Tomarus

Tomarus is a genus of scarab beetles in the subfamily Dynastinae, the rhinoceros beetles. They are native to the Americas, where they are distributed from the central United States to Argentina, and a few species occur in the Caribbean.

The adult beetles are nocturnal and attracted to lights; the larvae generally remain in the soil, often feeding on plant roots.

In order to identify species, the parameres of the male must be pulled out of the abdomen and examined. This process should be done carefully, because the parameres are quite fragile, "almost parchment-like".

Species

The following species belong to the genus Tomarus:

• Tomarus adoceteus <small>Ratcliffe & Cave, 2010</small><span style="color:gray">^{c g}</span>
• Tomarus bidentulus <small>(Fairmaire, 1892)</small><span style="color:gray">^c</span>
• Tomarus bituberculatus <small>(Palisot De Beauvois, 1805)</small><span style="color:gray">^{c g}</span>
• Tomarus burmeisteri <small>(Steinheil, 1872)</small><span style="color:gray">^{c g}</span>
• Tomarus cicatricosus <small>(Prell, 1937)</small><span style="color:gray">^{c g}</span>
• Tomarus colombianus <small>Lopez-Garcia & Gasca-Alvarez, 2014</small><span style="color:gray">^g</span>
• Tomarus columbianus <small>López-García & Gasca-Álvarez, 2014</small><span style="color:gray">^{c g}</span>
• Tomarus compositus <small>(Wickham, 1911)</small><span style="color:gray">^{c g}</span>
• Tomarus cuniculus <small>(Fabricius, 1801)</small><span style="color:gray">^{i c g b}</span>
• Tomarus discrepans <small>Escalona & Joly, 2006</small><span style="color:gray">^{c g}</span>
• Tomarus ebenus <small>(Degeer, 1774)</small><span style="color:gray">^{c g}</span> = type species
• Tomarus effetus <small>(Wickham, 1914)</small><span style="color:gray">^{c g}</span>
• Tomarus formosianus <small>Grouvelle, 1913</small><span style="color:gray">^g</span>
• Tomarus fossor <small>(Latreille, 1833)</small><span style="color:gray">^{c g}</span>
• Tomarus gyas <small>Erichson, 1848</small><span style="color:gray">^{c g}</span>
• Tomarus laevicollis <small>(Bates, 1888)</small><span style="color:gray">^{i c g}</span>
• Tomarus maimon <small>Erichson, 1847</small><span style="color:gray">^{c g}</span>
• Tomarus maternus <small>(Prell, 1937)</small><span style="color:gray">^{c g}</span>
• Tomarus nasutus <small>(Burmeister, 1847)</small><span style="color:gray">^{i c g}</span>
• Tomarus neglectus <small>(LeConte, 1847)</small><span style="color:gray">^{i c g b}</span>
• Tomarus peruvianus <small>(Endrödi, 1970)</small><span style="color:gray">^c</span>
• Tomarus pullus <small>(Prell, 1937)</small><span style="color:gray">^{c g}</span>
• Tomarus pumilus <small>(Prell, 1937)</small><span style="color:gray">^{c g}</span>
• Tomarus roigjunenti <small>Neita & Ratcliffe, 2017</small><span style="color:gray">^{c g}</span>
• Tomarus rosettae <small>(Endrödi, 1968)</small><span style="color:gray">^{c g}</span>
• Tomarus rostratus <small>Dupuis, 2014</small><span style="color:gray">^{c g}</span>
• Tomarus rubripes <small>(Boheman, 1858)</small><span style="color:gray">^{c g}</span>
• Tomarus sallaei <small>(Bates, 1888)</small><span style="color:gray">^{i c g b}</span>
• Tomarus selanderi <small>(Cartwright, 1959)</small><span style="color:gray">^{i c g}</span>
• Tomarus similis <small>(Endrödi, 1968)</small><span style="color:gray">^{c g}</span>
• Tomarus spinipenis <small>Neita & Ratcliffe, 2017</small><span style="color:gray">^{c g}</span>
• Tomarus subtropicus <small>(Blatchley, 1922)</small><span style="color:gray">^{i c g b}</span> (sugarcane grub)
• Tomarus villosus <small>(Burmeister, 1847)</small><span style="color:gray">^{c g}</span>

<small>Data sources: i = ITIS, c = Catalogue of Life, g = GBIF, b = Bugguide.net</small>

• Tomarus gibbosus <small>(LeConte, 1856)</small>, the carrot beetle
• Tomarus relictus <small>(Say, 1825)</small>

References