Taxonomy of Lucanidae

The Lucanidae are a family of beetles that include the stag beetles. The classification presented here follows Smith (2006), with the exception of the tribal classification within the Lucaninae.

Subfamily Aesalinae

Subfamily Aesalinae

Tribe Aesalini

:Genus Aesalus<small> (Fabricius, 1801)</small>

::*Aesalus asiaticus<small> Lewis, 1883</small>

::*Aesalus himalayicus<small> Kurosawa, 1985</small>

::*Aesalus imanishii<small> Inahara & Ratti, 1991</small>

::*Aesalus meridionalis<small> Bartolozzi, 1989</small>

::*Aesalus neotropicalis<small> Bates, 1886</small>

::*Aesalus saburoi

::*Aesalus sawaii<small> Fujita & Ichikawa, 1985</small>

::*Aesalus scarabaeoides<small> Panzer, 1793</small>

::*Aesalus sichuanensis

::*Aesalus smithi<small> Bates, 1889</small>

::*Aesalus trogoides<small> Albers, 1883</small>

::*Aesalus ulanowskii<small> Ganglbauer, 1886</small>

:Genus Cretaesalus (fossil)

:Genus Echinoaesalus<small> Zelenka, 1993</small>

::*Echinoaesalus barriesi<small> (Zelenka, 1993)</small>

::*Echinoaesalus hidakai<small> (Araya, 1993)</small>

::*Echinoaesalus jaechi<small> (Zelenka,1993)</small>

::*Echinoaesalus matsuii<small> (Araya, 1993)</small>

::*Echinoaesalus sabahensis<small> (Zelenka, 1994)</small>

::*Echinoaesalus schuhi<small> (Zelenka, 1994)</small>

::*Echinoaesalus timidus<small> (Krikken, 1975)</small>

::*Echinoaesalus yongi<small> (Araya, 1993)</small>

:Genus Lucanobium<small> Howden & Lawrence, 1974</small>

::*Lucanobium squamosum<small> Howden & Lawrence, 1974</small>

Tribe Ceratognathini

:Genus Ceratognathus<small> Westwood, 1838</small>

::*Ceratognathus abdominalis<small> Parry, 1870</small>

::*Ceratognathus bitumulatus<small> Carter, 1925</small>

::*Ceratognathus flabellatus<small> Boileau, 1905</small>

::*Ceratognathus frenchi<small> Blackburn, 1897</small>

::*Ceratognathus froggatti<small> Blackburn, 1894</small>

::*Ceratognathus gilesi<small> Blackburn, 1895</small>

::*Ceratognathus macrognathus<small> Boileau, 1905</small>

::*Ceratognathus mentiferus<small> Westwood, 1863</small>

::*Ceratognathus minutus<small> Lea, 1929</small>

::*Ceratognathus niger<small> Westwood, 1838</small>

::*Ceratognathus ocularis<small> Carter, 1925</small>

::*Ceratognathus rufipennis<small> Westwood, 1872</small>

::*Ceratognathus tasmanus<small> Benesh, 1943</small>

::*Ceratognathus westwoodi<small> Thomson, 1862</small>

:Genus Hilophyllus<small> Paulsen & Mondaca, 2006</small>

::*Hilophyllus argentinensis<small> (Martinez, 1981)</small>

::*Hilophyllus martinezi<small> Paulsen & Mondaca, 2006</small>

::*Hilophyllus penai<small> (Martinez, 1976)</small>

:Genus Holloceratognathus<small> Nikolajev, 1998</small>

::*Holloceratognathus cylindricus<small> (Broun, 1895)</small>

::*Holloceratognathus helotoides<small> (Thomson, 1862)</small>

::*Holloceratognathus passaliformis<small> (Holloway, 1962)</small>

:Genus Mitophyllus<small> Parry, 1843</small>

::*Mitophyllus alboguttatus<small> (Bates, 1867)</small>

::*Mitophyllus dispar<small> (Sharp, 1882)</small>

::*Mitophyllus foveolatus<small> (Broun, 1880)</small>

::*Mitophyllus gibbosus<small> (Broun, 1884)</small>

::*Mitophyllus insignis<small> Broun, 1921</small>

::*Mitophyllus irroratus<small> Parry, 1842</small>

::*Mitophyllus macrocerus<small> (Broun, 1886)</small>

::*Mitophyllus parrianus<small> Westwood, 1863</small>

::*Mitophyllus reflexus<small> Broun, 1908</small>

Tribe Nicagini

:Genus Nicagus<small> LeConte, 1861</small>

::*Nicagus obscurus<small> (LeConte, 1848)</small>

::*Nicagus occultus<small> Paulsen & Smith, 2005</small>

::*Nicagus japonicus<small> Nagel, 1928</small>

Subfamily Lampriminae

Subfamily Lampriminae

:Genus Dendroblax<small> White, 1846</small>

::*Dendroblax earlianus<small> White, 1846</small>

:Genus Homolamprima

::*Homolamprima crenulata<small> (MacLeay, 1885)</small>

:Genus Lamprima

::*Lamprima adolphinae<small> (Gestro, 1875)</small>

::*Lamprima aenea<small> Fabricius, 1792</small>

::*Lamprima aurata<small> Latreille, 1817</small>

::*Lamprima imberbis<small> Barber, 1926</small>

::*Lamprima insularis<small> Macleay, 1885</small>

:Genus Phalacrognathus

::*Phalacrognathus muelleri<small> MacLeay, 1885</small>

:Genus Streptocerus<small> Dejean, 1833</small>

::*Streptocerus speciosus<small> Fairmaire, 1850</small>

Subfamily Lucaninae

Subfamily Lucaninae <small>Latreille, 1804</small>

Tribe Aegini

:Genus Aegus<small> MacLeay, 1819</small>

Tribe Allotopini

:Genus Allotopus

::*Allotopus moellenkampi<small> (Fruhstorfer, 1894)</small>

::*Allotopus rosenbergi<small> (Vollenhoven, 1872)</small>

:Genus Mesotopus

Tribe Chiasognathini

:Genus Casignetus

:Genus Chiasognathus<small> Stephens, 1831</small>

::*Chiasognathus beneshi<small> Lacroix, 1978</small>

::*Chiasognathus grantii<small> Stephens, 1831</small>

::*Chiasognathus jousselini<small> Reiche, 1850</small>

::*Chiasognathus latreillei<small> Solier, 1851</small>

::*Chiasognathus mniszechii<small> Kiesche 1919</small>

:Genus Protognathinus

:Genus Sphaenognathus

Tribe Cladognathini

:Genus Aphanognathus

::*Aphanognathus laterotarsoides<small> (Houlbert, 1915)</small>

:Genus Capreolucanus

::*Capreolucanus sicardi<small> Didier, 1928</small>

:Genus Cladophyllus

:Genus Eligmodontus

:Genus Gonometopus

:Genus Macrodorcas<small> Motschulsky, 1862</small>

::*Macrodorcas adachii

::*Macrodorcas amamiana

::*Macrodorcas barbata

::*Macrodorcas bisignata

::*Macrodorcas bomansi

::*Macrodorcas davidi

::*Macrodorcas dierli

::*Macrodorcas elsiledis

::*Macrodorcas fulvonotata

::*Macrodorcas humilis

::*Macrodorcas intricata

::*Macrodorcas kobayashii

::*Macrodorcas koyamai

::*Macrodorcas melliana

::*Macrodorcas mochizukii

::*Macrodorcas moellenkampi

::*Macrodorcas montivaga

::*Macrodorcas pieli

::*Macrodorcas pseudaxis

::*Macrodorcas recta<small> Motschulsky, 1858</small>

::*Macrodorcas ruficrus

::*Macrodorcas songiana

::*Macrodorcas striatipennis<small> Motschulsky, 1862</small>

::*Macrodorcas ursulae

::*Macrodorcas vernicata

::*Macrodorcas yaeyamaensis

:Genus Palaeognathus

:Genus Prismognathus<small> Motschulsky, 1860</small>

::*Prismognathus alessandrae

::*Prismognathus angularis<small> Waterhouse, 1874</small>

::*Prismognathus branczicki

::*Prismognathus cheni

::*Prismognathus dauricus<small> Motschulsky, 1860</small>

::*Prismognathus davidis

::*Prismognathus delislei

::*Prismognathus formosanus

::*Prismognathus klapperichi

::*Prismognathus lucidus

::*Prismognathus morimotoi

::*Prismognathus nigerrimus

::*Prismognathus nigricolor

::*Prismognathus piluensis

::*Prismognathus platycephalus

::*Prismognathus ruficephalus

::*Prismognathus sinensis

::*Prismognathus subnitens

::*Prismognathus tokui

:Genus Prosopocoilus<small> Westwood, 1845</small>

:Genus Pseudorhaetus

:Genus Rhaetulus

:Genus Rhaetus

::*Rhaetus westwoodi<small> (Parry, 1862)</small>

:Genus Tetrarthrius

:Genus Weinreichius

::*Weinreichius perroti<small> Lacroix, 1978</small>

Tribe Colophonini

:Genus Colophon

Tribe Cyclommatini

:Genus Cyclommatus<small> Parry, 1863</small>

Tribe Dendeziini

:Genus Dendezia

:Genus Oonotus<small> Parry, 1864</small>

:Genus Xiphodontus

Tribe Dorcini

:Genus Bartolozziolucanus

:Genus Cantharolethrus

::*Cantharolethrus luxeri

:Genus Cyclommatus

::*Cyclommatus scutellaris

:Genus Dorcasoides

:Genus Dorcus<small> MacLeay, 1819</small>

::*Dorcus akahorii

::*Dorcus alcides

::*Dorcus alexisi

::*Dorcus amamianus

::*Dorcus antaeus

::*Dorcus anteus

::*Dorcus arfakianus

::*Dorcus binodulosus

::*Dorcus brevis

::*Dorcus bucephalus

::*Dorcus carinulatus

::*Dorcus consentaneus

::*Dorcus curvidens

::*Dorcus cylindricus

::*Dorcus davidis

::*Dorcus emikoae

::*Dorcus eurycephalus

::*Dorcus formosanus

::*Dorcus gracilicornis

::*Dorcus garndis

::*Dorcus hopei

::*Dorcus hyperion

::*Dorcus japonicus

::*Dorcus kyanrauensis

::*Dorcus metacostatus

::*Dorcus miwai

::*Dorcus musimon

::*Dorcus parallelus

::*Dorcus parallelipipedus<small> - lesser stag beetle. Linnaeus, 1758</small>

::*Dorcus parryi

::*Dorcus parvulus

::*Dorcus peyronis

::*Dorcus prochazkai

::*Dorcus ratiocinativus

::*Dorcus rectus

::*Dorcus reichei

::*Dorcus rudis

::*Dorcus sawaii

::*Dorcus schenklingi

::*Dorcus sewertzowi

::*Dorcus suturalis

::*Dorcus taiwanicus

::*Dorcus tanakai

::*Dorcus taurus

::*Dorcus tenuecostatus

::*Dorcus titanus

::*Dorcus tormosanus

::*Dorcus ursulus

::*Dorcus vavrai

::*Dorcus velutinus

::*Dorcus vicinus

::*Dorcus yamadai

:Genus Homoderus

:Genus Leptinopterus

:Genus Prosopocoilus

::*Prosopocoilus astacoides

::*Prosopocoilus biplagiatus

::*Prosopocoilus bison

::*Prosopocoilus giraffa

::*Prosopocoilus inclinatus

::*Prosopocoilus savagei

:Genus Rhaetulus

::*Rhaetulus crenatus

:Genus Serrognathus

:Genus Pycnosiphorus

:Genus Sclerostomus

Tribe Figulini

:Genus Cardanus

:Genus Dinonigidius

:Genus Figulus

:Genus Ganelius

:Genus Nigidionus

:Genus Nigidius

:Genus Novonigidius

:Genus Penichrolucanus

Tribe Lissapterini

:Genus Bomansius

:Genus Dorculus

:Genus Geodorcus

:Genus Hoplogonus

:Genus Lissapterus

:Genus Lissotes

:Genus Paralissotes

:Genus Pseudodorcus

Tribe Lucanini

:Genus Hexarthrius <small>Hope, 1842</small>

::*Hexarthrius aduncus<small>Jordan & Rothschild, 1894 </small>

::*Hexarthrius andreasi <small>Schenk, 2003</small>

::*Hexarthrius bowringii <small>Parry, 1862</small>

::*Hexarthrius buquettii <small>(Hope, 1843)</small>

::*Hexarthrius davisoni<small>Waterhouse, 1888</small>

::*Hexarthrius forsteri <small>(Hope, 1840)</small>

::*Hexarthrius howdeni <small>De Lisle, 1972</small>

::*Hexarthrius kirchneri <small>Schenk, 2003</small>

::*Hexarthrius mandibularis <small>Deyrolle, 1881</small>

::*Hexarthrius melchioritis <small>Séguy, 1954 </small>

::*Hexarthrius mniszechi<small>(Thomson, 1857)</small>

::*Hexarthrius nigritus <small>Lacroix, 1990</small>

::*Hexarthrius parryi <small>Hope, 1842</small>

::*Hexarthrius rhinoceros<small>(Olivier, 1789)</small>

::*Hexarthrius vitalisi <small>Didier, 1925</small>

:Genus Lucanus

::*Lucanus capreolus

::*Lucanus cervus

::*Lucanus elaphus

::*Lucanus formosanus

::*Lucanus maculifemoratus

::*Lucanus mazama

::*Lucanus placidus

::*Lucanus swinhoei

::*Lucanus tetraodon

:Genus Pseudolucanus

::*Pseudolucanus barbarossa<small> (Fabricius, 1801)</small>

Tribe Odontolabini

:Genus Calcodes

:Genus Neolucanus

::*Neolucanus castanopterus

:Genus Odontolabis

::*Odontolabis castelnaudi

::*Odontolabis cuvera

::*Odontolabis femoralis

::*Odontolabis siva

Tribe Platycerini

:Genus Platyceroides<small> Benesh, 1946</small>

:Genus Platyceropsis<small> Benesh, 1946 </small>

::*Platyceropsis keeni<small> (Casey, 1895)</small>

:Genus Platycerus<small> Geoffroy, 1762</small>

Tribe Ryssonotini

:Genus Cacostomus

::*Cacostomus floralis<small> (Lea, 1914)</small>

::*Cacostomus squamosus<small> Newman, 1840</small>

::*Cacostomus subvittata<small> (Moore, 1994)</small>

:Genus Ryssonotus

Subfamily Syndesinae

Subfamily Syndesinae

Tribe Ceruchini

:Genus Ceruchus<small> MacLeay, 1819</small>

::*Ceruchus piceus<small> (Weber, 1801)</small>

::*Ceruchus punctatus<small> LeConte, 1869</small>

::*Ceruchus striatus<small> LeConte, 1859</small>

::*Ceruchus yangi<small> Huang, Imura & Chen, 2011</small>

Tribe Sinodendronini

:Genus Sinodendron<small> Hellwig in Schneider, 1792</small>

::*Sinodendron cylindricum<small> (Linnaeus, 1758)</small>

::*Sinodendron rugosum<small> Mannerheim, 1843</small>

Tribe Syndesini

:Genus Psilodon<small> Perty, 1830</small>

::*Psilodon aequinoctiale<small> (Buquet, 1840)</small>

::*Psilodon gilberti<small> Boucher, 1993</small>

::*Psilodon schuberti<small> Perty, 1830</small>

::*Psilodon seguyi<small> (Didier, 1929) </small>

::*Psilodon xerophilicum<small> Martinez & Reyes-Castillo, 1985 </small>

:Genus Syndesus<small> MacLeay, 1819</small>

::*â€ÂSyndesus ambericus <small>Woodruff, 2009</small> (Miocene?; Dominican amber)

::*Syndesus cancellatus<small> Montrouzier, 1860 </small>

::*Syndesus cornutus<small> (Fabricius, 1801) </small>

::*Syndesus macleayi<small> Boileau, 1905</small>

References