List of Stygnidae species

This is a list of the described species of the harvestman family Stygnidae. The data is taken from Joel Hallan's Biology Catalog.

Heterostygninae

Heterostygninae <small>Roewer, 1913</small>

• Eutimesius <small>Roewer, 1913</small>

* Eutimesius albicinctus <small>(Roewer, 1915)</small> — Venezuela

* Eutimesius ephippiatus <small>(Roewer, 1915)</small> — Colombia

* Eutimesius ornatus <small>(Roewer, 1943)</small> — Colombia, Venezuela

* Eutimesius simoni <small>Roewer, 1913</small> — Brazil, Colombia, Ecuador, Peru

• Imeri <small>Pinto-da-Rocha & Tourinho, 2012</small> (! after Hallan)

* Imeri ajuba <small>Coronato-Ribeiro, Pinto-da-Rocha & Rheims, 2013</small> — Brazil (Amazonas)

* Imeri lomanhungae <small>Pinto-da-Rocha & Tourinho, 2012</small> — Brazil (Amazonas)

• Innoxius <small>Pinto-da-Rocha, 1997</small>

* Innoxius magnus <small>(Caporiacco, 1951)</small> — Venezuela

• Minax <small>Pinto-da-Rocha, 1997</small>

* Minax tetraspinosus <small>Pinto-da-Rocha, 1997</small> — Venezuela

• Stenostygnellus <small>Roewer, 1913</small>

* Stenostygnellus flavolimbatus <small>Roewer, 1913</small> — Venezuela

* Stenostygnellus macrochelis <small>(Roewer, 1917)</small> — Venezuela

• Stygnidius <small>Simon, 1879</small>

* Stygnidius guerinii <small>Sørensen, 1932</small> — French Guayana

* Stygnidius inflatus <small>(Guérin-Méneville, 1829-1843)</small> — Brazil, Venezuela, French Guiana

• Stygnoplus <small>Simon, 1879</small>

* Stygnoplus antiguanus <small>(Roewer, 1943)</small> — Leeward Islands, Antigua

* Stygnoplus biguttatus <small>Pinto-da-Rocha, 1997</small> — Venezuela

* Stygnoplus clavotibialis <small>(Goodnight & Goodnight, 1947)</small> — Trinidad

* Stygnoplus flavitarsis <small>(Simon, 1879)</small> — Leeward Islands, Guadeloupe

* Stygnoplus forcipatus <small>(C.L. Koch, 1845)</small> — Colombia

* Stygnoplus granulosus <small>Mello-Leitão, 1940</small> — Venezuela

* Stygnoplus longipalpus <small>(Goodnight & Goodnight, 1942)</small> — Guyana

* Stygnoplus meinerti <small>Sørensen, 1932</small> — Venezuela

* Stygnoplus triacanthus <small>(C.L. Koch, 1839)</small> — South America

* Stygnoplus tuberculatus <small>(Goodnight & Goodnight, 1942)</small> — Dominica (plus synonym Stygnoplus dominicanus <small>(Roewer, 1943)</small> per Pérez-González, 2000, after Hallan).

• Timesius <small>Simon, 1879</small>

* Timesius vesicularis <small>(Gervais, 1844)</small> — Colombia

• Yapacana <small>Pinto-da-Rocha, 1997</small>

* Yapacana tibialis <small>(Pinto-da-Rocha, 1997)</small> — Venezuela

Stygninae

Stygninae <small>Simon, 1879</small>

• Actinostygnoides <small>Goodnight & Goodnight, 1942</small>

* Actinostygnoides carus <small>Goodnight & Goodnight, 1942</small> — Guyana

• Auranus <small>Mello-Leitão, 1941</small>

* Auranus hoeferscovitorum <small>Pinto-da-Rocha, 1997</small> — Brazil

* Auranus parvus <small>Mello-Leitão, 1941</small> — Brazil

• Fortia <small>Villarreal et al., 2022</small> (! after Hallan)

* Fortia jedi <small>Villarreal et al., 2022</small> — Colombia

* Fortia sith <small>Villarreal et al., 2022</small> — Colombia

• Iguarassua <small>Roewer, 1943</small>

* Iguarassua schubarti <small>Roewer, 1943</small> — Brazil

• Jabbastygnus <small>Kury & Villarreal, 2015</small> (! after Hallan)

* Jabbastygnus huttorum <small>Kury & Villarreal, 2015</small> — Colombia (Boyacá)

• Kaapora <small>Pinto-da-Rocha, 1997</small>

* Kaapora minutissimus <small>(Roewer, 1943)</small> — Brazil

• Metaphareus <small>Roewer, 1912</small>

* Metaphareus albimanus <small>Roewer, 1912</small> — Colombia

* Metaphareus punctatus <small>Roewer, 1913</small> — Venezuela

• Niceforoiellus <small>Mello-Leitão, 1941</small>

* Niceforoiellus assimilis <small>Mello-Leitão, 1941</small> — Colombia

• Obidosus <small>Roewer, 1931</small> (! after Hallan, genus restored per Villarreal et al. 2019)

* Obidosus albilineatus <small>Roewer, 1957</small> — Peru, Ecuador, Brazil (Amazonas)

* Obidosus amigos <small>(Ochoa & Pinto-da-Rocha, 2013)</small> — Peru

* Obidosus amplichelis <small>Roewer, 1931</small> — Brazil (Amazonas)

* Obidosus apiacas <small>(Pinto-da-Rocha, 2000)</small> — Brazil (Mato Grosso)

* Obidosus bahiensis <small>(Pinto-da-Rocha & Villarreal, 2009)</small> — Brazil (Bahía)

* Obidosus boibumba <small>(Villarreal & Pinto-da-Rocha, 2006)</small> — Brazil (Acre), Peru

* Obidosus carnaval <small>(Villarreal & Pinto-da-Rocha, 2006)</small> — Brazil (Acre), Colombia (Amazonas)

* Obidosus cirio <small>(Villarreal & Pinto-da-Rocha, 2006)</small> — Brazil (Pará)

* Obidosus coxalis <small>(Roewer, 1931)</small> — Brazil (Amazonas)

* Obidosus evelineae <small>(Soares & Soares, 1978)</small> — Brazil (Pará)

* Obidosus foliadereis <small>(Villarreal & Pinto-da-Rocha, 2006)</small> — Brazil (Mato Grosso)

* Obidosus jirau <small>(Bragagnolo, 2013)</small> — Brazil (Rondonia)

* Obidosus junina <small>(Villarreal & Pinto-da-Rocha, 2006)</small> — Brazil (Bahía)

* Obidosus kakinte <small>(Ochoa & Pinto-da-Rocha, 2013)</small> — Peru

* Obidosus laevis <small>(Sørensen, 1932)</small> — Brazil (Pará, Matto Grosso)

* Obidosus longipalpis <small>(Roewer, 1943)</small> — Brazil (Amazonas)

* Obidosus machiguenga <small>(Ochoa & Pinto-da-Rocha, 2013)</small> — Peru

* Obidosus mendopictus <small>(Soares, 1978)</small> — Brazil (Bahía)

* Obidosus osvaldoi <small>(Kury & Pinto-da-Rocha, 2008)</small> — Brazil (Espírito Santo)

* Obidosus palpalis <small>(Roewer, 1931)</small> — Brazil (Amazonas)

* Obidosus regalo <small>(Bragagnolo, 2013)</small> — Brazil (Tocantins)

* Obidosus trocaraincola <small>(Pinto-da-Rocha, 1997)</small> — Brazil (Pará)

• Ortonia <small>Wood, 1869</small>

* Ortonia ferox <small>Wood, 1869</small> — Ecuador

• Paraphareus <small>Goodnight & Goodnight, 1943</small>

* Paraphareus tatei <small>Goodnight & Goodnight, 1943</small> — Guyana/Venezuela (Bolívar: Monte Roraima)

• Phareus <small>Simon, 1879</small> (! revised after Hallan)

* Phareus antrophilus <small>Villarreal & Rodríguez-T, 2006</small> — Colombia

* Phareus raptator <small>(Gervais, 1844)</small> — Colombia, Venezuela

• Pickeliana <small>Mello-Leitão, 1932</small> (! revised after Hallan)

* Pickeliana albimaculata <small>Hara & Pinto-da-Rocha, 2008</small>

* Pickeliana capito <small>(Soares & Soares, 1974)</small>

* Pickeliana pickeli <small>Mello-Leitão, 1932</small> — Brazil

• Planophareus <small>Goodnight & Goodnight, 1943</small>

* Planophareus pallidus <small>Goodnight & Goodnight, 1943</small> — Monte Roraima, Brazil

• Protimesius <small>Roewer, 1913</small> (! revised after Hallan)

* Protimesius gracilis <small>Roewer, 1913</small> — Brazil (Amapá), Suriname, French Guiana

* Protimesius lucifer <small>Villarreal, Ázara & Kury, 2019</small> — Brazil (Pará)

* Protimesius orcus <small>Villarreal, Ázara & Kury, 2019</small> — Brazil (Ceará)

• Ricstygnus <small>Kury, 2009</small> (! after Hallan)

* Ricstygnus quineti <small>Kury, 2009</small> — Brazil (Ceará)

• Sickesia <small>Soares, 1979</small>

* Sickesia helmuti <small>Soares, 1979</small> — Brazil

• Stenophareus <small>Goodnight & Goodnight, 1943</small>

* Stenophareus roraimus <small>Goodnight & Goodnight, 1943</small> — Roraima, Brazil

• Stenostygnoides <small>Roewer, 1913</small>

* Stenostygnoides cosmetitarsus <small>Roewer, 1913</small> — Surinam

• Stygnus <small>Perty, 1833</small>

* Stygnus aggerum <small>Sørensen, 1932</small> — Venezuela

* Stygnus armatus <small>Perty, 1833</small> — Brazil

* Stygnus brevispinis <small>Pinto-da-Rocha, 1997</small> — Bolivia

* Stygnus ferrugineus <small>(Perty, 1833)</small> — Brazil, French Guiana

* Stygnus gertschi <small>(Roewer, 1963)</small> — Colombia

* Stygnus grasshoffi <small>Pinto-da-Rocha, 1997</small> — Venezuela

* Stygnus klugi <small>(Goodnight & Goodnight, 1943)</small> — Peru

* Stygnus lesserti <small>(Roewer, 1943)</small> — Brazil

* Stygnus luteus <small>(Mello-Leitão, 1931)</small> — Brazil

* Stygnus marthae <small>Pinto-da-Rocha, 1997</small> — Brazil

* Stygnus mediocris <small>(Roewer, 1931)</small> — Ecuador

* Stygnus multispinosus <small>(Piza, 1938)</small> — Brazil

* Stygnus pectinipes <small>(Roewer, 1943)</small> — Surinam, Brazil, Colombia

* Stygnus peruvianus <small>(Roewer, 1957)</small> — Peru

* Stygnus polyacanthus <small>(Mello-Leitão, 1923)</small> — Brazil

* Stygnus simonis <small>Sørensen, 1932</small> — Peru

* Stygnus simplex <small>(Roewer, 1913)</small> — Ecuador, Colombia, Peru

* Stygnus weyrauchi <small>(Roewer, 1963)</small> — Peru

• Verrucastygnus <small>Pinto-da-Rocha, 1997</small>

* Verrucastygnus caliginosus <small>(R. Pinto-da-Rocha, 1990)</small> — Brazil

incertae sedis

incertae sedis

• Gaibulus <small>Roewer, 1943</small>

* Gaibulus schubarti <small>Roewer, 1943</small> — Brazil

Since excluded

Subsequently transferred are former Stygnidae: Nomoclastinae <small>Roewer, 1943</small>

• Nomoclastes <small>Sørensen, 1932</small>

* Nomoclastes quasimodo <small>Pinto-da-Rocha, 1997</small> — Colombia

* Nomoclastes taedifer <small>Sørensen, 1932</small> — Colombia

Moved to => Nomoclastidae <small>Roewer, 1943</small>

References

• Joel Hallan's Biology Catalog: Stygnidae