Scolopendra (from Ancient Greek (skólops), meaning "thorn", and (énteron), meaning "earthworm") is a species-rich genus of large tropical centipedes of the family Scolopendridae.
Description
The genus Scolopendra contains many species of centipedes found across the world's tropics and warmer temperate areas. The species vary considerably in coloration and size. Scolopendra are mostly very large centipedes. The largest species found in tropical climates can exceed and are the largest living centipedes in the world. All Scolopendra species can deliver a painful bite, injecting venom through their forcipules, which are not fangs or other mouthparts; rather, these are modified legs on the first body segment.
Ecology
Scolopendra species are active predators, feeding primarily on insects and other invertebrates. Larger specimens have been observed preying on frogs, tarantulas, lizards, birds, snakes, rodents, and even bats. Two southeast Asian species, S. cataracta and S. paradoxa, as well as S. alcyona from the Ryukyu Islands, are amphibious, as these species can travel underwater by swimming or walking.
Venom
The venom of most species is not medically significant; however, bites from several species can cause intense and long-lasting pain and swelling. Large Scolopendra species from Asian/Pacific regions, such as Scolopendra subspinipes and Scolopendra dehaani, are particularly potent, and have caused one reported fatality. In 2014, a fatality was reported for a bite from a Scolopendra gigantea. The venom of certain Scolopendra species was found to contain compounds such as serotonin, haemolytic phospholipase, a cardiotoxic protein, and a cytolysin.
Taxonomic history
Scolopendra was one of the genera created by Carl Linnaeus in his 1758 10th edition of Systema Naturae, the starting point for zoological nomenclature. Only two of the species originally assigned to the genus remain so: Scolopendra gigantea and S. morsitans; the latter was chosen to be the type species by Opinion 454 of the International Commission on Zoological Nomenclature, overruling a previous designation by Pierre André Latreille, in which he chose Linnaeus' Scolopendra forficata (now Lithobius forficatus) as the type species.
Species
The genus Scolopendra contains these species:
- Scolopendra abnormis <small>Lewis & Daszak, 1996</small>
- Scolopendra afer <small>(Meinert, 1886)</small>
- Scolopendra alcyona <small>Tsukamoto & Shimano, 2021</small>
- Scolopendra algerina <small>Newport, 1845</small>
- Scolopendra alternans <small>Leach, 1813</small>
- Scolopendra andhrensis <small>Jangi & Dass, 1984</small>
- Scolopendra angulata <small>Newport, 1844</small>
- Scolopendra angusticollis <small>Murray, 1887</small>
- Scolopendra anomia <small>Newport, 1844</small>
- Scolopendra appendiculata <small>Daday, 1891</small>
- Scolopendra arenicola <small>(Lawrence, 1975)</small>
- Scolopendra arthrorhabdoides <small>Ribaut, 1912</small>
- Scolopendra attemsi <small>Lewis, Minelli & Shelley, 2006</small>
- Scolopendra aztecorum <small>Verhoeff, 1934</small>
- Scolopendra calcarata <small>Porat, 1876</small>
- Scolopendra canidens <small>Newport, 1844</small>
- Scolopendra cataracta <small>Siriwut, Edgecombe & Panha, 2016</small>
- Scolopendra childreni <small>Newport, 1844</small>
- Scolopendra chlora <small>Chamberlin, 1942</small>
- Scolopendra chlorotes <small>C. L. Koch, 1856</small>
- Scolopendra cingulata <small>Latreille, 1829</small>
- Scolopendra clavipes <small>C. L. Koch, 1847</small>
- Scolopendra concolor <small>Newport, 1845</small>
- Scolopendra crassa <small>Templeton, 1846</small>
- Scolopendra cretica <small>Attems, 1902</small>
- Scolopendra cribrifera <small>Gervais, 1847</small>
- Scolopendra crudelis <small>C. L. Koch, 1847</small>
- Scolopendra dalmatica <small>C. L. Koch, 1847</small>
- Scolopendra dawydoffi <small>Kronmüller, 2012</small>
- Scolopendra dehaani <small>Brandt, 1840</small>
- Scolopendra ellorensis <small>Jangi & Dass, 1984</small>
- Scolopendra fissispina <small>L. Koch, 1865</small>
- Scolopendra foveolata <small>Verhoeff, 1937</small>
- Scolopendra galapagoensis <small>Bollman, 1889</small>
- Scolopendra gigantea <small>Linnaeus, 1758</small>
- Scolopendra gracillima <small>Attems, 1898</small>
- Scolopendra hainanum <small>Kronmüller, 2012</small>
- Scolopendra hardwickei <small>Newport, 1844</small>
- Scolopendra hermosa <small>Chamberlin, 1941</small>
- Scolopendra heros <small>Girard, 1853</small>
- Scolopendra horrida <small>C. L. Koch, 1847</small>
- Scolopendra inaequidens <small>Gervais, 1847</small>
- Scolopendra indiae <small>(Chamberlin, 1914)</small>
- Scolopendra indica <small>Meinert, 1886</small>
- Scolopendra inermipes <small>C. L. Koch, 1847</small>
- Scolopendra inermis <small>Newport, 1845</small>
- Scolopendra jangii <small>Khanna & Yadav, 1997</small>
- Scolopendra japonica <small>Koch, 1878</small>
- Scolopendra koreana <small>(Verhoeff, 1934)</small>
- Scolopendra labiata <small>C. L. Koch, 1863</small>
- Scolopendra laeta <small>Haase, 1887</small>
- Scolopendra langi <small>(Chamberlin, 1927)</small>
- Scolopendra latro <small>Meinert, 1886</small>
- Scolopendra leki <small>(Waldock & Edgecombe 2012)</small>
- Scolopendra limicolor <small>Wood, 1861</small>
- Scolopendra lucasii <small>Gervais, 1847</small>
- Scolopendra lufengia <small>S. Kang et al., 2017</small>
- Scolopendra lutea <small>(Attems, 1928)</small>
- Scolopendra madagascariensis <small>Attems, 1910</small>
- Scolopendra malkini <small>Chamberlin, 1955</small>
- Scolopendra mazbii <small>Gravely, 1912</small>
- Scolopendra media <small>(Muralewicz, 1926)</small>
- Scolopendra melionii <small>Lucas, 1853</small>
- Scolopendra metuenda <small>Pocock, 1895</small>
- Scolopendra michoacana <small>Chamberlin, 1941</small>
- Scolopendra mima <small>Chamberlin, 1942</small>
- Scolopendra mirabilis <small>(Porat, 1876)</small>
- Scolopendra monticola <small>(Lawrence, 1975)</small>
- Scolopendra morsitans <small>Linnaeus, 1758</small>
- Scolopendra multidens <small>Newport, 1844</small>
- Scolopendra negrocapitis <small>Zhang & Wang, 1999</small>
- Scolopendra nuda <small>(Jangi & Dass, 1980)</small>
- Scolopendra occidentalis <small>F. Meinert, 1886</small>
- Scolopendra octodentata <small>Verhoeff, 1934</small>
- Scolopendra oraniensis <small>Lucas, 1846</small>
- Scolopendra paradoxa <small>Doménech, 2018</small>
- Scolopendra pachygnatha <small>Pocock, 1895</small>
- Scolopendra paranuda <small>(Khanna & Tripathi, 1987)</small>
- Scolopendra pentagramma <small>Motschoulsky, 1886</small>
- Scolopendra pinguis <small>Pocock, 1891</small>
- Scolopendra polymorpha <small>Wood, 1861</small>
- Scolopendra pomacea <small>C. L. Koch, 1847</small>
- Scolopendra puncticeps <small>Wood, 1861</small>
- Scolopendra punensis <small>Jangi & Dass, 1984</small>
- Scolopendra robusta <small>Kraepelin, 1903</small>
- Scolopendra sanatillae <small>Bollman, 1893</small>
- Scolopendra silhetensis <small>Newport, 1845</small>
- Scolopendra spinipriva <small>Bücherl, 1946</small>
- Scolopendra spinosissima <small>Kraepelin, 1903</small>
- Scolopendra subspinipes <small>Leach, 1816</small>
- Scolopendra sumichrasti <small>Saussure, 1860</small>
- Scolopendra tenuitarsis <small>Pocock, 1895</small>
- Scolopendra teretipes <small>(Porat,1893)</small>
- Scolopendra valida <small>Lucas, 1840</small>
- Scolopendra violacea <small>Fabricius, 1798</small>
- Scolopendra viridicornis <small>Newport, 1844</small>
- Scolopendra viridipes <small>Dufour, 1820</small>
- Scolopendra viridis <small>Say, 1821</small>
- Scolopendra zuluana <small>(Lawrence, 1958)</small>
One fossil species, â Scolopendra proavita, is known from Baltic amber deposits from the Eocene of Poland. Fossil remains of a species tentatively assigned to S. morsitans (as S. (cf) morsitans) are also known from Pliocene-aged rocks in the Makapansgat of South Africa.
Species gallery
See also
References