Protandrena is a genus of mining bees in the family Andrenidae. Depending upon whose definition of the genus one follows, there are anywhere from 50 to 180 described species in Protandrena; traditional classifications recognize 7 subgenera, some of which are sometimes elevated to genus rank, and other classifications place many of these species in the related genus Pseudopanurgus (e.g.), leaving Protandrena with a much smaller constituency. In the most inclusive definition, they are found from Canada through Argentina. However, there is current disagreement whether the Protandrena in South America belong to different genera, in which case the genus extends only as far south as Panama.
Ecology
They are solitary bees, but some species nest in aggregations. They prefer to nest in sunny areas with sparse vegetation. The underground nests have cells lined with a chemical substance. This "wallpaper" acts as a barrier between fungi and bacteria. The eggs hatch, the larvae develop, and then overwinter as mature larvae with hardened skin. They are primarily active from May to October, but have been noted to be active in April in the region six of the United States. Cuckoo bees in the genus Holcopasites have been found as kleptoparasites in Protandrena nests.
There are specialists and generalists found in the genus Protandrena. One notable specialist is Protandrena abdominalis, a specialist on Monarda punctata (bee balm) that is placed in its own subgenus, Metapsaenythia.
Morphology
Protandrena are typically slender black bees. They frequently have yellow on the face and pronotum. They may have red on the metasoma. They rarely have a green or blue tint. The forewings will have two or three submarginal cells. The photo to the right shows submarginal cells (on Lasioglossum). The three submarginal cells are near the top of the wing.
Other Identifying characteristics include
- middle tibial spur on the female that is finely toothed basally and becomes coarser distally; an exception to this is the South American subgenus Austropanurgus, where the spur is finely toothed throughout.
- The male's last abdominal sternum has a pair of large distal lobes that are constricted at the base.
- The gonostyli are more than half as long as the gonocoxites, with the apices articulated or partly fused. The South American subgenus Parasarus is an exception to this, with the gonostyli being less than 1/3 as long as the .
Etymology
Protandrena means "basic Andrena". This is referring to the facial similarities to Andrena
Selected species
- Protandrena abdominalis <small>(Cresson, 1878)</small><span style="color:gray"><sup> i c g</sup></span>
- Protandrena amplipennis <small>Timberlake, 1976</small><span style="color:gray"><sup> i c g</sup></span>
- Protandrena angusticeps <small>Timberlake, 1976</small><span style="color:gray"><sup> i c g</sup></span>
- Protandrena avulsa <small>Ramos & Melo, 2006</small><span style="color:gray"><sup> i c g</sup></span>
- Protandrena bachue <small>Gonzalez & Ruz, 2007</small><span style="color:gray"><sup> i c g</sup></span>
- Protandrena bancrofti <small>Dunning, 1897</small><span style="color:gray"><sup> i c g</sup></span>
- Protandrena bicolor <small>(Timberlake, 1955)</small><span style="color:gray"><sup> i c g</sup></span>
- Protandrena bishoppi <small>Crawford, 1916</small><span style="color:gray"><sup> i c g</sup></span>
- Protandrena blandula <small>Timberlake, 1976</small><span style="color:gray"><sup> i c g</sup></span>
- Protandrena boharti <small>Timberlake, 1976</small><span style="color:gray"><sup> i c g</sup></span>
- Protandrena cockerelli <small>Dunning, 1897</small><span style="color:gray"><sup> i c g b</sup></span>
- Protandrena cognata <small>Timberlake, 1976</small><span style="color:gray"><sup> i c g</sup></span>
- Protandrena duplonotata <small>(Timberlake, 1955)</small><span style="color:gray"><sup> i c</sup></span>
- Protandrena durangoensis <small>Timberlake, 1976</small><span style="color:gray"><sup> i c g</sup></span>
- Protandrena eclepta <small>Timberlake, 1976</small><span style="color:gray"><sup> i c g</sup></span>
- Protandrena euphorbiae <small>(Timberlake, 1955)</small><span style="color:gray"><sup> i c</sup></span>
- Protandrena exigua <small>Timberlake, 1976</small><span style="color:gray"><sup> i c g</sup></span>
- Protandrena fasciata <small>Timberlake, 1976</small><span style="color:gray"><sup> i c g</sup></span>
- Protandrena foveata <small>Timberlake, 1976</small><span style="color:gray"><sup> i c g</sup></span>
- Protandrena guarnensis <small>Gonzalez & Ruz, 2007</small><span style="color:gray"><sup> i c g</sup></span>
- Protandrena heteromorpha <small>(Cockerell, 1896)</small><span style="color:gray"><sup> i c g</sup></span>
- Protandrena hurdi <small>Timberlake, 1976</small><span style="color:gray"><sup> i c g</sup></span>
- Protandrena impressa <small>Timberlake, 1976</small><span style="color:gray"><sup> i c g</sup></span>
- Protandrena irwini <small>Timberlake, 1976</small><span style="color:gray"><sup> i c g</sup></span>
- Protandrena kansensis <small>Timberlake, 1976</small><span style="color:gray"><sup> i c g</sup></span>
- Protandrena lateralis <small>Timberlake, 1976</small><span style="color:gray"><sup> i c g</sup></span>
- Protandrena leucopus <small>Timberlake, 1976</small><span style="color:gray"><sup> i c g</sup></span>
- Protandrena lipovskyi <small>Timberlake, 1976</small><span style="color:gray"><sup> i c g</sup></span>
- Protandrena maculata <small>Timberlake, 1976</small><span style="color:gray"><sup> i c g</sup></span>
- Protandrena marstoni <small>Timberlake, 1976</small><span style="color:gray"><sup> i c g</sup></span>
- Protandrena maurula <small>(Cockerell, 1896)</small><span style="color:gray"><sup> i c g</sup></span>
- Protandrena maximina <small>Gonzalez & Ruz, 2007</small><span style="color:gray"><sup> i c g</sup></span>
- Protandrena metanotalis <small>Timberlake, 1976</small><span style="color:gray"><sup> i c g</sup></span>
- Protandrena mexicanorum <small>(Cockerell, 1896)</small><span style="color:gray"><sup> i c g b</sup></span>
- Protandrena modesta <small>(Smith, 1879)</small><span style="color:gray"><sup> i c g</sup></span>
- Protandrena nudescens <small>Timberlake, 1976</small><span style="color:gray"><sup> i c g</sup></span>
- Protandrena pectidis <small>(Timberlake, 1955)</small><span style="color:gray"><sup> i c g</sup></span>
- Protandrena pernitens <small>Timberlake, 1976</small><span style="color:gray"><sup> i c g</sup></span>
- Protandrena persimilis <small>Timberlake, 1976</small><span style="color:gray"><sup> i c g</sup></span>
- Protandrena polita <small>Timberlake, 1976</small><span style="color:gray"><sup> i c g</sup></span>
- Protandrena protuberata <small>Timberlake, 1976</small><span style="color:gray"><sup> i c g</sup></span>
- Protandrena punctulata <small>Timberlake, 1976</small><span style="color:gray"><sup> i c g</sup></span>
- Protandrena rangeli <small>Gonzalez & Ruz, 2007</small><span style="color:gray"><sup> i c g</sup></span>
- Protandrena scutellata <small>Cockerell, 1916</small><span style="color:gray"><sup> i c g</sup></span>
- Protandrena semilevis <small>Timberlake, 1976</small><span style="color:gray"><sup> i c g</sup></span>
- Protandrena skinneri <small>Timberlake, 1976</small><span style="color:gray"><sup> i c g</sup></span>
- Protandrena sorocula <small>Timberlake, 1976</small><span style="color:gray"><sup> i c g</sup></span>
- Protandrena sphaeralceae <small>Timberlake, 1976</small><span style="color:gray"><sup> i c g</sup></span>
- Protandrena swenki <small>Crawford, 1913</small><span style="color:gray"><sup> i c g</sup></span>
- Protandrena tessellata <small>Timberlake, 1976</small><span style="color:gray"><sup> i c g</sup></span>
- Protandrena texana <small>Timberlake, 1976</small><span style="color:gray"><sup> i c g</sup></span>
- Protandrena tidestromiae <small>Timberlake, 1976</small><span style="color:gray"><sup> i c g</sup></span>
- Protandrena trifoliata <small>(Cockerell, 1896)</small><span style="color:gray"><sup> i c g</sup></span>
- Protandrena trilobata <small>Timberlake, 1976</small><span style="color:gray"><sup> i c g</sup></span>
- Protandrena unimaculata <small>Timberlake, 1976</small><span style="color:gray"><sup> i c g</sup></span>
- Protandrena verbesinae <small>(Timberlake, 1955)</small><span style="color:gray"><sup> i c g</sup></span>
- Protandrena wayruronga <small>Gonzalez & Ruz, 2007</small><span style="color:gray"><sup> i c g</sup></span>
- Protandrena xestops <small>Timberlake, 1976</small><span style="color:gray"><sup> i c g</sup></span>
<small>Data sources: i = ITIS, c = Catalogue of Life, g = GBIF, b = Bugguide.net</small>
References
Further reading
External links