Pyrgulopsis is a genus of freshwater snails with a gill and an operculum, aquatic gastropod mollusks in the family Hydrobiidae.
Etymology
The name Pyrgulopsis is composed from Pyrgula, another genus of snail, and opsis = of.
Description
Generic characters of the genus Pyrgulopsis are: the shell is minute, conically turreted, somewhat elongated, imperforate and unicarinate. The apex is acute. The aperture is ovate. The edge of the aperture, called the peritreme, is continuous. The operculum is ovate, thin, corneous and spiral, with polar point well forward and approximating the columella.
The jaw is thin and membranaceous. The radula is odontophore, with teeth are arranged in transverse rows, according to the formula 3 + 1 + 3. Formula for denticles of rhachidian: .
Distribution
The distribution of the genus Pyrgulopsis includes Western and South-western United States. Snails of species in the genus Pyrgulopsis occur in fresh water and in brackish water.
Species
Pyrgulopsis is the largest genus of freshwater gastropods in the North America. In 2014, 139 species were recognized in this genus.
Species in the genus Pyrgulopsis include:
- Pyrgulopsis aardahli <small>Hershler, 1989</small> - Benton Valley springsnail
- Pyrgulopsis aloba <small>Hershler, 1999</small> - Duckwater pyrg
- Pyrgulopsis amargosae <small>Hershler, 1989</small> - Amargosa springsnail
- Pyrgulopsis anguina <small>Hershler, 1989</small> - Longitudinal gland pyrg
- Pyrgulopsis archimedis <small>S. S. Berry, 1947</small> - archimedes pyrg
- Pyrgulopsis arizonae <small>(Taylor, 1987)</small> - Apache springsnail
- Pyrgulopsis avernalis <small>(Pilsbry, 1935)</small> - Moapa pebblesnail
- Pyrgulopsis bacchus <small>Hershler, 1988</small> - Grand Wash springsnail
- Pyrgulopsis bedfordensis <small>Hershler & Gustafson, 2001</small>
- Pyrgulopsis bernardina <small>(Taylor, 1987)</small> - San Bernardino springsnail
- Pyrgulopsis blainica <small>Hershler, Liu & Gustafson, 2008</small>
- Pyrgulopsis bruneauensis <small>Hershler, 1990</small> - Bruneau hot springsnail
- Pyrgulopsis bryantwalkeri <small>Hershler, 1994</small> - Cortez Hills pebblesnail
- Pyrgulopsis californiensis <small>(Gregg & Taylor, 1965)</small> - Laguna Mountain springsnail
- Pyrgulopsis carinifera <small>(Pilsbry, 1935)</small>
- Pyrgulopsis castaicensis <small>Hershler & Liu, 2010</small>
- Pyrgulopsis chamberlini <small>(Hershler, 1998)</small> - smooth Glenwood pyrg
- Pyrgulopsis chupaderae <small>(Taylor, 1987)</small> - Chupadera springsnail
- Pyrgulopsis coloradensis <small>Hershler, 1998</small> - Blue Point pyrg
- Pyrgulopsis conica <small>Hershler, 1988</small> - Kingman springsnail
- Pyrgulopsis cruciglans <small>Hershler, 1998</small> - Transverse grand pyrg
- Pyrgulopsis crystalis <small>Hershler & Sada, 1987</small> - Crystal springsnail
- Pyrgulopsis davisi <small>(Taylor, 1987)</small> - Limpia Creek springsnail
- Pyrgulopsis deaconi <small>Hershler, 1998</small> - Spring Mountains Pyrg
- Pyrgulopsis deserta <small>(Pilsbry, 1916)</small> - desert springsnail
- Pyrgulopsis diablensis <small>Hershler, 1995</small> - Diablo Range pyrg
- Pyrgulopsis eremica <small>Hershler, 1995</small> - Smoke Creek pyrg
- Pyrgulopsis erythropoma <small>(Pilsbry, 1899)</small> - Ash Meadows pebblesnail
- Pyrgulopsis fairbanksensis <small>Hershler & Sada, 1987</small> - Fairbanks springsnail
- Pyrgulopsis fusca <small>(Hershler, 1998)</small> - Otter Creek pyrg
- Pyrgulopsis gibba <small>Hershler, 1995</small> - Surprise Valley pyrg
- Pyrgulopsis gilae <small>(Taylor, 1987)</small> - Gila springsnail
- Pyrgulopsis giuliani <small>Hershler & Pratt, 1990</small> - southern Sierra Nevada springsnail
- Pyrgulopsis glandulosa <small>Hershler, 1988</small> - Verde Rim springsnail
- Pyrgulopsis greggi <small>Hershler, 1995</small> - Kern River pyrg
- Pyrgulopsis hamlinensis <small>Hershler, 1998</small> - Hamlin Valley pyrg
- Pyrgulopsis hendersoni <small>(Pilsbry, 1933)</small> - Harney Lake springsnail
- Pyrgulopsis idahoensis <small>(Pilsbry, 1933)</small> - Idaho springsnail
- Pyrgulopsis ignota <small>Hershler, Liu & Lang, 2010</small>
- Pyrgulopsis intermedia <small>(Tryon, 1865)</small> - Crooked Creek springsnail
- Pyrgulopsis isolata <small>Hershler & Sada, 1987</small> - elongate-gland springsnail
- Pyrgulopsis kolobensis <small>(Taylor, 1987)</small> - Toquerville springsnail
- Pyrgulopsis licina <small>Hershler, Liu & Bradford, 2013</small>
- Pyrgulopsis longae <small>Hershler, 1995</small> - Long Valley pyrg
- Pyrgulopsis longinqua <small>(Gould, 1855)</small>
- Pyrgulopsis marilynae <small>(Hershler, Ratcliffe, Liu, Lang and Hay, 2014)</small>
- Pyrgulopsis merriami <small>(Pilsbry & Beecher, 1892)</small> - Pahranagat pebblesnail
- Pyrgulopsis metcalfi <small>(Taylor, 1987)</small> - Naegele springsnail
- Pyrgulopsis micrococcus <small>(Pilsbry, 1893)</small> - Oasis Valley springsnail
- Pyrgulopsis milleri <small>Hershler & Liu, 2010</small>
- Pyrgulopsis minkleyi <small>(D. W. Taylor, 1966)</small>
- Pyrgulopsis montezumensis <small>Hershler, 1988</small> - Montezuma Well springsnail
- Pyrgulopsis morrisoni <small>Hershler, 1988</small> - Page springsnail
- Pyrgulopsis nanus <small>Hershler & Sada, 1987</small> - distal-gland springsnail
- Pyrgulopsis neomexicana <small>(Pilsbry, 1916)</small> - Socorro springsnail
- Pyrgulopsis nevadensis <small>(Stearns, 1883)</small> - corded pyrg - type species
- Pyrgulopsis nonaria <small>(Hershler, 1998)</small> - Ninemile pyrg
- Pyrgulopsis owensensis <small>Hershler, 1989</small> - Owens Valley springsnail
- Pyrgulopsis pecosensis <small>(Taylor, 1987)</small> - Pecos springsnail
- Pyrgulopsis perforata <small>Hershler, Liu & Bradford, 2013</small>
- Pyrgulopsis perturbata <small>Hershler, 1989</small> - Fish Slough springsnail
- Pyrgulopsis pilsbryana <small>(J. L. Baily & R. I. Baily, 1952)</small> - Bear Lake springsnail
- Pyrgulopsis pisteri <small>Hershler & Sada, 1987</small> - Median-gland Nevada springsnail
- Pyrgulopsis robusta <small>(Walker, 1908)</small> - Jackson Lake springsnail
- Pyrgulopsis roswellensis <small>(Taylor, 1987)</small> - Roswell springsnail
- Pyrgulopsis sanchezi <small>Hershler, Liu & Bradford, 2013</small>
- Pyrgulopsis saxatilis <small>(Hershler, 1998)</small> - sub-globose snake pyrg
- Pyrgulopsis similis <small>(Hershler, Ratcliffe, Liu, Lang and Hay, 2014)</small>
- Pyrgulopsis simplex <small>Hershler, 1988</small> - Fossil springsnail
- Pyrgulopsis sola <small>Hershler, 1988</small> - brown springsnail
- Pyrgulopsis stearnsiana <small>(Pilsbry, 1899)</small> - Yaqui springsnail
- Pyrgulopsis taylori <small>Hershler, 1995</small> - San Luis Obispo pyrg
- Pyrgulopsis texana <small>(Pilsbry, 1935)</small> - Phantom cave snail
- Pyrgulopsis thermalis <small>(Taylor, 1987)</small> - New Mexico hot springsnail
- Pyrgulopsis thompsoni <small>Hershler, 1988</small> - Huachuca springsnail
- Pyrgulopsis transversa <small>(Hershler, 1998)</small> - southern Bonneville pyrg
- Pyrgulopsis trivialis <small>(Taylor, 1987)</small> - Three Forks springsnail
- Pyrgulopsis turbatrix <small>Hershler, 1998</small> - Southeast Nevada Pyrg
- Pyrgulopsis variegata () - Northwest Bonneville pyrg
- Pyrgulopsis ventricosa <small>Hershler, 1995</small> - Clear Lake pyrg
- Pyrgulopsis wongi <small>Hershler, 1989</small>
Eastern North American species of Pyrgulopsis are considered to be in separate genus Marstonia according to the Thompson and Hershler (2002).
- Pyrgulopsis agarhecta <small>(F. G. Thompson, 1969)</small> - Ocmulgee marstonia - Marstonia agarhecta <small>F.G. Thompson, 1969</small>
- Pyrgulopsis arga - ghost marstonia - Marstonia arga <small>F.G. Thompson, 1977</small>
- Pyrgulopsis castor <small>(F. G. Thompson, 1977)</small> - beaverpond marstonia - Marstonia castor <small>F.G. Thompson, 1977</small>
- Pyrgulopsis halcyon <small>(F. G. Thompson, 1977)</small> - halcyon marstonia - Marstonia halcyon <small>F.G. Thompson, 1977</small>
- Pyrgulopsis hershleri <small>F. G. Thompson, 1995</small> - Coosa pyrg - Marstonia hershleri <small>(F.G. Thompson, 1995)</small>
- Pyrgulopsis letsoni <small>(Walker, 1901)</small> - gravel pyrg - Marstonia letsoni <small>(Walker, 1901)</small>
- Pyrgulopsis lustrica <small>(Pilsbry, 1890)</small> - boreal marstonia - Marstonia lustrica <small>(Pilsbry, 1890)</small>
- Pyrgulopsis ogmoraphe <small>(F. G. Thompson, 1977)</small> - royal springsnail - Marstonia ogmorhaphe <small>(F.G. Thompson, 1977)</small>
- Pyrgulopsis olivacea <small>(Pilsbry, 1895)</small> - olive marstonia - Marstonia olivacea <small>(Pilsbry, 1895)</small>
- Pyrgulopsis ozarkensis <small>Hinkley, 1915</small> - Ozark pyrg - Marstonia ozarkensis <small>(Hinkley, 1915)</small>
- Pyrgulopsis pachyta <small>(F. G. Thompson, 1977)</small> - armored marstonia - Marstonia pachyta <small>F.G. Thompson, 1977</small>
- Pyrgulopsis scalariformis <small>(Wolf, 1869)</small> - moss pyrg - Marstonia scalariformis <small>(Wolf, 1869)</small>
References
This article incorporates public domain text from the reference.
Further reading
External links