Thrixspermum, commonly known as hairseeds or ç½ç¹åÂ
°å± (bai dian lan shu), is a genus of flowering plants in the family Orchidaceae. Orchids in this genus are epiphytes, lithophytes or terrestrial plants with flat, leathery leaves and short-lived flowers with the sepals and petals more or less similar to each other. The labellum is rigidly fixed to the column and has three lobes. The side lobes are erect and the middle lobe is thick and fleshy. There are about 190 species distributed from tropical and subtropical Asia to the Western Pacific. Most species grow in lowland or tropical rainforests up to an altitude of 1,200 m.
Description
Orchids in the genus Thrixspermum are epiphytic or lithophytic, rarely terrestrial, monopodial herbs with long thick roots, and flat, fleshy leaves arranged in two ranks with their bases sheathing the stem. The flowers are arranged on a pendulous or arching flowering stem arising from a leaf . The flowers are usually short-lived and often open for less than a day. The sepals are free from and more or less similar to each other. The petals are free from each other and similar to, but slightly shorter than the sepals. The labellum is stiffly attached to the column with a pouched base and three lobes, the side lobes erect, usually short and blunt and the middle lobe thick and fleshy. The fruit is a long, thin capsule.
Taxonomy and naming
The genus Thrixspermum was first formally described in 1790 by João de Loureiro in Flora Cochinchinensis. The name Thrixspermum is derived from the Ancient Greek words (), meaning "hair" and () meaning "seed".
Species
The following is a list of species of Thrixspermum recognised by the Plants of the World Online as at January 2025:
- Thrixspermum aberrans <small>Schltr.</small>
- Thrixspermum acuminatissimum <small>(Rchb. f.) Rchb.f.</small>
- Thrixspermum acuminatissimum subsp. acuminatissimum
- Thrixspermum acuminatissimum subsp. affine <small>(Schltr.) P.O'Byrne & Gokusing</small>
- Thrixspermum acutilobum <small>J.J.Sm.</small>
- Thrixspermum agamense <small>J.J.Sm.</small>
- Thrixspermum agusanense <small>Ames</small>
- Thrixspermum alabense <small>J.J.Wood</small>
- Thrixspermum alboluteum <small>Toolmal, Schuit. & Culham</small>
- Thrixspermum amesianum <small>L.O.Williams</small>
- Thrixspermum amplexicaule <small>(Blume) Rchb.f.</small>
- Thrixspermum anceps <small>(Blume) Rchb.f.</small>
- Thrixspermum ancoriferum <small>(Guillaumin) Garay</small>
- Thrixspermum angustatum <small>L.O.Williams</small>
- Thrixspermum angustifolium <small>(Blume) Rchb.f.</small>
- Thrixspermum anjunganense <small>P.O'Byrne</small>
- Thrixspermum annamense <small>(Guillaum.) Garay</small>
- Thrixspermum arachnitiforme <small>Schltr.</small>
- Thrixspermum aurantiacum <small>J.J.Sm.</small>
- Thrixspermum bellamabantae <small>Cabactulan, Cootes & R.B.Pimentel</small>
- Thrixspermum benedictii <small>P.O'Byrne</small>
- Thrixspermum bicolor <small>P.O'Byrne, P.T.Ong & J.J.Verm.</small>
- Thrixspermum borneense <small>(Rolfe) Ridl.</small>
- Thrixspermum bosuangii <small>P.O'Byrne & Gokusing</small>
- Thrixspermum brevibracteatum <small>J.J.Sm.</small>
- Thrixspermum brevicapsularis <small>Holttum</small>
- Thrixspermum brevicaule <small>Carr</small>
- Thrixspermum brevipes <small>Schltr.</small>
- Thrixspermum breviscapum <small>(Carr) Kocyan & Schuit.</small>
- Thrixspermum bromeliforme <small>W.Suarez</small>
- Thrixspermum brunnescens <small>(Ridl.) J.J.Sm.</small>
- Thrixspermum calceolus <small>(Lindl.) Rchb.f.</small>
- Thrixspermum canaliculatum <small>J.J.Sm.</small>
- Thrixspermum carinatifolium <small>(Ridl.) Schltr.</small>
- Thrixspermum carnosum <small>(K.Schum.) Schltr.</small>
- Thrixspermum caudatum <small>P.O'Byrne & Gokusing</small>
- Thrixspermum celebicum <small>Schltr.</small>
- Thrixspermum centipeda <small>Lour</small>
- Thrixspermum cerinum <small>(Ridl.) J.J.Sm.</small>
- Thrixspermum changlangensis <small> K.Gogoi</small>
- Thrixspermum chanianum <small>P.O'Byrne & Gokusing</small>
- Thrixspermum clavatum <small>(J.Koenig) Garay</small>
- Thrixspermum clavilobum <small>J.J.Sm.</small>
- Thrixspermum complanatum <small>(J.Koenig) Schltr.</small>
- Thrixspermum congestum <small>(Bailey) Dockrill</small>
- Thrixspermum conigerum <small>J.J.Sm.</small>
- Thrixspermum cootesii <small>W.Suarez</small>
- Thrixspermum cordulatum <small>J.J.Sm.</small>
- Thrixspermum corneri <small>Holttum</small>
- Thrixspermum crassifolium <small>Ridl.</small>
- Thrixspermum crassilabre <small>(King & Pantl.) Ormerod</small>
- Thrixspermum crescentiforme <small>Ames & C.Schweinf.</small>
- Thrixspermum cryptophallon <small>P.O'Byrne & J.J.Verm.</small>
- Thrixspermum cymboglossum <small>Schltr.</small>
- Thrixspermum denticulatum <small>Schltr.</small>
- Thrixspermum doctersii <small>J.J.Sm.</small>
- Thrixspermum duplocallosum <small>Holttum</small>
- Thrixspermum elmeri <small>L.O.Williams</small>
- Thrixspermum elongatum <small>Ames</small>
- Thrixspermum erythrolomum <small>P.O'Byrne & J.J.Verm.</small>
- Thrixspermum eximium <small>L.O.Williams</small>
- Thrixspermum fantasticum <small>L.O.Williams</small>
- Thrixspermum fernandeziae <small>Cootes, Cabactulan & M.Leon</small>
- Thrixspermum filifolium <small>Schltr.</small>
- Thrixspermum filiforme <small>(Hook.f.) Kuntze</small>
- Thrixspermum fimbriatum <small>(Ridl.) J.J.Wood</small>
- Thrixspermum flaccidum <small>J.J.Sm.</small>
- Thrixspermum flammeum <small>Toolmal, Schuit. & Culham</small>
- Thrixspermum fleuryi <small>(Gagnep.) Tang & F.T.Wang</small>
- Thrixspermum formosanum <small>(Hayata) Schltr.</small>
- Thrixspermum fragrans <small>Ridl.</small>
- Thrixspermum fulgens <small>(Ridl.) Schltr.</small>
- Thrixspermum fuscum <small>(Ridl.) Merr.</small>
- Thrixspermum gombakense <small>J.J.Sm.</small>
- Thrixspermum gracilicaule <small>Schltr.</small>
- Thrixspermum graeffei <small>Rchb.f.</small>
- Thrixspermum grandiflorum <small>P.O'Byrne & J.J.Verm.</small>
- Thrixspermum hiepii <small>Aver. & Averyanova</small>
- Thrixspermum hystrix <small>(Blume) Rchb.f.</small>
- Thrixspermum incurvicalcar <small>J.J.Sm.</small>
- Thrixspermum indicum <small>Vik.Kumar, D.Verma & A.N.Rao</small>
- Thrixspermum indragiriense <small>Schltr.</small>
- Thrixspermum inquinatum <small>J.J.Sm.</small>
- Thrixspermum integrum <small>L.O.Williams</small>
- Thrixspermum iodochilus <small>Ridl.</small>
- Thrixspermum japonicum <small>(Miq.) Rchb.f.</small>
- Thrixspermum javanicum <small>J.J.Sm.</small>
- Thrixspermum kipandicum <small>P.O'Byrne & Gokusing</small>
- Thrixspermum kjellbergii <small>J.J.Sm.</small>
- Thrixspermum kocyanii <small>J.J.Wood & A.L.Lamb</small>
- Thrixspermum lampongense <small>J.J.Sm.</small>
- Thrixspermum lanatum <small>P.O'Byrne & Gokusing</small>
- Thrixspermum latifolium <small>J.J.Sm.</small>
- Thrixspermum latisaccatum <small>J.J.Sm.</small>
- Thrixspermum lengguanianum <small>P.T.Ong & P.O'Byrne</small>
- Thrixspermum leucorachne <small>Ridl.</small>
- Thrixspermum ligulatum <small>L.O.Williams</small>
- Thrixspermum linearifolium <small>Ames</small>
- Thrixspermum lingiae <small>P.O'Byrne & Gokusing</small>
- Thrixspermum linusii <small>J.J.Wood & A.L.Lamb</small>
- Thrixspermum lombokense <small>J.J.Sm.</small>
- Thrixspermum longicauda <small>Ridl.</small>
- Thrixspermum longilobum <small>J.J.Sm.</small>
- Thrixspermum longipedicellatum <small>(Joongku Lee, T.B.Tran & R.K.Choudhary) Kocyan & Schuit.</small>
- Thrixspermum longipilosum <small>J.J.Sm.</small>
- Thrixspermum loogemanianum <small>Schltr.</small>
- Thrixspermum lucidum <small>Schltr.</small>
- Thrixspermum luciferum <small>P.O'Byrne & J.J.Verm.</small>
- Thrixspermum maculatum <small>Schltr.</small>
- Thrixspermum majus <small>(Carr) Kocyan & Schuit.</small>
- Thrixspermum malayanum <small>J.J.Sm.</small>
- Thrixspermum megacarpum <small>P.O'Byrne, A.L.Lamb & P.T.Ong</small>
- Thrixspermum merapohense <small>P.O'Byrne & P.T.Ong</small>
- Thrixspermum mergitiferum <small>P.O'Byrne & J.J.Verm.</small>
- Thrixspermum merguense <small>(Hook.f .) Kuntze</small>
- Thrixspermum milneri <small>P.O'Byrne & Gokusing</small>)
- Thrixspermum montanum <small>Ridl.</small>
- Thrixspermum multicolor <small>(Ridl.) Merr.</small>
- Thrixspermum neohibernicum <small>Schltr.</small>
- Thrixspermum nicolasiorum <small>Calaramo, Cootes & Naive</small>
- Thrixspermum obtusum <small>(Blume) Rchb.f.</small>)
- Thrixspermum obyrneanum <small>Toolmal, P.T.Ong & Schuit.</small>
- Thrixspermum ochraceum <small>P.O'Byrne, P.T.Ong & J.J.Verm.</small>
- Thrixspermum odoratum <small>X.Q.Song, Q.W.Meng & Y.B.Luo</small>
- Thrixspermum olidum <small>P.O'Byrne</small>
- Thrixspermum oreadum <small>Schltr.</small>
- Thrixspermum pachychilum <small>P.O'Byrne & Gokusing</small>
- Thrixspermum pardale <small>(Ridl.) Schltr.</small>
- Thrixspermum patens <small>J.J.Sm.</small>
- Thrixspermum patkeiense <small>K.Gogoi</small>
- Thrixspermum pauciflorum <small>(Hook.f.) Kuntze</small>
- Thrixspermum pellicerae <small>Cootes, Cabactulan, R.B.Pimentel & M.Leon</small>
- Thrixspermum pensile <small>Schltr.</small>
- Thrixspermum pinocchio <small>P.O'Byrne & J.J.Verm.</small>
- Thrixspermum platycaule <small>Holttum</small>
- Thrixspermum platystachys <small>(F.M. Bailey) Schltr.</small>
- Thrixspermum poilanei <small>(Gagnep.) Tang & F.T.Wang</small>
- Thrixspermum polystictum <small>Toolmal, Schuit., & Suddee</small>
- Thrixspermum ponapense <small>Tuyama</small>
- Thrixspermum praetermissum <small>Toolmal, Schuit., & Suddee</small>
- Thrixspermum psiloglottis <small>(Ridl.) Schltr.</small>
- Thrixspermum pugionifolium <small>(Hook.f.) Schltr.</small>
- Thrixspermum pulchellum <small>(Thwaites) Schltr.</small>
- Thrixspermum pulchrum <small>Carr</small>
- Thrixspermum pulverulentum <small>(Carr) Kocyan & Schuit.</small>
- Thrixspermum punctatum <small>(Ridl.) J.J.Sm.</small>
- Thrixspermum purpurascens <small>Rchb. f.</small>
- Thrixspermum pusillum <small>(Guillaumin) Garay</small>
- Thrixspermum pygmaeum <small>(King & Pantl.) Holttum</small>
- Thrixspermum quinquelobum <small>Ames</small>
- Thrixspermum raciborskii <small>J.J.Sm</small>
- Thrixspermum raciborskii subsp. brevipollinium <small>P.O'Byrne & Ent</small>
- Thrixspermum raciborskii subsp. raciborskii
- Thrixspermum recurvum <small>(Hook.f.) Kuntze</small>
- Thrixspermum remotiflorum <small>J.J.Sm.</small>
- Thrixspermum rimauense <small>J.J.Wood & A.L.Lamb</small>
- Thrixspermum rimiae <small>P.O'Byrne & Ent</small>
- Thrixspermum robinsonii <small>Ames</small>
- Thrixspermum roseum <small>J.J.Sm.</small>
- Thrixspermum rostratum <small>Ames</small>
- Thrixspermum rubrocalcaratum <small>P.O'Byrne & Gokusing</small>
- Thrixspermum saccatum <small>P.O'Byrne & J.J.Verm.</small>
- Thrixspermum sagoense <small>J.J.Sm.</small>
- Thrixspermum samarindae <small>Schltr.</small>
- Thrixspermum sarawakense <small>Ames</small>
- Thrixspermum sarcophyllum <small>Garay</small>
- Thrixspermum saruwatarii <small>(Hayata) Schltr.</small>
- Thrixspermum scandens <small>P.O'Byrne</small>
- Thrixspermum scopa <small>(Rchb.f. ex Hook.f.) Holttum</small>
- Thrixspermum scortechinii <small>(Hook.f.) Kuntze</small>
- Thrixspermum semiteretifolium <small>J.J.Wood & A.L.Lamb</small>
- Thrixspermum simile <small>Tollmal, Schuit. & Utteridge</small>
- Thrixspermum simum <small>J.J.Sm.</small>
- Thrixspermum squarrosum <small>J.J.Sm.</small>
- Thrixspermum stelidioides <small>Aver. & Averyanova</small>
- Thrixspermum subteres <small>J.J.Sm.</small>
- Thrixspermum subulatum <small>(Blume) Rchb.f.</small>
- Thrixspermum sumatranum <small>J.J.Sm.</small>
- Thrixspermum sutepense <small>(Rolfe ex Downie) Tang & F.T.Wang</small>
- Thrixspermum taeniophyllum <small>Jun Y.Zhang, H.He & Yue H.Cheng</small>
- Thrixspermum tahanense <small>Carr</small>
- Thrixspermum taianum <small>P.O'Byrne & P.T.Ong</small>
- Thrixspermum tenuicalcar <small>Carr</small>
- Thrixspermum teretifolium <small>P.O'Byrne & J.J.Verm.</small>
- Thrixspermum torajaense <small>P.O'Byrne</small>
- Thrixspermum tortum <small>J.J.Sm.</small>
- Thrixspermum triangulare <small>Ames & C.Schweinf.</small>
- Thrixspermum trichoglottis <small>(Hook.f.)Kuntze</small>
- Thrixspermum tsii <small>W.H.Chen & Y.M.Shui</small>
- Thrixspermum tubulatum <small>P.O'Byrne</small>
- Thrixspermum tylophorum <small>Schltr.</small>
- Thrixspermum validum <small>J.J.Sm.</small>
- Thrixspermum vanoverberghii <small>Ames</small>
- Thrixspermum ventricosum <small>J.J.Wood & A.L.Lamb</small>
- Thrixspermum walkeri <small>Seidenf. & P.Ormerod</small>
- Thrixspermum warianum <small>Schltr.</small>
- Thrixspermum weberi <small>Ames</small>
- Thrixspermum wenzelii <small>Ames</small>
- Thrixspermum westenenkii <small>(J.J.Sm.) Kocyan & Schuit.</small>
- Thrixspermum williamsianum <small>(Kores) Ormerod</small>
- Thrixspermum xantholeucum <small>Schltr.</small>
- Thrixspermum xantholomum <small>P.O'Byrne & J.J.Verm.</small>
Distribution
Orchids in the genus Thrixspermum are found in China, Japan (including the Ryukyu Islands), Korea, Taiwan, India (including the Andaman and Nicobar Islands), Bangladesh, Nepal, Sri Lanka, Cambodia, Laos, Myanmar, Thailand, Vietnam, Borneo, Indonesia, Malaysia, the Philippines, New Guinea, the Solomon Islands, northern Australia (including Christmas Island), Fiji, New Caledonia, Samoa, Vanuatu and the Caroline Islands. About fourteen species, two of which are endemic occur in China, nine are found in Taiwan and three in Australia.
References