Paoliida

Paoliida is an extinct order of winged insects that lived in the late Paleozoic. Historically, both their systematic position and composition were controversial – for instance they had been considered as palaeodictyopterans, as basal Neoptera, or as stem-group of Pterygota – but recent studies have resolved them as the sister group of Dictyoptera.

Families and genera

After Prokop et al. (2014), Guan et al. (2016) and Nel & Poschmann (2021):

• †Anthracoptilidae <small>Handlirsch, 1922</small> (Synonyms: Strephocladidae <small>Martynov, 1938</small>; Strephoneuridae <small>Martynov, 1940</small>; ?Adiphlebiidae <small>Handlirsch, 1906</small>)
• ?†Adiphlebia <small>Scudder, 1885</small>
• †Afrocladus <small>Nel, Garrouste & Prokop, 2015</small>
• †Anthracoptilus <small>Lameere, 1917</small>
• †Carrizocladus <small>Rasnitsyn in Rasnitsyn et al., 2004</small>
• †Graticladus <small>Novokshonov and Aristov, 2004</small>
• †Homocladus <small>Carpenter, 1966</small>
• †Jarmilacladus <small>Rasnitsyn and Aristov, 2004</small>
• †Lodevocladus <small>Prokop et al., 2015</small>
• †Mesoptilus <small>Lameere, 1917</small>
• †Mycteroptila <small>Rasnitsyn, 1977</small>
• †Paracladus <small>Carpenter, 1966</small>
• †Pruvostia <small>Bolton, 1921</small>
• †Pseudomesoptilus <small>Guan et al., 2016</small>
• †Rhinomaloptila <small>Rasnitsyn, 1977</small>
• †Spargoptilon <small>Kukalová, 1965</small>
• †Strephocladus <small>Scudder, 1885</small>
• †Strephoneura <small>Martynov, 1940</small>
• †Strephoptilus <small>Rasnitsyn and Aristov, 2013</small>
• †Westphaloptilus <small>Guan et al., 2016</small>
• †Blattinopsidae <small>Bolton, 1925</small> (Synonym: Klebsiellidae <small>Handlirsch, 1906</small>)
• †Alienus <small>Handlirsch, 1906</small>
• †Avionblattinopsis <small>Quispe et al., 2021</small>
• †Balduriella <small>Meunier, 1925</small>
• †Blattinopsis <small>Giebel, 1867</small>
• †Glaphyrophlebia <small>Handlirsch, 1906</small>
• †Klebsiella <small>Meunier, 1908</small>
• †Microblattina <small>Scudder, 1895</small>
• †Protoblattiniella <small>Meunier, 1912</small>
• †Rhipidioptera <small>Brongniart, 1894</small>
• †Stephanopsis <small>Kukalová, 1958</small>
• †Westphaloblattinopsis <small>Béthoux and Jarzembowski, 2010</small>
• †Paoliidae <small>Handlirsch, 1906</small> (Synonyms: Ideliidae <small>Zalessky, 1928</small>; Herbstialidae <small>Schmidt, 1953</small>; Permulidae <small>Aristov & Storozhenko, 2011</small>; Permotermopsidae <small>Martynov, 1937</small>)
• †Acropermula <small>Aristov and Storozhenko, 2011</small>
• †Aenigmidelia <small>Sharov, 1961</small>
• †Archidelia <small>Sharov, 1961</small>
• †Carbonidelia <small>Nel, Garrouste & Jouault, 2023</small>
• †Darekia <small>Prokop et al., 2012</small>
• †Herbstiala <small>Schmidt, 1952</small>
• †Holasicia <small>Kukalová, 1958</small>
• †Idelina <small>Storozhenko, 1992</small>
• †Kemperala <small>Brauckmann, 1984</small>
• †Khosaridelia <small>Storozhenko, 1992</small>
• †Kortshakolia <small>Sharov, 1961</small>
• †Kochopteron <small>Brauckmann, 1984</small>
• †Mertovia <small>Prokop and Nel, 2007</small>
• †Micaidelia <small>Aristov 2004</small>
• †Mongoloidelia <small>Storozhenko, 1992</small>
• †Olinka <small>Kukalová, 1958</small>
• ?†Paolekia <small>Riek, 1976</small>
• †Paolia <small>Smith, 1871</small>
• †Paoliola <small>Handlirsch, 1919</small>
• †Permotermopsis <small>Martynov, 1937</small>
• †Permula <small>Handlirsch, 1919</small>
• †Protoblattina <small>Meunier, 1909</small>
• †Pseudofouquea <small>Handlirsch, 1906</small>
• †Silesiapteron <small>Prokop et al., 2013</small>
• †Simplexpaolia <small>Santos et al., 2023</small>
• †Sojanidelia <small>Storozhenko, 1992</small>
• †Stenaropodites <small>Martynov, 1928</small>
• †Sustaia <small>Kukalová, 1958</small>
• †Sylvidelia <small>Martynov, 1940</small>
• †Zdenekia <small>Kukalová, 1958</small>

References