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Umenocoleidae

Umenocoleidae is an extinct family of dictyopteran insects known from the Cretaceous. They are considered to be closely related to the Alienopteridae. They were originally considered to be beetles due to their beetle-like morphology, with sclerotised elytra-like forewings. This was probably an adaptation for living under bark and in other tight spaces.

Systematics

After

  • †Umenocoleus <small>Chen and Tan 1973</small> Dalazi Formation, China, Early Cretaceous (Aptian) Zhonggou Formation, China, Early Cretaceous (Albian), Jinju Formation, South Korea, Early Cretaceous (Albian)
  • †Ponopterix <small>VrÅ¡anský and Grimaldi 1999</small> Crato Formation, Brazil, Early Cretaceous (Aptian)
  • †Blattapterix <small>VrÅ¡anský 2003</small> Xiagou Formation, China, Early Cretaceous (Aptian)
  • †Elytropterix <small>VrÅ¡anský 2003</small> Dzun-Bain Formation, Mongolia, Aptian
  • †Petropterix <small>VrÅ¡anský 2003</small> Dzun-Bain Formation, Mongolia, Zaza Formation, Russia, Kitadani Formation, Japan, Early Cretaceous (Aptian) Zhonggou Formation, China, Early Cretaceous (Albian), Jinju Formation, South Korea, Early Cretaceous (Albian)
  • †Compunctiotypus <small>Kaddumi 2005</small> Jordanian amber, Early Cretaceous (Albian)
  • †Umenopterix <small>Lee 2016</small> Crato Formation, Brazil, Early Cretaceous (Aptian) Burmese amber, Myanmar, mid Cretaceous (latest Albian-earliest Cenomanian)
  • †Alienopterix <small>Mlynský, VrÅ¡anský et Wang, 2018</small> Burmese amber, Myanmar, mid Cretaceous (latest Albian-earliest Cenomanian)
  • †Enervipraeala <small>Luo, Xu et Jarzembowski, 2021</small> Burmese amber, Myanmar, mid Cretaceous (latest Albian-earliest Cenomanian)
  • †Laticephalana <small>Luo, Beutel, Xu et Jarzembowski, 2021</small> Burmese amber, Myanmar, mid Cretaceous (latest Albian-earliest Cenomanian)
  • †Archaeospinapteryx <small>Sendi et al. 2023</small> Burmese amber, Myanmar, mid Cretaceous (latest Albian-earliest Cenomanian)
  • †Trapezionotum <small>Sendi et al. 2023</small> Burmese amber, Myanmar, mid Cretaceous (latest Albian-earliest Cenomanian)
  • †Poikiloprosopon <small>Sendi et al. 2023</small> Burmese amber, Myanmar, mid Cretaceous (latest Albian-earliest Cenomanian)
  • †?Vzrkadlenie Burmese amber, Myanmar, mid Cretaceous (latest Albian-earliest Cenomanian) (Also placed in Alienopteridae or Cratovitismidae)
  • †Nigropterix <small>Sendi et al. 2023</small> Kzyl-Zhar, Kazakhstan, Late Cretaceous (Turonian)
  • †Umenotypus <small>Sendi et al. 2023</small> Kzyl-Zhar, Kazakhstan, Late Cretaceous (Turonian)
  • †Pseudoblattapterix <small>Lee et al. 2024</small> Jinju Formation, South Korea, Early Cretaceous (Albian)

Jantaropterix <small>Vršanský and Grimaldi 2003</small>, Perspicuus <small>Koubová, 2020</small>, Lepidopterix <small>Sendi, 2020</small> and Antophiloblatta <small>Sendi, 2020</small> have been transferred to Cratovitismidae, while Vitisma <small>Vršanský 1999</small> and Permoponopterix <small>Nel, Prokop et Kirejtshuk, 2014</small> have been excluded from the family.

References