Philotheca is a genus of about fifty species of flowering plants in the family Rutaceae. Plants in this genus are shrubs with simple leaves arranged alternately along the stems, flowers that usually have five sepals, five petals and ten stamens that curve inwards over the ovary. All species are endemic to Australia and there are species in every state, but not the Northern Territory.
Description
Plants in the genus Philotheca are shrubs that are either or have tiny, simple hairs. The leaves are arranged alternately along the stems, narrow oblong to almost cylindrical and sessile or on a very short petiole. From a single to many flowers are arranged in leaf s or on the ends of the branchlets. The flowers have five sepals and five petals (except in P. virgata which has four). The sepals are free from each other and the petals usually overlap at their bases. There are ten stamens that curve inwards over the ovary with anthers that have an appendage called the "apiculum". The ovary contains five carpels fused near their bases. The seeds are long and are released explosively from their capsule.
Taxonomy and naming
The genus Philotheca was first formally described in 1816 by Edward Rudge from a specimen collected near Port Jackson and the description was published in Transactions of the Linnean Society of London. The first species Rudge described was P. australis but this name is considered a nomen illegitimum and a taxonomic synonym of Philotheca salsolifolia by the Australian Plant Census.
The name Philotheca should have been written Psilotheca after the Ancient Greek words psilos meaning "bare", "smooth", "bald" or "naked" and theke meaning "case", "container", "envelope" or "sheath", referring to "the smooth tube of the stamens".
Many plants formerly in Eriostemon are now in this genus.
Distribution
Plants in the genus Philotheca are found in every state of Australia, but not in the Northern Territory.
Species
The following is a list of Philotheca species accepted by the Australian Plant Census as at April 2019:
- Philotheca acrolopha<small> Paul G.Wilson</small> (Qld.)
- Philotheca angustifolia<small> (Paul G. Wilson) Paul G. Wilson</small> â narrow-leaf wax flower
- Philotheca angustifolia <small> (Paul G. Wilson) Paul G. Wilson</small> subsp. angustifolia (Vic., S.A.)
- Philotheca angustifolia subsp. montana<small> (Paul G. Wilson) Paul G. Wilson</small> (Vic.)
- Philotheca apiculata<small> (Paul G. Wilson) Paul G. Wilson</small> (W.A.)
- Philotheca basistyla<small> Mollemans</small> (W.A.)
- Philotheca brevifolia<small> (A.Cunn. ex Endl.) Paul G. Wilson</small> (N.S.W.)
- Philotheca brucei<small> (F.Muell.) Paul G.Wilson</small> (W.A.)
- Philotheca brucei subsp. brevifolia<small> (Paul G. Wilson) Paul G. Wilson</small>
- Philotheca brucei <small> (F.Muell.) Paul G. Wilson</small> subsp. brucei
- Philotheca brucei subsp. cinerea<small> (Paul G. Wilson) Paul G. Wilson</small>
- Philotheca buxifolia<small> (Sm.) Paul G.Wilson</small> (N.S.W.)
- Philotheca buxifolia <small> (Sm.) Paul G. Wilson</small> subsp. buxifolia â box-leaf wax flower
- Philotheca buxifolia subsp. falcata<small> Paul G. Wilson</small>
- Philotheca buxifolia subsp. obovata<small> (G.Don) Paul G. Wilson</small>
- Philotheca ciliata<small> Hook.</small> (Qld., N.S.W.)
- Philotheca citrina<small> Paul G.Wilson</small> (W.A.)
- Philotheca coateana<small> Paul G. Wilson</small> (W.A.)
- Philotheca coccinea<small> (C.A.Gardner) Paul G.Wilson</small> (W.A.)
- Philotheca conduplicata<small> (Paul G. Wilson) P.I.Forst.</small> (Qld., N.S.W.)
- Philotheca cuticularis<small> Paul G. Wilson</small> (Qld.)
- Philotheca cymbiformis<small> (Paul G.Wilson) Paul G.Wilson</small> (W.A.)
- Philotheca deserti<small> (E. Pritzel) Paul G.Wilson</small> (W.A.)
- Philotheca deserti subsp. brevifolia <small>Paul G.Wilson</small>
- Philotheca deserti <small>(E.Pritz.) Paul G.Wilson</small> subsp. deserti
- Philotheca difformis<small> (A.Cunn. ex Endl.) Paul G.Wilson</small>
- Philotheca difformis <small> (A.Cunn. ex Endl.) Paul G. Wilson</small> subsp. difformis â small-leaf wax-flower (Qld., N.S.W., Vic., S.A.)
- Philotheca difformis subsp. smithiana<small> (Benth.) Paul G. Wilson</small> (Qld., N.S.W.)
- Philotheca epilosa<small> (Paul G. Wilson) P.I.Forst.</small> (Qld., N.S.W.)
- Philotheca eremicola<small> Paul G.Wilson</small> (W.A.)
- Philotheca ericifolia<small> (A.Cunn. ex Benth.) Paul G.Wilson</small> (N.S.W.)
- Philotheca falcata<small> (Paul G.Wilson) Paul G.Wilson</small> (W.A.)
- Philotheca fitzgeraldii<small> (C.R.P.Andrews) Paul G.Wilson</small> (W.A.)
- Philotheca freyciana<small> Rozefelds</small> (Tas.)
- Philotheca gardneri<small> (Paul G.Wilson) Paul G.Wilson</small> (W.A.)
- Philotheca gardneri <small> (A.Cunn. ex Endl.) Paul G. Wilson</small> subsp. gardneri
- Philotheca gardneri subsp. globosa<small> Paul G. Wilson</small>
- Philotheca glabra<small> (Paul G.Wilson) Paul G.Wilson</small> (W.A.)
- Philotheca glasshousiensis<small> (Domin) P.I.Forst.</small> (Qld.)
- Philotheca hispidula<small> (Sieber ex Spreng.) Paul G.Wilson</small> (N.S.W.)
- Philotheca kalbarriensis<small> Paul G.Wilson</small> (W.A.)
- Philotheca langei<small> Mollemans</small> (W.A.)
- Philotheca linearis<small> (A.Cunn. ex Endl.) Paul G.Wilson</small> (W.A., S.A., N.S.W.)
- Philotheca myoporoides<small> (DC.) M.J.Bayly</small>
- Philotheca myoporoides subsp. acuta<small> (Blakely) Bayly</small> (N.S.W.)
- Philotheca myoporoides subsp. brevipedunculata<small> Bayly</small> (N.S.W.)
- Philotheca myoporoides subsp. euroensis<small> Bayly</small> (Vic.)
- Philotheca myoporoides <small> (DC.) Bayly</small> subsp. myoporoides (N.S.W., Vic.)
- Philotheca myoporoides subsp. petraea <small> Rozefelds</small> (Vic.)
- Philotheca nodiflora<small> (Lindl.) Paul G.Wilson</small> (W.A.)
- Philotheca nodiflora subsp. calycina<small> (Turcz.) Paul G.Wilson</small>
- Philotheca nodiflora subsp. lasiocalyx<small> (Domin) Paul G.Wilson</small>
- Philotheca nodiflora subsp. latericola<small> Paul G.Wilson</small>
- Philotheca nodiflora <small> (Lindl.) Paul G.Wilson</small> subsp. nodiflora
- Philotheca nutans<small> (Paul G.Wilson) Paul G.Wilson</small> (W.A.)
- Philotheca obovalis<small> (A.Cunn.) Paul G.Wilson</small> (N.S.W.)
- Philotheca obovatifolia<small> (Bayly) P.I.Forst.</small> (Qld., N.S.W.)
- Philotheca pachyphylla<small> (Paul G.Wilson) Paul G.Wilson</small> (W.A.)
- Philotheca papillata<small> I.Telford & L.M.Copel.</small> (N.S.W.)
- Philotheca pinoides<small> (Paul G.Wilson) Paul G.Wilson</small> (W.A.)
- Philotheca pungens<small> (Lindl.) Paul G.Wilson</small> â prickly waxflower (Vic., S.A.)
- Philotheca queenslandica<small> (C.T.White) P.I.Forst.</small> (Qld.)
- Philotheca reichenbachii<small> Sieber ex Spreng.</small> (N.S.W.)
- Philotheca rhomboidea<small> (Paul G.Wilson) Paul G.Wilson</small> (W.A.)
- Philotheca salsolifolia<small> (Sm.) Druce</small> (N.S.W., A.C.T.)
- Philotheca salsolifolia subsp. pedicellata<small> Paul G.Wilson</small>
- Philotheca salsolifolia <small> (Sm.) Druce</small> subsp. salsolifolia
- Philotheca scabra<small> (Paxton) Paul G.Wilson</small> (N.S.W.)
- Philotheca scabra subsp. latifolia<small> (Paul G.Wilson) Paul G.Wilson</small>
- Philotheca scabra <small> (Paxton) Paul G.Wilson </small>subsp. scabra
- Philotheca sericea<small> (Paul G.Wilson) Paul G.Wilson</small> (W.A.)
- Philotheca spicata<small> (A.Rich.) Paul G.Wilson</small> (W.A.)
- Philotheca sporadica<small> (M.J.Bayly) Paul G.Wilson</small> (Qld.)
- Philotheca thryptomenoides<small> (S.Moore) Paul G.Wilson</small> (W.A.)
- Philotheca tomentella<small> (Diels) Paul G.Wilson</small> (W.A.)
- Philotheca trachyphylla<small> (F.Muell.) Paul G.Wilson</small> (N.S.W., Vic.)
- Philotheca tubiflora<small> A.S.George</small> (W.A.)
- Philotheca verrucosa<small> (A.Rich.) Paul G.Wilson</small> (Vic., S.A., Tas.)
- Philotheca virgata<small> (Hook.f.) Paul G.Wilson</small> (N.S.W., Vic., Tas.)
- Philotheca wonganensis<small> (Paul G.Wilson) Paul G.Wilson</small> (W.A.)
References