Rhipicephalus is a genus of hard ticks in the family Ixodidae, consisting of about 90 species. While many species are native to tropical Africa, the genus is cosmopolitan, being found across the world. Several species are vectors of significant human and animal pathogens.
Morphology
Rhipicephalus ticks are commonly called 'brown ticks' as they are mostly brown in colour. Most adult ticks in this genus do not have colour patterns on their scutum (inornate). Many taxa are very similar, without clear distinguishing features. Identification is often further frustrated by intraspecies variation. While males are reliably distinct, instar and female specimens are sometimes impossible to identify due to the lack of differential morphological features.
Disease potential
Rhipicephalus parasitism is of significant economic, medical, and veterinary importance. Many species are vectors of pathogens of humans and animals, including causing East Coast fever, anaplasmosis, babesiosis, rickettsiosis, Boutonneuse fever, Lyme disease, Q fever, Rocky Mountain spotted fever, NSDV and CrimeanâÂÂCongo hemorrhagic fever. In addition to the infectious agents spread by ticks, tick bites can lead to allergic and toxic reactions.
Some species can cause tick paralysis by injecting a neurotoxin with their bite, such as Rhipicephalus evertsi and Rhipicephalus warburtoni.
Sex pheromone studies
A number of Rhipicephalus spp. females produce phenol and p-cresol after being fed for six days. These compounds may act as sex pheromones since these compounds were shown to be the sex pheromone of Rhipicephalus appendiculatus.
Etymology
The name Rhipicephalus is derived from the Greek word rhiphis, meaning "fan-like", and úõÃÂñûî, kephalÃÂ, meaning "head". The two terms are related to the hexagonal basis capituli of Rhipicephalus.
Species
As of 2025, the genus contains 90 recognised species:
- Rhipicephalus afranicus <small>Bakkes, 2020</small>
- Rhipicephalus (Boophilus) annulatus <small>Say, 1821</small>
- Rhipicephalus appendiculatus <small>Neumann 1901</small> – brown ear tick
- Rhipicephalus aquatilis <small>Walker, Keirans & Pegram, 1993</small>
- Rhipicephalus armatus <small>Pocock, 1900</small>
- Rhipicephalus arnoldi <small>Theiler & Zumpt, 1949</small>
- Rhipicephalus aurantiacus <small>Neumann, 1907</small>
- Rhipicephalus australis <small>Fuller, 1899</small>
- Rhipicephalus bequaerti <small>Zumpt, 1949</small>
- Rhipicephalus bergeoni <small>Morel & Balis, 1976</small>
- Rhipicephalus boueti <small>Morel, 1957</small>
- Rhipicephalus bursa <small>Canestrini & Fanzago, 1878</small>
- Rhipicephalus camicasi <small>Morel, Mouchet & Rodhain, 1976</small>
- Rhipicephalus capensis <small>Koch, 1844</small>
- Rhipicephalus carnivoralis <small>Walker, 1966</small>
- Rhipicephalus cliffordi <small>Morel, 1965</small>
- Rhipicephalus complanatus <small>Neumann, 1911</small>
- Rhipicephalus compositus <small>Neumann, 1897</small>
- Rhipicephalus congolensis <small>Apanaskevich, Horak & Mulumba-Mfumu, 2013</small>
- Rhipicephalus cuspidatus <small>Neumann, 1906</small>
- Rhipicephalus (Boophilus) decoloratus <small>Koch, 1844</small>
- Rhipicephalus deltoideus <small>Neumann, 1910</small>
- Rhipicephalus distinctus <small>Bedford 1932</small>
- Rhipicephalus duttoni <small>Neumann, 1907</small>
- Rhipicephalus dux <small>Dönitz, 1910</small>
- Rhipicephalus evertsi <small>Neumann, 1897</small>
- Rhipicephalus exophthalmos <small>Keirans & Walker 1993</small>
- Rhipicephalus follis <small>Dönitz, 1910</small>
- Rhipicephalus fulvus <small>Neumann, 1913</small>
- Rhipicephalus (Boophilus) geigyi <small>Aeschlimann & Morel, 1965</small>
- Rhipicephalus gertrudae <small>Feldman-Muhsam, 1960</small>
- Rhipicephalus glabroscutatus <small>Du Toit, 1941</small>
- Rhipicephalus guilhoni <small>Morel & Vassiliades, 1963</small>
- Rhipicephalus haemaphysaloides <small>Supino, 1897</small>
- Rhipicephalus hibericus <small>Millán, Rodriguez-Pastor & Estrada-Peña, 2024</small>
- Rhipicephalus humeralis <small>Rondelli, 1926</small>
- Rhipicephalus hurti <small>Wilson, 1954</small>
- Rhipicephalus interventus <small>Walker, Pegram & Keirans, 1995</small>
- Rhipicephalus jeanneli <small>Neumann 1913</small>
- Rhipicephalus kochi <small>Dönitz, 1905</small>
- Rhipicephalus (Boophilus) kohlsi <small>Hoogstraal & Kaiser, 1960</small>
- Rhipicephalus leporis <small>Pomerantsev, 1946</small>
- Rhipicephalus linnaei <small>(Audouin, 1826)</small>
- Rhipicephalus longiceps <small>Warburton 1912</small>
- Rhipicephalus longicoxatus <small>Neumann, 1905</small>
- Rhipicephalus longus <small>Neumann, 1907</small>
- Rhipicephalus lounsburyi <small>Walker, 1990</small>
- Rhipicephalus lunulatus <small>Neumann 1907</small>
- Rhipicephalus maculatus <small>Neumann, 1901</small>
- Rhipicephalus masseyi <small>Nuttall & Warburton 1908</small>
- Rhipicephalus (Boophilus) microplus <small>Canestrini, 1888</small>
- Rhipicephalus moucheti <small>Morel, 1965</small>
- Rhipicephalus muehlensi <small>Zumpt, 1943</small>
- Rhipicephalus muhsamae <small>Morel & Vassiliades, 1965</small>
- Rhipicephalus neumanni <small>Walker, 1990</small>
- Rhipicephalus nitens <small>Neumann, 1904</small>
- Rhipicephalus oculatus <small>Neumann 1901</small>
- Rhipicephalus oreotragi <small>Walker & Horak, 2000</small>
- Rhipicephalus pilans <small>Schulze, 1935</small>
- Rhipicephalus planus <small>Neumann, 1907</small>
- Rhipicephalus praetextatus <small>Gerstäcker, 1873</small>
- Rhipicephalus pravus <small>Dönitz, 1910</small>
- Rhipicephalus pseudolongus <small>Santos Dias, 1953</small>
- Rhipicephalus pulchellus <small>Gerstäcker, 1873</small>
- Rhipicephalus pumilio <small>Schulze, 1935</small>
- Rhipicephalus punctatus <small>Warburton 1912</small>
- Rhipicephalus pusillus <small>Gil Collado, 1936</small>
- Rhipicephalus ramachandrai <small>Dhanda, 1966</small>
- Rhipicephalus rossicus <small>Yakimov & Kol-Yakimova, 1911</small>
- Rhipicephalus rutilus <small>Koch, 1844</small>
- Rhipicephalus sanguineus <small>Latreille, 1806</small> – brown dog tick
- Rhipicephalus scalpturatus <small>Santos Dias, 1959</small>
- Rhipicephalus schulzei <small>Olenev 1929</small>
- Rhipicephalus sculptus <small>Warburton 1912</small>
- Rhipicephalus secundus <small>Feldman-Muhsam</small><small>, 1952</small>
- Rhipicephalus senegalensis <small>Koch, 1844</small>
- Rhipicephalus serranoi <small>Santos Dias 1950</small>
- Rhipicephalus simpsoni <small>Nuttall 1910</small>
- Rhipicephalus simus <small>Koch, 1844</small>
- Rhipicephalus sulcatus <small>Neumann, 1908</small>
- Rhipicephalus supertritus <small>Neumann, 1907</small>
- Rhipicephalus tetracornus <small>Kitaoka & Suzuki, 1983</small>
- Rhipicephalus theileri <small>Bedford & Hewitt, 1925</small>
- Rhipicephalus tricuspis <small>Dönitz, 1906</small>
- Rhipicephalus turanicus <small>Pomerantsev, 1936</small>
- Rhipicephalus walkerae <small>Horak, Apanaskevich and Kariuki, 2013</small>
- Rhipicephalus warburtoni <small>Walker & Horak, 2000</small>
- Rhipicephalus zambeziensis <small>Walker, Norval & Corwin, 1981</small>
- Rhipicephalus ziemanni <small>Neumann, 1904</small>
- Rhipicephalus zumpti <small>Santos Dias, 1950</small>
Unrecognised
Rhipicephalus sanguineus species complex
Rhipicephalus includes a clade centred around the brown dog tick, Rhipicephalus sanguineus. The taxonomy is currently in flux as phylogenies are developed. The species complex contains 16 known species-level lineages, but these often do not map singularly or wholly onto known species. Some species that are prima facie in the complex are yet to be sequenced:
- R. afranicus <small>Bakkes, 2020</small> - 'Afrotropical lineage'
- R. camicasi <small>Morel, Mouchet & Rodhain, 1976</small>
- R. guilhoni <small>Morel & Vassiliades, 1963</small>
- R. leporis <small>Pomerantsev, 1946</small>
- R. linnaei <small>Audouin, 1826</small> - 'tropical lineage'
- R. moucheti <small>Morel, 1965</small>
- R. pumilio <small>Schulze, 1935</small>
- R. pusillus <small>Gil Collado, 1936</small>
- R. rossicus <small>Yakimov & Kol-Yakimova, 1911</small>
- R. rutilus <small>Koch, 1844</small> - 'southeastern European lineage'
- R. sanguineus sensu stricto <small>Latreille, 1806</small>
- R. schulzei <small>Olenev 1929</small>
- R. secundus <small>Feldman-Muhsam</small><small>, 1952</small>
- R. sulcatus <small>Neumann, 1908</small>
- R. turanicus <small>Pomerantsev, 1936</small>
Suspect
- Rhipicephalus bergeoni <small>Morel & Balis, 1976 </small>
- Rhipicephalus hibericus <small>Millán, Rodriguez-Pastor & Estrada-Peña, 2024</small>
Subgenus Boophilus
Boophilus was once considered a separate genus, but studies in the early 2000s resulted in Boophilus being made a subgenus of Rhipicephalus. Although Boophilus has been included in the Rhipicephalus genus, their morphology is different from the typical Rhipicephalus species.
Notes
References
Further reading
External links