorum (P.J.H. Hurter) Kyal. & Boatwr.Ã¢ÂÂEbutsini acacia

Vachellia edgeworthii (T.Anders.) Kyal. & Boatwr.

Vachellia elatior (Brenan) Kyal. & Boatwr.

subsp. elatior (Brenan) Kyal. & Boatwr.

subsp. turkanae (Brenan) Kyal. & Boatwr.

Vachellia erioloba (E.Mey.) P.J.H. HurterÃ¢ÂÂcamel thorn (Kameeldoring)

Vachellia erythrophloea (Brenan) Kyal. & Boatwr.

Vachellia etbaica (Schweinf.) Kyal. & Boatwr.Ã¢ÂÂsavannah thorn

subsp. australis (Brenan) Kyal. & Boatwr.

subsp. etbaica (Schweinf.) Kyal. & Boatwr.

subsp. platycarpa (Brenan) Kyal. & Boatwr.

subsp. uncinata (Brenan) Kyal. & Boatwr.

Vachellia exuvialis (Verdoorn) Kyal. & Boatwr.Ã¢ÂÂflaky-bark acacia

Vachellia farnesiana (L.) Wight & Arn.Ã¢ÂÂhuisache

var. farnesiana (L.) Wight & Arn.

var. guanacastensis (H.D.Clarke et al.) Wight & Arn.

var. minuta (M.E.Jones) Wight & Arn.

var. pinetorum (F.J.Herm.) Wight & Arn.Ã¢ÂÂpineland wattle

Vachellia fischeri (Harms) Kyal. & Boatwr.Ã¢ÂÂflat-topped thorn

Vachellia flava (Forssk.) Kyal. & Boatwr.

Vachellia gentlei (Standley) Seigler & EbingerÃ¢ÂÂgentle acacia

Vachellia gerrardii (Benth.) P.J.H. HurterÃ¢ÂÂred acacia

var. calvescens (Brenan) P.J.H. Hurter

var. gerrardii (Benth.) P.J.H. Hurter

var. latisiliqua (Brenan) P.J.H. Hurter

Vachellia glandulifera (S. Watson) Seigler & Ebinger

Vachellia globulifera (Saff.) Seigler & EbingerÃ¢ÂÂglobular acacia

Vachellia grandicornuta (Gerstner) Seigler & EbingerÃ¢ÂÂhorned-thorn acacia

Vachellia guanacastensis (Clark, Seigler, & Ebinger) Seigler & Ebinger

Vachellia gummifera (Willd.) Kyal. & Boatwr.Ã¢ÂÂgum-bearing acacia

Vachellia haematoxylon (Willd.) Seigler & EbingerÃ¢ÂÂgray camel thorn, giraffe thorn

Vachellia harmandiana (Pierre) Maslin, Seigler & Ebinger

Vachellia hebeclada (DC.) Kyal. & Boatwr.Ã¢ÂÂcandle-pod acacia

subsp. chobiensis (O.B.Miller) Kyal. & Boatwr.

subsp. hebeclada (DC.) Kyal. & Boatwr.

subsp. tristis (A.Schreiber) Kyal. & Boatwr.

Vachellia hindsii (Benth.) Seigler & EbingerÃ¢ÂÂHinds acacia

Vachellia hockii (De Wild.) Seigler & Ebinger

Vachellia horrida (L.) Kyal. & Boatwr.Ã¢ÂÂlong white-galled acacia

subsp. benadirensis (Chiov.) Kyal. & Boatwr.

subsp. horrida (L.) Kyal. & Boatwr.

Vachellia hydaspica (J.R. Drumm. ex R. Parker) Ali

Vachellia inopinata (Prain) Maslin, Seigler & Ebinger

Vachellia insulae-iacobi (L. Riley) Seigler & Ebinger

Vachellia jacquemontii (Benth.) aliÃ¢ÂÂbaonÃ¡Â¸Â·ÃÂ«, raati-banwali

Vachellia janzenii (Ebinger & Seigler) Seigler & EbingerÃ¢ÂÂJanzen acacia

Vachellia karroo (Hayne) Banfi & GalassoÃ¢ÂÂKarroo Bush

Vachellia kingii (Prain) Maslin, Seigler & Ebinger

Vachellia kirkii (Oliv.) Kyal. & Boatwr.Ã¢ÂÂflood plain acacia

subsp. kirkii (Oliv.) Kyal. & Boatwr.

var. kirkii (Oliv.) Kyal. & Boatwr.

var. sublaevis (Brenan) Kyal. & Boatwr.

subsp. mildbraedii (Harms) Kyal. & Boatwr.

Vachellia koltermanii R. GarcÃÂa, M. MejÃÂa, Ebinger, & Seigler

Vachellia kosiensis (P.P.Sw. ex Coates Palgr.) Kyal. & Boatwr.Ã¢ÂÂdune acacia, dune sweet-thorn

Vachellia lahai (Steud. & Hochst. ex. Benth.) Kyal. & Boatwr.Ã¢ÂÂred-thorn acacia

Vachellia lasiopetala (Oliv.) Kyal. & Boatwr.

Vachellia latispina (J.E.Burrows & S.M.Burrows) Kyal. & Boatwr.

Vachellia leucophloea (Roxb.) Maslin, Seigler & EbingerÃ¢ÂÂpilang

var. leucophloea (Roxb.) Maslin, Seigler & Ebinger

var. microcephala (Kurz) Maslin, Seigler & Ebinger

Vachellia leucospira (Brenan) Kyal. & Boatwr.

Vachellia luederitzii (Engl.) Kyal. & Boatwr.Ã¢ÂÂbastard umbrella thorn

var. luederitzii (Engl.) Kyal. & Boatwr.Ã¢ÂÂKalahari-sand acacia

var. retinens (Sim) Kyal. & Boatwr.Ã¢ÂÂballoon-thorn acacia

Vachellia macracantha (Humb. & Bonpl. ex Willd.) Seigler & EbingerÃ¢ÂÂlongspine acacia, French casha, long-spine acacia, porknut, cambrÃÂ³n, long-spined acacia, (Jamaica) parknut, (Virgin Islands) wild tamarind, (Netherlands Antilles) Creole casha, Spanish casha, steel acacia, (Virgin Islands) stink casha, strink casha

Vachellia macrothyrsa (Harms) Kyal. & Boatwr.

Vachellia malacocephala (Harms) Kyal. & Boatwr.Ã¢ÂÂblack-galled acacia

Vachellia mayana (Lundell) Seigler & EbingerÃ¢ÂÂMaya Acacia

Vachellia mbuluensis (Brenan) Kyal. & Boatwr.Ã¢ÂÂhairy-galled acacia

Vachellia melanoceras (Beurl.) Seigler & EbingerÃ¢ÂÂblackthorn acacia, bullhorn acacia

Vachellia montana (P.P.Swartz) Kyal. & Boatwr.

Vachellia myaingii (Lace) Maslin, Seigler & Ebinger

Vachellia myrmecophila (R.Vig.) Boatwr.

Vachellia natalitia (E.Mey.) Kyal. & Boatwr.Ã¢ÂÂpale-bark acacia, pale-bark sweet thorn

Vachellia nebrownii (Burtt Davy) Seigler & EbingerÃ¢ÂÂwater acacia, water thorn

Vachellia negrii (Pichi-Sermolli) Kyal. & Boatwr.

Vachellia nilotica (L.) P.J.H. Hurter & Mabb.Ã¢ÂÂscented-pod acacia, gum Arabic tree, babul, Amrad gum, thorny mimosa of India

subsp. adstringens (Schumach. & Thonn.) P.J.H. Hurter & Mabb.

subsp. cupressiformis (J.L.Stewart) P.J.H. Hurter & Mabb.

subsp. hemispherica (Ali & Faruqi) P.J.H. Hurter & Mabb.

subsp. indica (Benth.) P.J.H. Hurter & Mabb.Ã¢ÂÂBabul, prickly acacia

subsp. kraussiana (Benth.) P.J.H. Hurter & Mabb.

subsp. leiocarpa (Brenan) P.J.H. Hurter & Mabb.

subsp. nilotica (L.) P.J.H. Hurter & Mabb.

subsp. subalata (Vatke) P.J.H. Hurter & Mabb.

subsp. tomentosa (Benth.) P.J.H. Hurter & Mabb.

Vachellia nubica (Benth.) Kyal. & Boatwr.

Vachellia oerfota (Forssk) Kyal. & Boatwr.

var. brevifolia (Boulos) Kyal. & Boatwr.

var. oerfota (Forssk) Kyal. & Boatwr.

Vachellia origena (Hunde) Kyal. & Boatwr.

Vachellia ormocarpoides (P.J.H. Hurter) Kyal. & Boatwr.Ã¢ÂÂLeolo thorn

Vachellia oviedoensis (R. GarcÃÂa & M. MejÃÂa) Seigler & Ebinger

Vachellia pacensis (Rudd & Carter) Seigler & Ebinger

Vachellia planifrons Wight & Arn.

Vachellia pachyphloia (W. Fitzg.) Kodela

subsp. brevipinnula (Tindale & Kodela) Kodela

subsp. pachyphloia (W. Fitzg.) Kodela

Vachellia pallidifolia (Tindale) Kodela

Vachellia paolii (Chiov.) Kyal. & Boatwr.

subsp. paolii (Chiov.) Kyal. & Boatwr.

subsp. paucijuga (Brenan) Kyal. & Boatwr.

Vachellia pennatula (Schltdl. & Cham.) Seigler & EbingerÃ¢ÂÂfeather acacia

var. parvicephala Seigler & Ebinger

var. pennatula (Schltdl. & Cham.) Seigler & Ebinger

Vachellia permixta (Burtt Davy) Kyal. & Boatwr.Ã¢ÂÂslender acacia

Vachellia pilispina (Pichi-Sermolli) Kyal. & Boatwr.Ã¢ÂÂmpande

Vachellia polypyrigenes (Greenm.) Seigler & Ebinger

Vachellia prasinata (Hunde) Kyal. & Boatwr.

Vachellia pringlei (Rose) Seigler & EbingerÃ¢ÂÂPringle acacia

var. californica Ã¢ÂÂCalifornia Pringle acacia

var. pringlei (Rose) Seigler & EbingerÃ¢ÂÂtypical Pringle acacia

Vachellia pseudofistula (Harms) Kyal. & Boatwr.Ã¢ÂÂant-galled acacia

Vachellia qandalensis (Thulin) Kyal. & Boatwr.

Vachellia quintanilhae (Torre) Kyal. & Boatwr.

Vachellia reficiens (Wawra) Kyal. & Boatwr.Ã¢ÂÂred-bark acacia

subsp. misera (Vatke) Kyal. & Boatwr.

subsp. reficiens (Wawra) Kyal. & Boatwr.

Vachellia rehmanniana (Schinz) Kyal. & Boatwr.Ã¢ÂÂsilky acacia

Vachellia rigidula (Benth.) Seigler & EbingerÃ¢ÂÂblackbrush acacia, blackbrush

Vachellia robbertsei (P.P.Swartz) Kyal. & Boatwr.Ã¢ÂÂSekhukhune acacia

Vachellia robusta (Burch.) Kyal. & Boatwr.Ã¢ÂÂsplendid acacia

subsp. clavigera (E.Mey.) Kyal. & Boatwr.Ã¢ÂÂriver acacia

subsp. robusta (Burch.) Kyal. & Boatwr.Ã¢ÂÂrobust acacia

subsp. usambarensis (Taub.) Kyal. & Boatwr.

Vachellia roigii (LÃÂ©on) Seigler & EbingerÃ¢ÂÂRoig acacia

Vachellia rorudiana (Christopherson) Seigler & EbingerÃ¢ÂÂGalapagos acacia

Vachellia ruddiae (D. H. Janzen) Seigler & EbingerÃ¢ÂÂRudd acacia

Vachellia schaffneri (S. Watson) Seigler & EbingerÃ¢ÂÂSchaffner's wattle, twisted acacia

var. bravoensis (Isely) Seigler & Ebinger

var. schaffneri (S. Watson) Seigler & Ebinger

Vachellia schottii (Torr.) Seigler & EbingerÃ¢ÂÂSchott's wattle

Vachellia sekhukhuniensis (P.J.H. Hurter) Kyal. & Boatwr.Ã¢ÂÂSekhukhune thorn

Vachellia seyal (Delile) P.J.H. HurterÃ¢ÂÂwhite whistling thorn

var. fistula (Schweinf.) P.J.H. Hurter

var. seyal (Delile) P.J.H. Hurter

Vachellia siamensis (Craib) Maslin, Seigler & Ebinger

Vachellia sieberiana (DC.) Kyal. & Boatwr.Ã¢ÂÂlongpod thorn, false paperbark thorn

var. sieberiana (DC.) Kyal. & Boatwr.

var. villosa (A.Chev.) Kyal. & Boatwr.

var. woodii (Burtt Davy) Kyal. & Boatwr.Ã¢ÂÂpaperbark acacia, paperbark thorn

Vachellia sphaerocephala (Schltdl. & Cham.) Seigler & EbingerÃ¢ÂÂroundhead acacia, bee wattle

Vachellia stuhlmannii (Taub.) Kyal. & Boatwr.Ã¢ÂÂolive-barked thorn, vlei acacia

Vachellia suberosa (A. Cunn. ex Benth.) KodelaÃ¢ÂÂcorkybark wattle

Vachellia sutherlandii (F. Muell.) KodelaÃ¢ÂÂcorkwood wattle

Vachellia swazica (Burtt Davy) Kyal. & Boatwr.Ã¢ÂÂSwazi acacia

Vachellia tenuispina (Verdoorn) Kyal. & Boatwr.Ã¢ÂÂturf acacia

Vachellia tephrophylla (Thulin) Kyal. & Boatwr.

Vachellia theronii (P.P.Sw.) Boatwr.Ã¢ÂÂslender mountain thorn

Vachellia tomentosa (Rottler) Maslin, Seigler & EbingerÃ¢ÂÂklampis

Vachellia torrei (Brenan) Kyal. & Boatwr.Ã¢ÂÂMozambique sticky thorn

Vachellia tortilis (Forssk.) Galasso & BanfiÃ¢ÂÂumbrella thorn, umbrella acacia

subsp. heteracantha (Burch.) Galasso & Banfi

subsp. raddiana (Savi) Galasso & Banfi

var. pubescens (A.Chev.) Galasso & Banfi

var. raddiana (Savi) Galasso & Banfi

subsp. spirocarpa (Hochst. ex A.Rich.) Galasso & Banfi

var. crinita (Chiov.) Galasso & Banfi

var. spirocarpa (Hochst. ex. A.Rich.) Galasso & Banfi

subsp. tortilis (Forssk.) Galasso & Banfi

Vachellia tortuosa (L.) Seigler & EbingerÃ¢ÂÂtwisted acacia, acacia bush, casia, catclaw, Dutch casha, huisachillo, Rio Grande acacia, sweet briar, sweet-briar, wild poponax

Vachellia turnbulliana (Brenan) Kyal. & Boatwr.Ã¢ÂÂvelvet pod acacia

Vachellia valida (Tindale & Kodela) Kodela

Vachellia vernicosa (Britton & Rose) Seigler & EbingerÃ¢ÂÂviscid acacia

Vachellia viguieri (Villiers & Du Puy) Boatwr.

Vachellia villaregalis (McVaugh) Seigler & Ebinger

Vachellia walwalensis (Gilliland) Kyal. & Boatwr.

Vachellia xanthophloea (Benth.) P.J.H. HurterÃ¢ÂÂfever tree

Vachellia zanzibarica (S.Moore) Kyal. & Boatwr.Ã¢ÂÂcoastal whistling thorn

var. microphylla (Brenan) Kyal. & Boatwr.

var. zanzibarica (S.Moore) Kyal. & Boatwr.

Vachellia zapatensis (Urb. & Ekman) Seigler & Ebinger

Incertae sedis

These species are suspected to belong to Vachellia, but have not been formally transferred.

Acacia callicoma Meisn.

Acacia harala Thulin & Gifri

Acacia hunteri Oliv.

Acacia johnwoodii Boulos

Acacia planifrons Koenig ex Wight & Arn.

Acacia pseudo-eburnea J.R. Drumm. ex Dunn

Acacia tanjorensis Ragup., Thoth. & Mahad.

Acacia yemenensis Boulos

Hybrids

Vachellia ÃÂ cedilloi (Rico Arce) Seigler & Ebinger

Vachellia campechiana ÃÂ pennatula

Vachellia erioloba ÃÂ haematoxylon

Vachellia ÃÂ gladiata (Saff.) Seigler & Ebinger

Vachellia kirkii ÃÂ seyal

Vachellia macracantha ÃÂ pennatula

Vachellia seyal var. fistula ÃÂ xanthophloea

Vachellia ÃÂ standleyi (Saff.) Seigler & Ebinger

References