Tmarus is a genus of crab spiders in the family Thomisidae. The genus was established by Eugène Simon in 1875, with Tmarus piger <small>(Walckenaer, 1802)</small> as the type species.
The genus includes more than 230 described species distributed across all continents except Antarctica.
Taxonomy
The genus Tmarus has undergone several taxonomic revisions. The generic name Martus <small>Mello-Leitão, 1943</small> was considered a nomen nudum and synonymized with Tmarus. Additionally, Peritraeus <small>Simon, 1895</small> was found to be a junior synonym of Tmarus.
Description
Tmarus spiders are small to moderate-sized crab spiders with body lengths ranging from 3.7 to 7.3 millimeters. Females are typically larger than males, with males being slightly smaller in size. The spiders display cryptic coloration ranging from cream to dull grey, often mottled with cream, brown or black patterns that resemble bark, providing excellent camouflage.
The cephalothorax is as wide as long and rather convex, declining abruptly towards the posterior border. The clypeus is porrect (projecting forward), and the ocular area is well elevated. The lateral eyes are situated on well-defined tubercles, with both eye rows appearing almost straight or only slightly recurved. The lateral eyes are larger than the median eyes, while the anterior median eyes are the smallest.
The opisthosoma often features a posterior tubercle dorsally, though some species may have an elongated body form. The abdomen is clothed with numerous short, stiff setae, which are sometimes situated on small tubercles. The legs are spinous, with legs I and II being almost the same length and longer than legs III and IV. In Tmarus species, the posterior median eyes (PME) are situated nearer to each other than to the posterior lateral eyes (PLE), which distinguishes them from the closely related genus Monaeses.
Distribution and habitat
Tmarus species are found worldwide, with the highest diversity in tropical and subtropical regions. They typically inhabit vegetation, where they hunt by ambush rather than building webs. Many species are found on bark or leaves, where their cryptic coloration provides camouflage.
Species
, this genus includes 230 species and two subspecies.
These species have articles on Wikipedia:
- Tmarus africanus <small>Lessert, 1919</small> â Tanzania, Botswana, South Africa
- Tmarus angulatus <small>(Walckenaer, 1837)</small> â North America
- Tmarus cameliformis <small>Millot, 1942</small> â Ghana, Guinea, DR Congo, Tanzania, Malawi, Botswana, South Africa, Eswatini
- Tmarus cancellatus <small>Thorell, 1899</small> â Ivory Coast, Gabon, Nigeria, Cameroon, Rep. Congo, Equatorial Guinea (Bioko), Zimbabwe, South Africa
- Tmarus comellinii <small>Garcia-Neto, 1989</small> â Congo to South Africa
- Tmarus floridensis <small>Keyserling, 1884</small> â United States
- Tmarus foliatus <small>Lessert, 1928</small> â Senegal, Ivory Coast, DR Congo, Burundi, Tanzania, Zimbabwe, South Africa, Lesotho, Comoros
- Tmarus guineensis <small>Millot, 1942</small> â Guinea, South Africa
- Tmarus hystrix <small>(Simon, 1895)</small> â India, Sri Lanka
- Tmarus longicaudatus <small>Millot, 1942</small> â Mauritania, Niger, Namibia, South Africa, Yemen, Saudi Arabia, United Arab Emirates
- Tmarus minutus <small>Banks, 1904</small> â United States
- Tmarus natalensis <small>Lessert, 1925</small> â South Africa
- Tmarus piger <small>(Walckenaer, 1802)</small> â Europe, Turkey, Caucasus, Russia (Europe to Far East), Kazakhstan, China, Korea, Japan (type species)
- Tmarus planetarius <small>Simon, 1903</small> â Guinea Bissau, Sierra Leone, Ivory Coast, Nigeria, Cameroon, DR Congo, Kenya, Malawi, South Africa
- Tmarus aberrans <small>Mello-Leitão, 1944</small> â Brazil
- Tmarus aculeatus <small>Chickering, 1950</small> â Panama
- Tmarus africanus <small>Lessert, 1919</small> â Tanzania, Botswana, South Africa
- Tmarus albidus <small>(L. Koch, 1876)</small> â Australia (Queensland)
- Tmarus albifrons <small>Piza, 1944</small> â Brazil
- Tmarus albisterni <small>Mello-Leitão, 1942</small> â Argentina
- Tmarus albolineatus <small>Keyserling, 1880</small> â Brazil
- Tmarus alticola <small>Mello-Leitão, 1929</small> â Brazil
- Tmarus amazonicus <small>Mello-Leitão, 1929</small> â Brazil
- Tmarus ampullatus <small>Soares, 1943</small> â Brazil
- Tmarus angulatus <small>(Walckenaer, 1837)</small> â North America
- Tmarus angulifer <small>Simon, 1895</small> â Australia (Queensland)
- Tmarus aporus <small>Soares & Camargo, 1948</small> â Brazil
- Tmarus atypicus <small>Mello-Leitão, 1929</small> â Brazil
- Tmarus australis <small>Mello-Leitão, 1941</small> â Argentina
- Tmarus baptistai <small>Silva-Moreira, 2010</small> â Brazil
- Tmarus bedoti <small>Lessert, 1928</small> â DR Congo
- Tmarus berlandi <small>Lessert, 1928</small> â DR Congo
- Tmarus bifasciatus <small>Mello-Leitão, 1929</small> â Peru, Brazil
- Tmarus bifidipalpus <small>Mello-Leitão, 1943</small> â Brazil
- Tmarus biocellatus <small>Mello-Leitão, 1929</small> â Brazil
- Tmarus bisectus <small>Piza, 1944</small> â Brazil
- Tmarus borgmeyeri <small>Mello-Leitão, 1929</small> â Brazil
- Tmarus bucculentus <small>Chickering, 1950</small> â Panama
- Tmarus byssinus <small>Tang & Li, 2009</small> â China
- Tmarus caeruleus <small>Keyserling, 1880</small> â Brazil
- Tmarus cameliformis <small>Millot, 1942</small> â Ghana, Guinea, DR Congo, Tanzania, Malawi, Botswana, South Africa, Eswatini
- Tmarus camellinus <small>Mello-Leitão, 1929</small> â Brazil
- Tmarus cancellatus <small>Thorell, 1899</small> â Ivory Coast, Gabon, Nigeria, Cameroon, Rep. Congo, Equatorial Guinea (Bioko), Zimbabwe, South Africa
- T. c. congoensis <small>Comellini, 1955</small> â DR Congo
- Tmarus candefactus <small>Caporiacco, 1954</small> â French Guiana
- Tmarus candidissimus <small>Mello-Leitão, 1947</small> â Brazil
- Tmarus caporiaccoi <small>Comellini, 1955</small> â DR Congo
- Tmarus caretta <small>Mello-Leitão, 1929</small> â Brazil
- Tmarus caxambuensis <small>Mello-Leitão, 1929</small> â Brazil
- Tmarus cinerasceus <small>(L. Koch, 1876)</small> â Australia (Queensland)
- Tmarus cinereus <small>Mello-Leitão, 1929</small> â Brazil, Guyana
- Tmarus circinalis <small>Song & Chai, 1990</small> â China
- Tmarus clarus <small>(Mello-Leitão, 1937)</small> â Brazil
- Tmarus clavimanus <small>Mello-Leitão, 1929</small> â Brazil
- Tmarus clavipes <small>Keyserling, 1891</small> â Brazil
- Tmarus cognatus <small>Chickering, 1950</small> â Panama
- Tmarus comellinii <small>Garcia-Neto, 1989</small> â Congo to South Africa
- Tmarus contortus <small>Chickering, 1950</small> â Panama
- Tmarus corruptus <small>O. Pickard-Cambridge, 1892</small> â Mexico, Panama
- Tmarus craneae <small>Chickering, 1966</small> â Trinidad
- Tmarus cretatus <small>Chickering, 1965</small> â Panama
- Tmarus curvus <small>Chickering, 1950</small> â Panama
- Tmarus decens <small>O. Pickard-Cambridge, 1892</small> â Panama
- Tmarus decoloratus <small>Keyserling, 1883</small> â Peru
- Tmarus decorus <small>Chickering, 1965</small> â Panama
- Tmarus dejectus <small>(O. Pickard-Cambridge, 1885)</small> â India
- Tmarus digitatus <small>Mello-Leitão, 1929</small> â Brazil
- Tmarus digitiformis <small>Yang, Zhu & Song, 2005</small> â China
- Tmarus dostinikus <small>Barrion & , 1995</small> â Philippines
- Tmarus ehecatltocatl <small>Jiménez, 1992</small> â Mexico
- Tmarus elongatus <small>Mello-Leitão, 1929</small> â Brazil
- Tmarus eques <small>Thorell, 1890</small> â Indonesia (Java)
- Tmarus espiritosantensis <small>B. A. M. Soares & H. E. M. Soares, 1946</small> â Brazil
- Tmarus estylifer <small>Mello-Leitão, 1929</small> â Brazil
- Tmarus fallax <small>Mello-Leitão, 1929</small> â Brazil, Guyana
- Tmarus fanjing <small>Yang & Yu, 2022</small> â China
- Tmarus farri <small>Chickering, 1966</small> â Jamaica
- Tmarus fasciolatus <small>Simon, 1906</small> â India
- Tmarus femellus <small>Caporiacco, 1941</small> â Ethiopia
- Tmarus floridensis <small>Keyserling, 1884</small> â United States
- Tmarus foliatus <small>Lessert, 1928</small> â Senegal, Ivory Coast, DR Congo, Burundi, Tanzania, Zimbabwe, South Africa, Lesotho, Comoros
- Tmarus formosus <small>Mello-Leitão, 1917</small> â Brazil
- Tmarus gajdosi <small>Marusik & , 2002</small> â Mongolia
- Tmarus galapagosensis <small>Baert, 2013</small> â Galapagos
- Tmarus geayi <small>Caporiacco, 1954</small> â French Guiana
- Tmarus gladiatus <small>Tang & Li, 2010</small> â China
- Tmarus grandis <small>Mello-Leitão, 1929</small> â Brazil
- Tmarus guineensis <small>Millot, 1942</small> â Guinea, South Africa
- Tmarus hastatus <small>Tang & Li, 2009</small> â China
- Tmarus hazevensis <small>, 1973</small> â Israel
- Tmarus hirsutus <small>Mello-Leitão, 1929</small> â Brazil
- Tmarus histrix <small>Caporiacco, 1954</small> â French Guiana
- Tmarus hiyarensis <small>Ileperuma Arachchi & Benjamin, 2019</small> â Sri Lanka
- Tmarus holmbergi <small>Schiapelli & Gerschman, 1941</small> â Argentina
- Tmarus homanni <small>Chrysanthus, 1964</small> â Indonesia (New Guinea)
- Tmarus humphreyi <small>Chickering, 1965</small> â Panama
- Tmarus hystrix <small>(Simon, 1895)</small> â India, Sri Lanka
- Tmarus impedus <small>Chickering, 1965</small> â Panama
- Tmarus incertus <small>Keyserling, 1880</small> â Colombia
- Tmarus incognitus <small>Mello-Leitão, 1929</small> â Brazil
- Tmarus ineptus <small>O. Pickard-Cambridge, 1892</small> â Panama, Colombia, Bonaire
- Tmarus infrasigillatus <small>Mello-Leitão, 1947</small> â Brazil
- Tmarus innotus <small>Chickering, 1965</small> â Panama
- Tmarus innumus <small>Chickering, 1965</small> â Panama
- Tmarus insuetus <small>Chickering, 1966</small> â Trinidad
- Tmarus intentus <small>O. Pickard-Cambridge, 1892</small> â Guatemala, Panama
- Tmarus interritus <small>Keyserling, 1880</small> â Panama, Brazil
- Tmarus jabalpurensis <small>Gajbe & Gajbe, 1999</small> â India
- Tmarus jelskii <small>(Taczanowski, 1872)</small> â French Guiana
- Tmarus jocosus <small>O. Pickard-Cambridge, 1898</small> â Costa Rica
- Tmarus karolae <small>, 1964</small> â Ivory Coast
- Tmarus komi <small>Ono, 1996</small> â Japan (Ryukyu Is.)
- Tmarus koreanus <small>Paik, 1973</small> â China, Korea
- Tmarus kotigeharus <small>Tikader, 1963</small> â India
- Tmarus lanyu <small>Zhang, Zhu & Tso, 2006</small> â Taiwan
- Tmarus lapadui <small>Jézéquel, 1964</small> â Ivory Coast
- Tmarus latifrons <small>Thorell, 1895</small> â Myanmar, Indonesia (Krakatau)
- Tmarus lawrencei <small>Comellini, 1955</small> â DR Congo
- Tmarus levii <small>Chickering, 1965</small> â Panama
- Tmarus lichenoides <small>Mello-Leitão, 1929</small> â Brazil
- Tmarus littoralis <small>Keyserling, 1880</small> â Brazil
- Tmarus locketi <small>Millot, 1942</small> â Guinea, DR Congo
- T. l. djuguensis <small>Comellini, 1955</small> â DR Congo
- Tmarus longicaudatus <small>Millot, 1942</small> â Mauritania, Niger, Namibia, South Africa, Yemen, Saudi Arabia, United Arab Emirates
- Tmarus longipes <small>Caporiacco, 1947</small> â Tanzania
- Tmarus longqicus <small>Song & Zhu, 1993</small> â China
- Tmarus longus <small>Chickering, 1965</small> â Panama
- Tmarus loriae <small>Thorell, 1890</small> â Malaysia
- Tmarus macilentus <small>(L. Koch, 1876)</small> â Australia (Queensland)
- Tmarus maculosus <small>Keyserling, 1880</small> â Colombia
- Tmarus makiharai <small>Ono, 1988</small> â Japan (Ryukyu Is.)
- Tmarus malleti <small>Lessert, 1919</small> â Guinea, DR Congo, Tanzania
- Tmarus manojkaushalyai <small>Ileperuma Arachchi & Benjamin, 2019</small> â Sri Lanka
- Tmarus marmoreus <small>(L. Koch, 1876)</small> â Australia (Queensland)
- Tmarus menglae <small>Song & Zhao, 1994</small> â China
- Tmarus menotus <small>Chickering, 1966</small> â Jamaica
- Tmarus metropolitanus <small>Mello-Leitão, 1929</small> â Brazil
- Tmarus milloti <small>Comellini, 1955</small> â Cameroon, DR Congo
- Tmarus minensis <small>Mello-Leitão, 1929</small> â Brazil
- Tmarus minutus <small>Banks, 1904</small> â United States
- Tmarus misumenoides <small>Mello-Leitão, 1927</small> â Brazil
- Tmarus montericensis <small>Keyserling, 1880</small> â Peru
- Tmarus morosus <small>Chickering, 1950</small> â Panama, Colombia
- Tmarus mourei <small>Mello-Leitão, 1947</small> â Brazil
- Tmarus mundulus <small>O. Pickard-Cambridge, 1892</small> â Panama
- Tmarus mutabilis <small>Soares, 1944</small> â Brazil
- Tmarus natalensis <small>Lessert, 1925</small> â South Africa
- Tmarus neocaledonicus <small>Kritscher, 1966</small> â New Caledonia
- Tmarus nigrescens <small>Mello-Leitão, 1929</small> â Brazil
- Tmarus nigridorsi <small>Mello-Leitão, 1929</small> â Brazil
- Tmarus nigristernus <small>Caporiacco, 1947</small> â Uganda
- Tmarus nigrofasciatus <small>Mello-Leitão, 1929</small> â Brazil
- Tmarus nigroviridis <small>Mello-Leitão, 1929</small> â Brazil
- Tmarus ningshaanensis <small>Wang & Xi, 1998</small> â China
- Tmarus obesus <small>Mello-Leitão, 1929</small> â Brazil, French Guiana
- Tmarus oblectator <small>Logunov, 1992</small> â Russia (Far East)
- Tmarus obsecus <small>Chickering, 1965</small> â Panama
- Tmarus okinawanus <small>, 2024</small> â Japan (Ryukyu Is.)
- Tmarus orientalis <small>Schenkel, 1963</small> â China, Korea
- Tmarus pallidus <small>Mello-Leitão, 1929</small> â Brazil
- Tmarus parallelus <small>Mello-Leitão, 1943</small> â Brazil
- Tmarus parki <small>Chickering, 1950</small> â Panama
- Tmarus paulensis <small>Piza, 1935</small> â Brazil
- Tmarus pauper <small>O. Pickard-Cambridge, 1892</small> â Panama
- Tmarus perditus <small>Mello-Leitão, 1929</small> â Brazil
- Tmarus peregrinus <small>Chickering, 1950</small> â Panama
- Tmarus peruvianus <small>Berland, 1913</small> â Peru
- Tmarus piger <small>(Walckenaer, 1802)</small> â Europe, Turkey, Caucasus, Russia (Europe to Far East), Kazakhstan, China, Korea, Japan (type species)
- Tmarus piochardi <small>(Simon, 1866)</small> â Mediterranean, Turkey, Georgia, Iran
- Tmarus pizai <small>Soares, 1941</small> â Brazil
- Tmarus planetarius <small>Simon, 1903</small> â Guinea Bissau, Sierra Leone, Ivory Coast, Nigeria, Cameroon, DR Congo, Kenya, Malawi, South Africa
- Tmarus planifrons <small>Mello-Leitão, 1943</small> â Brazil
- Tmarus planquettei <small>Jézéquel, 1966</small> â Ivory Coast
- Tmarus pleuronotatus <small>Mello-Leitão, 1941</small> â Brazil
- Tmarus plurituberculatus <small>Mello-Leitão, 1929</small> â Brazil
- Tmarus polyandrus <small>Mello-Leitão, 1929</small> â Brazil
- Tmarus posticatus <small>Simon, 1929</small> â Brazil
- Tmarus primitivus <small>Mello-Leitão, 1929</small> â Brazil
- Tmarus probus <small>Chickering, 1950</small> â Panama
- Tmarus productus <small>Chickering, 1950</small> â Panama
- Tmarus prognathus <small>Simon, 1929</small> â Brazil
- Tmarus projectus <small>(L. Koch, 1876)</small> â Australia (Queensland)
- Tmarus protobius <small>Chickering, 1965</small> â Panama
- Tmarus pugnax <small>Mello-Leitão, 1929</small> â Brazil
- Tmarus pulchripes <small>Thorell, 1895</small> â Singapore
- Tmarus punctatissimus <small>(Simon, 1870)</small> â Portugal, Spain, France, Caucasus, Russia (Europe to Far East), Iran, Turkmenistan, Korea, Japan
- Tmarus punctatus <small>(Nicolet, 1849)</small> â Chile
- Tmarus qinlingensis <small>Song & Wang, 1994</small> â China
- Tmarus rainbowi <small>Mello-Leitão, 1929</small> â Australia (South Australia)
- Tmarus rarus <small>B. A. M. Soares & H. E. M. Soares, 1946</small> â Brazil
- Tmarus riccii <small>Caporiacco, 1941</small> â Ethiopia
- Tmarus rimosus <small>Paik, 1973</small> â Russia (Urals to Far East), China, Korea, Japan
- Tmarus rostratus <small>Y. C. Zhou, Qin, G. C. Zhou, Peng & Liu, 2025</small> â China
- Tmarus rubinus <small>Chickering, 1965</small> â Panama
- Tmarus rubromaculatus <small>Keyserling, 1880</small> â United States
- Tmarus salai <small>Schick, 1965</small> â United States
- Tmarus schoutedeni <small>Comellini, 1955</small> â DR Congo
- Tmarus scopullitus <small>Liu & Zhang, 2025</small> â China
- Tmarus semiroseus <small>Simon, 1909</small> â Vietnam
- Tmarus separatus <small>Banks, 1898</small> â Panama
- Tmarus serratus <small>Yang, Zhu & Song, 2005</small> â China
- Tmarus shimojanai <small>Ono, 1997</small> â Japan (Ryukyu Is.)
- Tmarus sigillatus <small>Chickering, 1950</small> â Panama
- Tmarus simoni <small>Comellini, 1955</small> â Sierra Leone
- Tmarus songi <small>Han & Zhu, 2009</small> â China
- Tmarus soricinus <small>Simon, 1906</small> â India
- Tmarus spicatus <small>Tang & Li, 2009</small> â China
- Tmarus spinosus <small>Comellini, 1955</small> â DR Congo
- Tmarus staintoni <small>(O. Pickard-Cambridge, 1873)</small> â Spain, France, Italy, Morocco, Algeria
- Tmarus stellio <small>Simon, 1875</small> â Southern, Central and Eastern Europe, Turkey, Caucasus, Iran
- Tmarus stolzmanni <small>Keyserling, 1880</small> â Peru
- Tmarus striolatus <small>Mello-Leitão, 1943</small> â Brazil
- Tmarus studiosus <small>O. Pickard-Cambridge, 1892</small> â Panama
- Tmarus subqinlingensis <small>R. Zhang & F. Zhang, 2023</small> â China
- Tmarus taibaiensis <small>Song & Wang, 1994</small> â China
- Tmarus taishanensis <small>Zhu & Wen, 1981</small> â Russia (South Siberia), China
- Tmarus taiwanus <small>Ono, 1977</small> â China, Taiwan
- Tmarus tamazolinus <small>Jiménez, 1988</small> â Mexico
- Tmarus thorelli <small>Comellini, 1955</small> â DR Congo
- Tmarus tinctus <small>Keyserling, 1880</small> â Peru
- Tmarus tonkinus <small>Simon, 1909</small> â Vietnam
- Tmarus toschii <small>Caporiacco, 1949</small> â Kenya
- Tmarus trifidus <small>Mello-Leitão, 1929</small> â Brazil
- Tmarus trituberculatus <small>Mello-Leitão, 1929</small> â Brazil
- Tmarus truncatus <small>(L. Koch, 1876)</small> â Australia (Queensland)
- Tmarus tuberculitibiis <small>Caporiacco, 1940</small> â Ethiopia
- Tmarus undatus <small>Tang & Li, 2009</small> â China
- Tmarus ungulatus <small>Chen, Liu & Hu, 2025</small> â China
- Tmarus unicus <small>Gertsch, 1936</small> â United States
- Tmarus vachoni <small>Millot, 1942</small> â Ivory Coast
- Tmarus variabilis <small>(L. Koch, 1876)</small> â Australia (Queensland)
- Tmarus variatus <small>Keyserling, 1891</small> â Brazil
- Tmarus verrucosus <small>Mello-Leitão, 1948</small> â Guyana
- Tmarus vertumus <small>Chickering, 1966</small> â Puerto Rico
- Tmarus vexillifer <small>(Butler, 1876)</small> â Mauritius (Rodriguez)
- Tmarus villasboasi <small>Mello-Leitão, 1949</small> â Brazil
- Tmarus viridis <small>Keyserling, 1880</small> â Peru, Brazil
- Tmarus viridomaculatus <small>Ileperuma Arachchi & Benjamin, 2019</small> â Sri Lanka
- Tmarus vitusus <small>Chickering, 1965</small> â Mexico, Panama
- Tmarus wiedenmeyeri <small>Schenkel, 1953</small> â Venezuela
- Tmarus yaginumai <small>Ono, 1977</small> â Korea, Japan
- Tmarus yani <small>Yin, Peng, Gong & Kim, 2004</small> â China
- Tmarus yerohamus <small>Levy, 1973</small> â Israel
- Tmarus yiminhensis <small>Zhu & Wen, 1981</small> â China
- Tmarus yueluensis <small>Y. C. Zhou, Qin, G. C. Zhou, Peng & Liu, 2025</small> â China
- Tmarus zhui <small>Sherwood & Li, 2021</small> â China
References
External links