Gracilaria, also known as Irish moss or ogonori, is a genus of red algae in the family Gracilariaceae. It is notable for its economic importance as an agarophyte meaning that it is used to make agar, as well as its use as a food for humans and various species of shellfish. Various species in the genus are cultivated among Asia, South America, Africa and Oceania. They produce over 90% of the world's agar.
Taxonomy
Gracilaria contains the following subtaxa:
- Gracilaria abbottiana <small>M.D.Hoyle</small>
- Gracilaria abyssalis <small>Gurgel & Yoneshigue-Valentin</small>
- Gracilaria aculeata <small>(Hering) Papenfuss</small>
- Gracilaria aggregata <small>Hooker f. & Harvey</small>
- Gracilaria ambigua <small>Greville</small>
- Gracilaria apiculata <small>P.Crouan & H.Crouan</small>
- Gracilaria apiculata subsp. candelabriformis <small>Gurgel, Fredericq & J.N.Norris</small>
- Gracilaria apiculifera <small>J.Agardh</small>
- Gracilaria arcuata f. rhizophora <small>Børgesen</small>
- Gracilaria arcuata var. attenuata <small>Umamaheswara Rao</small>
- Gracilaria arcuata var. snackeyi <small>Weber Bosse</small>
- Gracilaria arcuata <small>Zanardini</small>
- Gracilaria armata <small>(C.Agardh) Greville</small>
- Gracilaria articulata <small>C.F.Chang & B.M.Xia</small>
- Gracilaria ascidiicola <small>E.Y.Dawson</small>
- Gracilaria attenuata <small>(M.Umamaheswara Rao) V.Krishnamurthy</small>
- Gracilaria austromaritima <small>Przhemenstskaya</small>
- Gracilaria babae <small>(H.Yamamoto) P.-K. Ng, P.-E. Lim & S.-M. Phang</small>
- Gracilaria baiana <small>Lyra, Gurgel, M.C.Oliveira & Nunes</small>
- Gracilaria beckeri <small>(J.Agardh) Papenfuss</small>
- Gracilaria birdiae <small>E.M.Plastino & E.C.Oliveira</small>
- Gracilaria blodgettii <small>Harvey</small>
- Gracilaria brasiliensis <small>Gurgel & Yoneshigue-Valentin</small>
- Gracilaria brevis <small>W.R.Taylor</small>
- Gracilaria bursa-pastoris <small>(S.G.Gmelin) P.C.Silva</small>
- Gracilaria camerunensis <small>Pilger</small>
- Gracilaria canaliculata <small>Sonder</small>
- Gracilaria capensis <small>F.Schmitz ex Mazza</small>
- Gracilaria capitata <small>Zanardini</small>
- Gracilaria cearensis <small>(A.B.Joly & Pinheiro) A.B.Joly & Pinheiro</small>
- Gracilaria cerrosiana <small>W.R.Taylor</small>
- Gracilaria cervicornis <small>(Turner) J.Agardh</small>
- Gracilaria changii <small>(B.M.Xia & I.A.Abbott) I.A.Abbott, J.Zhang & B.M.Xia</small>
- Gracilaria chilensis <small>C.J.Bird, McLachlan & E.C.Oliveira</small>
- Gracilaria chondracantha <small>(Kützing) A.J.K.Millar</small>
- Gracilaria chondroides <small>(Kützing) P.Crouan & H.Crouan</small>
- Gracilaria chorda var. exilis <small>Yamamoto</small>
- Gracilaria chouae <small>Zhang & B.M.Xia</small>
- Gracilaria cliftonii <small>Withell, A.J.K.Millar & Kraft</small>
- Gracilaria comosa <small>Withell, A.J.K.Millar & Kraft</small>
- Gracilaria compressa var. lyra <small>J.Agardh</small>
- Gracilaria conferta <small>(Schousboe ex Montagne) Montagne</small>
- Gracilaria confervoides f. ecortica <small>V.M.May</small>
- Gracilaria confervoides var. ramulosa <small>Kützing</small>
- Gracilaria coppejansii <small>Muangmai, Lewmanomont, Prathep, Terada & Zuccarello</small>
- Gracilaria corallicola <small>Zanardini</small>
- Gracilaria cornea <small>J.Agardh</small>
- Gracilaria corniculata <small>(C.Agardh) J.Agardh</small>
- Gracilaria coronopifolia <small>J.Agardh</small>
- Gracilaria corticata <small>(J.Agardh) J.Agardh</small>
- Gracilaria corticata var. cylindrica <small>Umamaheswara Rao</small>
- Gracilaria corticata var. linearis <small>J.Agardh</small>
- Gracilaria corticata var. ramalinioides <small>J.Agardh</small>
- Gracilaria corymbiata <small>(N.RodrÃÂguez de Rios) E.K.Ganesan</small>
- Gracilaria crassa f. conglomerata <small>Børgesen</small>
- Gracilaria crassissima <small>(P.Crouan & H.Crouan) P.Crouan & H.Crouan</small>
- Gracilaria crispata <small>Setchell & Gardner</small>
- Gracilaria crockeri <small>E.Y.Dawson</small>
- Gracilaria cuneata <small>ÃÂÃÂAreschoug</small>
- Gracilaria cuneifolia <small>(Okamura) I.K.Lee & Kurogi</small>
- Gracilaria curtissiae <small>J.Agardh</small>
- Gracilaria cylindrica <small>Børgesen</small>
- Gracilaria damicornis <small>J.Agardh</small>
- Gracilaria dawsonii <small>Hoyle</small>
- Gracilaria debilis <small>(ForsskÃÂ¥l) Børgesen</small>
- Gracilaria dendroides <small>Gargiulo, De Masi & Tripodi</small>
- Gracilaria denticulata <small>F.Schmitz ex Mazza</small>
- Gracilaria disputabilis <small>(M.Bodard) M.Bodard</small>
- Gracilaria disticha <small>(J.Agardh) J.Agardh</small>
- Gracilaria divaricata <small>Harvey</small>
- Gracilaria divergens <small>(C.Agardh) J.Agardh</small>
- Gracilaria domingensis <small>(Kützing) Sonder ex Dickie</small>
- Gracilaria dotyi <small>Hoyle</small>
- Gracilaria dumosa <small>Harvey & Bailey</small>
- Gracilaria dura <small>(C.Agardh) J.Agardh</small>
- Gracilaria dura f. prolificans <small>Reinbold</small>
- Gracilaria dura var. fruticosa <small>J.Agardh</small>
- Gracilaria ecuadoreana <small>(W.R.Taylor) E.Y.Dawson</small>
- Gracilaria edulis <small>(S.G.Gmelin) P.C.Silva</small>
- Gracilaria ephemera <small>Skelton, G.R.South & A.J.K.Millar</small>
- Gracilaria epihippisora <small>Hoyle</small>
- Gracilaria excavata <small>(Setchell & Gardner) G.M.Lyra, C.Iha, M.C. Oliveira, J.M.C.Nunes</small>
- Gracilaria falconii <small>Ardito, Núñez-Resendiz, Dreckmann & SentÃÂes</small>
- Gracilaria fanii <small>B.M.Xia & Pan</small>
- Gracilaria ferox <small>J.Agardh</small>
- Gracilaria firma <small>C.F.Chang & B.-M.Xia</small>
- Gracilaria fisheri <small>(B.M.Xia & I.A.Abbott) I.A.Abbott, J.Zhang & B.M.Xia</small>
- Gracilaria flabelliformis <small>(P.Crouan & H.Crouan) Fredericq & Gurgel</small>
- Gracilaria flabelliformis subsp. aionana <small>Gurgel, Fredericq & J.N.Norris</small>
- Gracilaria flabelliformis subsp. simplex <small>Gurgel, Fredericq & J.N.Norris</small>
- Gracilaria flagelliformis <small>(Sonder) Womersley</small>
- Gracilaria flexuosa <small>E.M.Holmes</small>
- Gracilaria foliifera <small>(ForsskÃÂ¥l) Børgesen</small>
- Gracilaria foliifera f. granatea <small>(J.V.Lamouroux) Børgesen</small>
- Gracilaria fruticosa <small>Harvey</small>
- Gracilaria galetensis <small>Gurgel, Fredericq, & J.N.Norris</small>
- Gracilaria gardneri <small>(Setchell) Gurgel, J.N.Norris & Fredericq</small>
- Gracilaria gigartinoides <small>(P.Crouan & H.Crouan) P.Crouan & H.Crouan</small>
- Gracilaria gigas <small>Harvey</small>
- Gracilaria glomerata <small>Zhang & B.M.Xia</small>
- Gracilaria gracilis <small>(Stackhouse) Steentoft, L.M.Irvine & Farnham</small>
- Gracilaria gurgelii <small>Freshwater</small>
- Gracilaria hainanensis <small>C.F.Chang & B.M.Xia</small>
- Gracilaria halogenea <small>A.J.K.Millar</small>
- Gracilaria hancockii <small>E.Y.Dawson</small>
- Gracilaria hauckii <small>P.C.Silva</small>
- Gracilaria hayi <small>Gurgel, Fredericq & J.N.Norris</small>
- Gracilaria hermonii <small>Millar</small>
- Gracilaria heteroclada <small>(Montagne) J.Feldmann & G.Feldmann</small>
- Gracilaria hikkaduwensis <small>Durairatnam</small>
- Gracilaria howensis <small>A.H.S.Lucas</small>
- Gracilaria huangii <small>S.-M.Lin & De Clerck</small>
- Gracilaria hummii <small>Hommersand & Freshwater</small>
- Gracilaria incrustata <small>J.Agardh</small>
- Gracilaria incurvata <small>Okamura</small>
- Gracilaria indica <small>M.Umamaheswara Rao</small>
- Gracilaria intermedia <small>J.Agardh</small>
- Gracilaria intermedia subsp. ganesanana <small>Gurgel, Fredericq & J.Norris</small>
- Gracilaria isabellana <small>Gurgel, Fredericq & J.N.Norris</small>
- Gracilaria kanyakumariensis <small>Umamaheswara Rao</small>
- Gracilaria khanjanapajiae <small>Saengkaew, Muangmai & Zuccarello</small>
- Gracilaria kilakkaraiensis <small>V.Krishnamurthy & Rajendran</small>
- Gracilaria lacerata <small>Setchell & N.L.Gardner</small>
- Gracilaria lantaensis <small>Muangmai, Zuccarello, Noiraksa & Lewmanomont</small>
- Gracilaria latifrons <small>P.Crouan & H.Crouan</small>
- Gracilaria longa <small>Gargiulo, De Masi & Tripodi</small>
- Gracilaria longirostris <small>Zhang & Wang</small>
- Gracilaria mammillaris <small>(Montagne) M.Howe</small>
- Gracilaria manilaensis <small>Yamamoto & Trono</small>
- Gracilaria mannarensis <small>Umamaheswara Rao</small>
- Gracilaria maramae <small>G.R.South</small>
- Gracilaria marcialana <small>E.Y.Dawson</small>
- Gracilaria mayae <small>A.J.K.Millar</small>
- Gracilaria megaspora <small>(E.Y.Dawson) Papenfuss</small>
- Gracilaria mexicana <small>(Kützing) P.Crouan & H.Crouan</small>
- Gracilaria microcarpa <small>Dreckmann, Núñez-Resendiz & SentÃÂes</small>
- Gracilaria microdendron <small>J.Agardh</small>
- Gracilaria millardetii <small>(Montagne) J.Agardh</small>
- Gracilaria millardetii f. crenulata <small>J.Agardh</small>
- Gracilaria millardetii f. exposita <small>Børgesen</small>
- Gracilaria millardetii f. latifolia <small>Børgesen</small>
- Gracilaria millardetii f. linearifolia <small>J.Agardh</small>
- Gracilaria minor <small>(Sonder) Durairatnam</small>
- Gracilaria minuta <small>Lewmanomont</small>
- Gracilaria mixta <small>I.A.Abbott, J.Zhang & B.M.Xia</small>
- Gracilaria multifurcata <small>Børgesen</small>
- Gracilaria multipartita <small>(Clemente) Harvey</small>
- Gracilaria multipartita f. prolifera <small>E.S.Sinova</small>
- Gracilaria multipartita f. supernediliatata <small>E.S.Sinova</small>
- Gracilaria multipartita var. crispa <small>(Clemente) Cremades</small>
- Gracilaria multipartita var. elongata <small>(Clemente) Cremades</small>
- Gracilaria occidentalis <small>(Børgesen) M.Bodard</small>
- Gracilaria oliveirarum <small>Gurgel, Fredericq & J.N.Norris</small>
- Gracilaria opuntia <small>Durairatnam</small>
- Gracilaria ornata <small>ÃÂÃÂAreschoug</small>
- Gracilaria pachydermatica <small>Setchell & N.L.Gardner</small>
- Gracilaria pacifica <small>I.A.Abbott</small>
- Gracilaria palmettoides <small>P.Crouan & H.Crouan</small>
- Gracilaria papenfussii <small>I.A.Abbott</small>
- Gracilaria parva <small>Freshwater, B.Williamson, P.W.Gabrielson & Margarita Brandt</small>
- Gracilaria parvispora <small>I.A.Abbott</small>
- Gracilaria patens <small>P.Crouan & H.Crouan</small>
- Gracilaria pauciramosa <small>(N.RodrÃÂguez de RÃÂos) A.M.Bellorin, M.C.Oliveira, & E.C.Oliveira</small>
- Gracilaria percurrens <small>(I.A.Abbott) I.A.Abbott</small>
- Gracilaria perplexa <small>K.Byrne & Zuccarello</small>
- Gracilaria peruana <small>Piccone & Grunow</small>
- Gracilaria phuquocensis <small>N.H.Le, N.Muangmai & G.C.Zuccarello</small>
- Gracilaria pinnata <small>Setchell & N.L.Gardner</small>
- Gracilaria preissiana <small>(Sonder) Womersley</small>
- Gracilaria prolifica <small>(Kützing) P.Crouan & H.Crouan</small>
- Gracilaria protea <small>J.Agardh</small>
- Gracilaria pudumadamensis <small>V.Krishnamurthy & N.R.Rajendran</small>
- Gracilaria pulvinata <small>Skottsberg</small>
- Gracilaria punctata <small>(Okamura) Yamada</small>
- Gracilaria pygmaea <small>Børgesen</small>
- Gracilaria ramisecunda <small>E.Y.Dawson</small>
- Gracilaria ramulosa <small>(Martius) Greville</small>
- Gracilaria rangiferina <small>(Kützing) Piccone</small>
- Gracilaria reptans <small>(Weber Bosse) P.C.Silva</small>
- Gracilaria rhodocaudata <small>Yamamoto & Kudo</small>
- Gracilaria rhodymenioides <small>A.J.K.Millar</small>
- Gracilaria robusta <small>Setchell</small>
- Gracilaria rubra <small>C.F.Chang & B.M.Xia</small>
- Gracilaria rubrimembra <small>E.Y.Dawson</small>
- Gracilaria salicornia <small>(C.Agardh) E.Y.Dawson</small>
- Gracilaria salzmannii <small>Bornet ex Möbius</small>
- Gracilaria secunda <small>(C.Agardh) Zanardini</small>
- Gracilaria secundata <small>Harvey</small>
- Gracilaria secundata <small>Setchell & N.L.Gardner</small>
- Gracilaria shimodensis <small>Terada, R.& Yamamoto, H.</small>
- Gracilaria silviae <small>Lyra, Gurgel, M.C.Oliveira & J.M.C.Nunes</small>
- Gracilaria skottsbergii <small>W.R.Taylor</small>
- Gracilaria spinigera <small>E.Y.Dawson</small>
- Gracilaria spinuligera <small>Børgesen</small>
- Gracilaria spinulosa <small>(Okamura) Chang & B.-M.Xia</small>
- Gracilaria srilankia <small>(C.F.Chang & B.M.Xia) A.F.Withell, A.J.K.Millar & Kraft</small>
- Gracilaria stellata <small>I.A.Abbott, Zhang & B.M.Xia</small>
- Gracilaria stipitata <small>A.F.Withell, A.J.K.Millar & Kraft</small>
- Gracilaria sublittoralis <small>Yamada & Segawa ex H.Yamamoto</small>
- Gracilaria subsecundata <small>Setchell & N.L.Gardner</small>
- Gracilaria subtilis <small>(B.M.Xia & I.A.Abbott) B.M.Xia & I.A.Abbott</small>
- Gracilaria sullivanii <small>Yamamoto & Trono</small>
- Gracilaria suzanneae <small>L.P.Soares, C.F.D.Gurgel & M.T.Fujii</small>
- Gracilaria symmetrica <small>E.Y.Dawson</small>
- Gracilaria taiwanensis <small>S.-M.Lin, L.-C.Liu & Payri</small>
- Gracilaria tenuissima <small>Croaun & Crouan</small>
- Gracilaria tenuistipitata <small>C.F.Chang & B.-M.Xia</small>
- Gracilaria tenuistipitata var. liui <small>Zhang & Xia</small>
- Gracilaria tenuistipitata var. tenuistipitata <small>C.F.Chang & B.M.Xia</small>
- Gracilaria tepocensis <small>(E.Y.Dawson) E.Y.Dawson</small>
- Gracilaria textorii <small>(Suringar) Hariot</small>
- Gracilaria textorii f. tenuis <small>V.May</small>
- Gracilaria textorii var. cunninghamii <small>(Farlow) E.Y.Dawson</small>
- Gracilaria tikvahiae <small>McLachlan</small>
- Gracilaria transtasmanica <small>(M.Preuss, N.Muangmai and Zuccarello) G. M.Lyra, C.Iha, J.M.C.Nunes, C.C.Davis</small>
- Gracilaria tridactylites <small>P.Crouan & H.Crouan ex J.Agardh</small>
- Gracilaria truncata <small>Kraft</small>
- Gracilaria tsudae <small>(I.A.Abbott & I.Meneses) I.A.Abbott</small>
- Gracilaria tuberculosa <small>(Hampe ex Kützing) J.Agardh</small>
- Gracilaria turgida <small>E.Y.Dawson</small>
- Gracilaria tuticorinensis <small>V.Krishnamurthy & Rajendran</small>
- Gracilaria urvillei <small>(Montagne) I.A.Abbott</small>
- Gracilaria usneoides <small>(C.Agardh) J.Agardh</small>
- Gracilaria usneoides var. succosa <small>J.Agardh</small>
- Gracilaria vanbosseae <small>(I.A.Abbott) I.A.Abbott</small>
- Gracilaria veloroae <small>E.Y.Dawson</small>
- Gracilaria venezuelensis <small>W.R.Taylor</small>
- Gracilaria vermiculata <small>P.J.L.Dangeard</small>
- Gracilaria vermiculophylla <small>(Ohmi) Papenfuss</small>
- Gracilaria vermiculophylla var. zhengii <small>(J.F.Zhang & B.M.Xia) Yoshida</small>
- Gracilaria verrucosa var. procerrima <small>(Esper) M.P.Reis</small>
- Gracilaria verrucosa var. ramulosa <small>(C.Agardh) M.P.Reis</small>
- Gracilaria vieillardii <small>P.C.Silva</small>
- Gracilaria vieirae <small>M.P.Reis</small>
- Gracilaria viridis <small>A.Sfriso, M.A.Wolf, K.Sciuto, M.Morabito, C.Andreoli & I.Moro</small>
- Gracilaria vivesii <small>M.Howe</small>
- Gracilaria webervanbosseae <small>Huisman & G.W.Saunders</small>
- Gracilaria wrightii <small>(Turner) J.Agardh</small>
- Gracilaria xiae-abbotii <small>G.M.Lyra, J.M.C.Nunes & C.C.Davis</small>
- Gracilaria yamamotoi <small>Zhang & B.M.Xia</small>
- Gracilaria yinggehaiensis <small>Xia & Wang</small>
- Gracilaria yoneshigueana <small>Gurgel, Fredericq & J.N.Norris S </small>
Distribution
Gracilaria are found in warm waters throughout the world, though they also occur seasonally in temperate waters. It can not tolerate temperatures below . Gracilaria are found in all oceans except the Arctic. Their center of diversity is the Western Pacific, where they have been traditionally cultivated as a source of agar.
Use
Gracilaria is used as a food in Filipino, Hawaiian, Japanese, Korean and Sri Lankan cuisines. In Japanese cuisine, it is called ogonori or ogo, and used to make tokoroten. In the Philippines, it is called gulaman and used to make a gelatin substitute. In Jamaica, it is known as Irish moss. In Korea, it is known as kkosiraegi.
Gracilaria oligosaccharides with degree of polymerization 6 prepared by agarase digestion from agar-bearing Gracilaria sp. polysaccharides have been shown to be an effective prophylactic agent during in vitro and in vivo experiments against Japanese encephalitis viral infection. The sulfated oligosaccharides from Gracilaria sp. seem to be promising candidates for further development as antiviral agents.
In the Philippines, Gracilaria have been harvested and used as food for centuries, eaten both fresh or sun-dried and turned into jellies. The earliest historical attestation is from the Vocabulario de la lengua tagala (1754) by the Jesuit priests Juan de Noceda and Pedro de Sanlucar, where golaman or gulaman was defined as "una yerva, de que se haze conserva a modo de Halea, naze en la mar" ("an herb, from which a jam-like preserve is made, grows in the sea"), with an additional entry for guinolaman to refer to food made with the jelly.
In Japan, Gracilaria has been used to produce funori (), an agar-based glue, since the 17th century.
In Sri Lanka, Gracilaria has been used to make a seaweed soup that also incorporates coconut cream and lime. It is also used to create seaweed jelly, a local sweetmeat in the Puttalam District of northwestern Sri Lanka.
Irish moss has long been used as a clarifying agent during the brewing of beer, especially among home brewers, as an alternative to mechanical filtering.
Gracilaria tikvahiae has been used as a model species to determine how well seaweed could be used as a resilient food in sun-blocking scenarios like nuclear winter.
Aquarium trade
Gracilaria commonly appears as a macroalgae for sale in the aquarium trade. It is highly palatable to tangs and many other herbivorous fish, and its nutrient uptake ability makes it a suitable choice for a refugium.
Ecology
Gracilaria are susceptible to infection by the parasitic oomycete Pythium porphyrae. Reproduction by Gracilaria gracilis is supported by Idotea balthica â the first known case of an animal helping algae reproduce.
References
External links