Pachygnatha is a genus of long-jawed orb-weavers that was first described by Carl Jakob Sundevall in 1823.
Distribution
Spiders in this genus are found in Eurasia, Africa and North America.
Life style
Pachygnatha species are typically found close to the ground in leaf litter, hollow plant stems, under stones, dead wood, or leaves. They are usually more numerous in damp areas.
Adult spiders are secondarily webless, with only immature spiders constructing orb webs.
African species
Very little is known about the biology and behaviour of Afrotropical species. Studies have shown that Pachygnatha species are found mainly in the lower litter layer, with Pachygnatha leleupi being the most common species. This species occurs throughout equatorial Africa at high altitudes, from Cameroon to Malawi. Specimens were much more abundant where water was available.
Description
The carapace bears eyes that are closely grouped, with median eyes separated by their own diameter. The chelicerae are notably large and consistently feature three teeth on the anterior margin in females. In males, the chelicerae are modified and expanded, sometimes bearing a spur on the outside above the base of the fang. The abdomen is subspherical and slightly flattened.
The dorsal surface displays a folium bordered by dark marks with adjacent silver spots. The folium typically has posterior lobes, and the cardiac mark is sometimes outlined by dark pigment or features a median dark mark. Paired white spots are often present. The legs are rather thin and lack macrosetae.
Species
, this genus includes 44 species and two subspecies:
- Pachygnatha amurensis <small>Strand, 1907</small> â Russia (Far East), China
- Pachygnatha atromarginata <small>Bosmans & Bosselaers, 1994</small> â Cameroon
- Pachygnatha autumnalis <small>Marx, 1884</small> â Canada, United States, Cuba
- Pachygnatha bispiralis <small>Nzigidahera & , 2014</small> â Burundi
- Pachygnatha bonneti <small>Senglet, 1973</small> â Spain
- Pachygnatha brevis <small>Keyserling, 1884</small> â Canada, United States
- Pachygnatha calusa <small>Levi, 1980</small> â United States
- Pachygnatha clercki <small>Sundevall, 1823</small> â North America, Europe, Caucasus, Russia (Europe to Far East), Kazakhstan, Iran, Central Asia, China, Korea, Japan (type species)
- Pachygnatha clerckoides <small>Wunderlich, 1985</small> â Albania, North Macedonia, Bulgaria, Russia (Europe, Caucasus), Iran
- Pachygnatha degeeri <small>Sundevall, 1830</small> â Azores, Europe, Turkey, Caucasus, Russia (Europe to Far East), Iran, Central Asia, China
- P. d. dysdericolor <small>Jocqué, 1977</small> â Morocco
- Pachygnatha dorothea <small>McCook, 1894</small> â Canada, United States
- Pachygnatha fengzhen <small>Zhu, Song & Zhang, 2003</small> â China
- Pachygnatha furcillata <small>Keyserling, 1884</small> â Canada, United States
- Pachygnatha goedeli <small>Bosmans & Bosselaers, 1994</small> â Cameroon
- Pachygnatha hexatracheata <small>Bosmans & Bosselaers, 1994</small> â Cameroon
- Pachygnatha intermedia <small>Nzigidahera & Jocqué, 2014</small> â Burundi
- Pachygnatha jansseni <small>Bosmans & Bosselaers, 1994</small> â Cameroon
- Pachygnatha kiwuana <small>Strand, 1913</small> â DR Congo
- Pachygnatha leleupi <small>Lawrence, 1952</small> â Cameroon, DR Congo, Malawi, Zimbabwe, South Africa
- Pachygnatha listeri <small>Sundevall, 1830</small> â Europe, Turkey, Caucasus, Russia (Europe to Far East), Kazakhstan
- Pachygnatha longipes <small>Simon, 1894</small> â Madagascar
- Pachygnatha monticola <small>Baba & , 2018</small> â Japan
- Pachygnatha mucronata <small>Tullgren, 1910</small> â East Africa
- P. m. comorana <small>Schmidt & Krause, 1993</small> â Comoros
- Pachygnatha ochongipina <small>Barrion & , 1995</small> â Philippines
- Pachygnatha okuensis <small>Bosmans & Bosselaers, 1994</small> â Cameroon
- Pachygnatha opdeweerdtae <small>Bosmans & Bosselaers, 1994</small> â Cameroon
- Pachygnatha palmquisti <small>Tullgren, 1910</small> â Kenya, Tanzania
- Pachygnatha procincta <small>Bosmans & Bosselaers, 1994</small> â Cameroon, Burundi, Kenya
- Pachygnatha quadrimaculata <small>( & Strand, 1906)</small> â Russia (Far East), China, Korea, Japan
- Pachygnatha rotunda <small>Saito, 1939</small> â Japan
- Pachygnatha ruanda <small>Strand, 1913</small> â Rwanda
- Pachygnatha shengtangensis <small>Huang, Wu, Yin, Xu & Chen, 2024</small> â China
- Pachygnatha simoni <small>Senglet, 1973</small> â Spain
- Pachygnatha sundevalli <small>Senglet, 1973</small> â Portugal, Spain
- Pachygnatha tenera <small>Karsch, 1879</small> â Russia (Far East), China, Korea, Japan
- Pachygnatha terilis <small>Thaler, 1991</small> â Switzerland, Austria, Italy
- Pachygnatha tristriata <small>C. L. Koch, 1845</small> â Canada, United States
- Pachygnatha tullgreni <small>Senglet, 1973</small> â Portugal, Spain
- Pachygnatha unciniformis <small>Huang, Wu, Yin, Xu & Chen, 2024</small> â China
- Pachygnatha ventricosa <small>Nzigidahera & Jocqué, 2014</small> â Burundi
- Pachygnatha vorax <small>Thorell, 1895</small> â Myanmar
- Pachygnatha xanthostoma <small>C. L. Koch, 1845</small> â Canada, United States
- Pachygnatha zappa <small>Bosmans & Bosselaers, 1994</small> â Cameroon, Kenya, Malawi, South Africa
- Pachygnatha zhui <small>Zhu, Song & Zhang, 2003</small> â China
In synonymy:
- P. gaoi <small>Zhu, Song & Zhang, 2003</small> = Pachygnatha amurensis <small>Strand, 1907</small>
- P. kuratai <small>Levi, 1951</small> = Pachygnatha dorothea <small>McCook, 1894</small>
- P. nipponica <small>(Kishida, 1936)</small> = Pachygnatha quadrimaculata <small>(Bösenberg & Strand, 1906)</small>
- P. sewardi <small>Chamberlin & Ivie, 1947</small> = Pachygnatha clercki <small>Sundevall, 1823</small>
- P. yiliensis <small>(Hu & Wu, 1985)</small> = Pachygnatha degeeri <small>Sundevall, 1830</small>
References