Tityus is a large genus of thick-tailed scorpions (family Buthidae), the namesake of its subfamily Tityinae. As of 2021, Tityus contains more than 220 described species distributed in Central America and South America, from Costa Rica to Argentina. Species in the genus Tityus have been studied for hundreds of years, long before the taxonomic classification was put in place. Tityus tend to be of medium size for scorpions, roughly 50 to 70 millimeters long.ÃÂ They are dark brown or red in color, and can exhibit sexual dimorphism. They can live in a variety of environments, ranging from urban to arid mountains to the Amazon Rainforest. Tityus scorpions are best known for their venom and potent sting. The genus contains several dangerously venomous scorpions, the best known of which is the Brazilian yellow scorpion, T. serrulatus. Its venom can cause severe illness (including pancreatitis), and in the young, old and infirm even death. Some experts have argued that the genus as a whole may be paraphyletic, which could explain the knowledge gaps related to Tityus. ÃÂ
Geography
Scorpions in the genus Tityus can live in several distinct environments across South America. However, there appears to be a clear geographic distinction that exists between species. A species that inhabits the Andes Mountains will not also live in the Amazon Rainforest. In some South American countries, such as Argentina, the geographic range of Tityus scorpions is expanding. This creates a problem for the general public and healthcare. When scorpion stings become more frequent, it puts more strain on hospitals and healthcare facilities.ÃÂ
Venom
The genus Tityus is most well known for its venomous species. Tityus serrulatus venom contains a powerful neurotoxin that affects almost all anatomical body systems. The most dangerous species in the genus Tityus is serrulatus. The nature of their venom and its ability to impact the entire body make Tityus serrulatus a particularly dangerous species. However, their stings are not often lethal, which may be due to low venom mass injected. Young children and seniors are at a higher risk of death than the general population. Many factors are important for determining how dangerous a scorpion sting will be. Variables such as venom composition, location of the sting, and the overall health makeup of the victim in question play a role in determining the lethality of a sting. Scorpion stings are the most common cause of envenomation in Brazil, and are seen as a risk in urban environments. Scorpions in the genus Tityus have been studied by medical researchers for the purpose of identifying and understanding the toxins produced by various species.
Behavior
Little is yet understood about scorpion behavior in the genus Tityus. This is especially true of reproductive aspects, although some general patterns are known. Unlike the majority of arachnids that are oviparous, (i.e. hatching from eggs), in contrast all scorpions including those of the genus Tityus are viviparous, with live births. Notably, several species of Tityus (including Tityus serrulatus) are parthenogenic, a process that allows females to reproduce without fertilization. This survival strategy may contribute to their success in challenging environments.ÃÂ When in contrast the two sexes undergo a mating process, the females can use chemical communication to induce male courtship.ÃÂ After being exposed to compounds associated with female scorpions, the males will alter their activity and perform behaviors characteristic of courtship.ÃÂ
Tityus, like other scorpions use the stingers for both subduing prey items but also for defense, which is a behavior demonstrated by both sexes.ÃÂ Venom is energetically expensive to create, so scorpions only use their stinger defensively when seriously threatened.
Diet
Scorpions in the genus Tityus are carnivores.ÃÂ They feed on insects such as cockroaches and crickets.ÃÂ It is believed that members of the genus Tityus can survive for over a year without food, and even give birth after lengthy periods of starvation.ÃÂ This is another survival technique that helps them outlast unfavorable conditions.
Species
- Tityus aba <small>Candido, Lucas, de Souza, Diaz & Lira-da-Silva, 2005</small>
- Tityus abudi <small>Armas, 1999</small>
- Tityus acananensis <small>González-Sponga, 2009</small>
- Tityus adisi <small>Lourenço, 2002</small>
- Tityus adrianoi <small>Lourenço, 2003</small>
- Tityus ahincoi <small>González-Sponga, 2001</small>
- Tityus altithronus <small>Armas, 1999</small>
- Tityus anasilviae <small>Armas & Abud Antun, 2004</small>
- Tityus androcottoides <small>(Karsch, 1879)</small>
- Tityus anduzei <small>González-Sponga, 1997</small>
- Tityus angelesae <small>Santiago-Blay, 2009</small>
- Tityus anneae <small>Lourenço, 1997</small>
- Tityus antioquensis <small>Lourenço & Otero Patiño, 1998</small>
- Tityus apiacas <small>Lourenço, 2002</small>
- Tityus apozonalli <small>Riquelme, Villegas & González, 2015</small>
- Tityus arellanoparrai <small>González-Sponga, 1985</small>
- Tityus argentinus <small>Borelli, 1899</small>
- Tityus asthenes <small>Pocock, 1893</small>
- Tityus atriventer <small>Pocock, 1897</small>
- Tityus bahiensis <small>(Perty, 1833)</small>
- Tityus bahoruco <small>Teruel & Armas, 2006</small>
- Tityus barquisimetanus <small>González-Sponga, 1994</small>
- Tityus bastosi <small>Lourenço, 1984</small>
- Tityus bellulus <small>Armas, 1999</small>
- Tityus betschi <small>Lourenço, 1992</small>
- Tityus birabeni <small>Abalos, 1955</small>
- Tityus blanci <small>Lourenço, 1994</small>
- Tityus blaseri <small>Mello-Leitão, 1931</small>
- Tityus boconoensis <small>González-Sponga, 1981</small>
- Tityus bolivianus <small>Kraepelin, 1895</small>
- Tityus brazilae <small>Lourenço & Eickstedt, 1984</small>
- Tityus breweri <small>González-Sponga, 1997</small>
- Tityus cachipalensis <small>González-Sponga, 2002</small>
- Tityus caesarbarrioi <small>González-Sponga, 2001</small>
- Tityus canopensis <small>Lourenço, 2002</small>
- Tityus carabobensis <small>González-Sponga, 1987</small>
- Tityus carinatoides <small>Mello-Leitão, 1945</small>
- Tityus caripitensis <small>Quiroga, deSousa & Parrilla-Alvarez, 2000</small>
- Tityus carvalhoi <small>Mello-Leitão, 1945</small>
- Tityus cerroazul <small>Lourenço, 1986</small>
- Tityus championi <small>Pocock, 1898</small>
- Tityus charalaensis <small>Mello-Leitão, 1940</small>
- Tityus charreyroni <small>Vellard, 1932</small>
- Tityus chilensis <small>Lourenço, 2005</small>
- Tityus cisandinus <small>Lourenço & Ythier, 2017</small>
- Tityus clathratus <small>C. L. Koch, 1844</small>
- Tityus columbianus <small>(Thorell, 1876)</small>
- Tityus confluens <small>Borelli, 1899</small>
- Tityus costatus <small>(Karsch, 1879)</small>
- Tityus crassicauda <small>(Lourenço, 2013)</small>
- Tityus crassimanus <small>(Thorell, 1876)</small>
- Tityus cuellari <small>Lourenço, 1994</small>
- Tityus culebrensis <small>González-Sponga, 1994</small>
- Tityus cylindricus <small>(Karsch, 1879)</small>
- Tityus dasyurus <small>Pocock, 1897</small>
- Tityus dedoslargos <small>Francke & Stockwell, 1987</small>
- Tityus demangei <small>Lourenço, 1981</small>
- Tityus dinizi <small>Lourenço, 1997</small>
- Tityus discrepans <small>(Karsch, 1879)</small>
- Tityus dorae <small>González-Sponga, 2001</small>
- Tityus duacaensis <small>González-Sponga, 2007</small>
- Tityus dulceae <small>González-Sponga, 2006</small>
- Tityus dupouyi <small>González-Sponga, 1987</small>
- Tityus ebanoverde <small>Armas, 1999</small>
- Tityus ecuadorensis <small>Kraepelin, 1896</small>
- Tityus elii <small>(Armas & Marcano Fondeur, 1992)</small>
- Tityus elizabethae <small>Lourenço & Ramos, 2004</small>
- Tityus elizabethebravoi <small>González-Sponga & Wall Gonzalez, 2007</small>
- Tityus engelkei <small>Pocock, 1902</small>
- Tityus estherae <small>Santiago-Blay, 2009</small>
- Tityus evandroi <small>Mello-Leitão, 1945</small>
- â Tityus exstinctus <small>Lourenço, 1995</small>
- Tityus fasciolatus <small>Pessôa, 1935</small>
- Tityus festae <small>Borelli, 1899</small>
- Tityus filodendron <small>González-Sponga, 1981</small>
- Tityus florezi <small>Lourenço, 2000</small>
- Tityus footei <small>Chamberlin, 1916</small>
- Tityus forcipula <small>(Gervais, 1843)</small>
- Tityus fuhrmanni <small>Kraepelin, 1914</small>
- Tityus funestus <small>Hirst, 1911</small>
- Tityus gaffini <small>Lourenço, 2000</small>
- Tityus gasci <small>Lourenço, 1982</small>
- Tityus gonzalespongai <small>Quiroga, de Sousa, Parrilla-Alvarez & Manzanilla, 2004</small>
- Tityus guaricoensis <small>Gonzalez-Sponga, 2004</small>
- Tityus horacioi <small>Lourenço & Leguin, 2011</small>
- Tityus imei <small>Borges, de Sousa & Manzanilla, 2006</small>
- Tityus indecisus <small>Mello-Leitão, 1934</small>
- Tityus insignis <small>(Pocock, 1889)</small>
- Tityus intermedius <small>Borelli, 1899</small>
- Tityus irapaensis <small>González-Sponga, 2002</small>
- Tityus isabelceciliae <small>González-Sponga, D'Suze & Sevcik, 2001</small>
- Tityus ivani <small>González-Sponga, 2008</small>
- Tityus ivicnancor <small>González-Sponga, 1997</small>
- Tityus jeanvellardi <small>Lourenço, 2001</small>
- Tityus julianae <small>Lourenço, 2005</small>
- Tityus juliorum <small>Santiago-Blay, 2009</small>
- Tityus jussarae <small>Lourenço, 1988</small>
- Tityus kaderkai <small>Kovarik, 2005</small>
- Tityus kalettai <small>González-Sponga, 2007</small>
- Tityus kukututee <small>Ythier, Chevalier & Gangadin, 2020</small>
- Tityus kuryi <small>Lourenço, 1997</small>
- Tityus lancinii <small>González-Sponga, 1972</small>
- Tityus lokiae <small>Lourenço, 2005</small>
- Tityus longidigitus <small>González-Sponga, 2008</small>
- Tityus lourencoi <small>Flórez, 1996</small>
- Tityus lutzi <small>Giltay, 1928</small>
- Tityus macrochirus <small>Pocock, 1897</small>
- Tityus magnimanus <small>Pocock, 1897</small>
- Tityus maimirensis <small>González-Sponga, 2007</small>
- Tityus manakai <small>González-Sponga, 2004</small>
- Tityus maniapurensis <small>González-Sponga, 2009</small>
- Tityus marajoensis <small>Lourenço & da Silva, 2007</small>
- Tityus maranhensis <small>Lourenço, de Jesus Junior & Limeira-de-Oliveira, 2006</small>
- Tityus martinpaechi <small>Lourenço, 2001</small>
- Tityus matthieseni <small>Lourenço & Pinto-da-Rocha, 2000</small>
- Tityus mattogrossensis <small>Borelli, 1901</small>
- Tityus maturinensis <small>González-Sponga, 2008</small>
- Tityus melanostictus <small>Pocock, 1893</small>
- Tityus melici <small>Lourenço, 2003</small>
- Tityus metuendus <small>Pocock, 1897</small>
- Tityus michelii <small>Armas, 1982</small>
- Tityus microcystis <small>Lutz & Mello, 1922</small>
- Tityus monaguensis <small>González-Sponga, 1974</small>
- Tityus mongei <small>Lourenço, 1996</small>
- Tityus mucusunamensis <small>González-Sponga, 2006</small>
- Tityus munozi <small>Lourenço, 1997</small>
- Tityus neblina <small>Lourenço, 2008</small>
- Tityus neglectus <small>Mello-Leitão, 1932</small>
- Tityus neibae <small>Armas, 1999</small>
- Tityus nelsoni <small>Lourenço, 2005</small>
- Tityus nematochirus <small>Mello-Leitão, 1940</small>
- Tityus neoespartanus <small>González-Sponga, 1996</small>
- Tityus nororientalis <small>González-Sponga, 1996</small>
- Tityus obispoi <small>González-Sponga, 2006</small>
- Tityus obscurus <small>(Gervais, 1843)</small>
- Tityus obtusus <small>(Karsch, 1879)</small>
- Tityus ocelote <small>Francke & Stockwell, 1987</small>
- Tityus osmanus <small>González-Sponga, 1996</small>
- Tityus oteroi <small>Lourenço, 1998</small>
- Tityus ottenwalderi <small>Armas, 1999</small>
- Tityus pachyurus <small>Pocock, 1897</small>
- Tityus pampanensis <small>González-Sponga, 2007</small>
- Tityus paraguayensis <small>Kraepelin, 1895</small>
- Tityus parvulus <small>Kraepelin, 1914</small>
- Tityus paulistorum <small>Lourenço & Qi, 2006</small>
- Tityus perijanensis <small>González-Sponga, 1994</small>
- Tityus pictus <small>Pocock, 1893</small>
- Tityus pintodarochai <small>Lourenço, 2005</small>
- Tityus pittieri <small>González-Sponga, 1981</small>
- Tityus pococki <small>Hirst, 1907</small>
- Tityus portoplatensis <small>Armas & Marcano Fondeur, 1992</small>
- Tityus potameis <small>Lourenço & Leão Giupponi, 2004</small>
- Tityus prancei <small>Lourenço, 2000</small>
- Tityus proseni <small>Abalos, 1954</small>
- Tityus pugilator <small>Pocock, 1898</small>
- Tityus pusillus <small>Pocock, 1893</small>
- Tityus quiriquirensis <small>González-Sponga, 2008</small>
- Tityus quirogae <small>De Sousa, Manzanilla & Parrilla-Alvarez, 2006</small>
- Tityus quisqueyanus <small>(Armas, 1982)</small>
- Tityus ramirezi <small>Esquivel de Verde, 1969</small> [nomen dubium]
- Tityus raquelae <small>Lourenço, 1988</small>
- Tityus rebieri <small>Lourenço, 1997</small>
- Tityus riocaurensis <small>González-Sponga, 1996</small>
- Tityus rionegrensis <small>Lourenço, 2006</small>
- Tityus riverai <small>Teruel & Sanchez, 2009</small>
- Tityus roigi <small>Maury & Lourenço, 1987</small>
- Tityus rojasi <small>González-Sponga, 1996</small>
- Tityus romeroi <small>González-Sponga, 2008</small>
- Tityus rondonorum <small>Rojas-Runjaic & Armas, 2007</small>
- Tityus rufofuscus <small>Pocock, 1897</small>
- Tityus rugosus <small>Schenkel, 1932</small>
- Tityus rusmelyae <small>González-Sponga, D'Suze & Sevcik, 2001</small>
- Tityus sabinae <small>Lourenço, 1994</small>
- Tityus sanarensis <small>González-Sponga, 1997</small>
- Tityus sanfernandoi <small>González-Sponga, 2008</small>
- Tityus sarisarinamensis <small>González-Sponga, 2002</small>
- Tityus sastrei <small>Lourenço & Flórez, 1990</small>
- Tityus septentrionalis <small>Armas & Abud Antun, 2004</small>
- Tityus serrulatus <small>Lutz & Mello, 1922</small>
- Tityus shiriana <small>González-Sponga, 1991</small>
- Tityus silvestris <small>Pocock, 1897</small>
- Tityus simonsi <small>Pocock, 1900</small>
- Tityus smithii <small>Pocock, 1893</small>
- Tityus soratensis <small>Kraepelin, 1912</small>
- Tityus stigmurus <small>(Thorell, 1876)</small>
- Tityus strandi <small>Werner, 1939</small>
- Tityus surimeridensis <small>González-Sponga, 2002</small>
- Tityus surorientalis <small>González-Sponga, 1996</small>
- Tityus sylviae <small>Lourenço, 2005</small>
- Tityus tamayoi <small>González-Sponga, 1987</small>
- Tityus tayrona <small>Lourenço, 1991</small>
- Tityus tenuicauda <small>Prendini, 2001</small>
- Tityus timendus <small>Pocock, 1898</small>
- Tityus tocantim <small>Lourenço, 2022</small>
- Tityus trinitatis <small>Pocock, 1897</small>
- Tityus trivittatus <small>Kraepelin, 1898</small>
- Tityus tucurui <small>Lourenço, 1988</small>
- Tityus uniformis <small>Mello-Leitão, 1931</small>
- Tityus unus <small>Lourenço & Pinto-da-Rocha, 2000</small>
- Tityus uquirensis <small>González-Sponga, 2001</small>
- Tityus urachichensis <small>González-Sponga, 2007</small>
- Tityus urbinai <small>Scorza, 1952</small>
- Tityus uruguayensis <small>Borelli, 1901</small>
- Tityus vaissadei <small>Lourenço, 2002</small>
- Tityus valerae <small>Scorza, 1954</small>
- Tityus venamensis <small>González-Sponga, 1981</small>
- Tityus ventuarensis <small>González-Sponga, 2009</small>
- Tityus walli <small>González-Sponga & Wall Gonzalez, 2007</small>
- Tityus wayuu <small>Rojas-Runjaic & Armas, 2007</small>
- Tityus yerenai <small>González-Sponga, 2009</small>
- Tityus ythieri <small>Lourenço, 2007</small>
- Tityus zulianus <small>González-Sponga, 1981</small>
[?] Tityus thelyacanthus <small>Mello-Leitão, 1933</small>
References
External links