Leptoconops (black gnat) is a midge genus in the family Ceratopogonidae. It has a mostly tropical or subtropical distribution worldwide, but some species occur as far north as Moscow region in Russia and the Yukon Territory in Canada.
This genus is relictual, having had a pantropical distribution during the Cretaceous. The presence of Leptoconops, along with Austroconops, in ancient Lebanese amber makes these the earliest existing lineages of biting midges. Extinct species have also been described from amber from Siberia, New Jersey, Canada, Hungary, Sakhalin, France, and Spain.
Adult Leptoconops females are diurnal feeders, and suck vertebrate blood. Adults of both sexes in some species rest by burying themselves in sand. Larvae feed on algae, fungi, and bacteria. They burrow in moist, usually saline, sand or mud of desert areas and coastal and inland beaches.
Species
Leptoconops contains the following species:
- Leptoconops acer <small>Clastrier, 1973</small>
- Leptoconops albiventris <small>de Meijere, 1915</small>
- Leptoconops algeriensis <small>Clastrier, 1975</small>
- Leptoconops altuneshanensis <small>Yu and Shao, 1988</small>
- Leptoconops americanus <small>Carter, 1921</small>
- Leptoconops amplifemoralis <small>Chanthawanich and Delfinado, 1967</small>
- â Leptoconops amplificatus <small>Borkent, 2001</small>
- Leptoconops andersoni <small>Clastrier and Wirth, 1978</small>
- â Leptoconops antiquus <small>Borkent, 2001</small>
- Leptoconops arnaudi <small>Clastrier and Wirth, 1978</small>
- Leptoconops ascius <small>Yu and Hui, 1988</small>
- Leptoconops asilomar <small>Clastrier and Wirth, 1978</small>
- Leptoconops atchleyi <small>Clastrier and Wirth, 1978</small>
- Leptoconops auster <small>Clastrier, 1981</small>
- Leptoconops aviarum <small>Gutsevich, 1973</small>
- Leptoconops bahreinensis <small>Clastrier and Boorman, 1987</small>
- Leptoconops bossoi <small>Clastrier, 1981</small>
- Leptoconops belkini <small>Wirth and Atchley, 1973</small>
- Leptoconops bequaerti <small>(Kieffer), 1925</small>
- Leptoconops bezzii <small>(Noè), 1905</small>
- Leptoconops bidentatus <small>Gutsevich, 1960</small>
- Leptoconops binangulus <small>Yu, 1989</small>
- Leptoconops binisiculus <small>Yu and Liu, 1988</small>
- Leptoconops borealis <small>Gutsevich, 1945</small>
- Leptoconops boreus <small>Kalugina, 1991</small>
- Leptoconops brasiliensis <small>(Lutz), 1913</small>
- Leptoconops brevistylus <small>Mazumdar, Saha & Chaudhuri, 2010</small>
- Leptoconops bullsbrookensis <small>Smee, 1966</small>
- Leptoconops bundyensis <small>Smee, 1966</small>
- â Leptoconops burmiticus <Small>Szadziewski, 2004</small>
- Leptoconops californiensis <small>Wirth and Atchley, 1973</small>
- Leptoconops camelorum <small>(Kieffer), 1921</small>
- Leptoconops capensis <small>de Meillon and Hardy, 1953</small>
- Leptoconops carteri <small>Hoffman, 1926</small>
- Leptoconops casali <small>Cavalieri and Chiossone, 1966</small>
- Leptoconops catawbae <small>(Boesel), 1948</small>
- Leptoconops chenfui <small>Yu and Xiang, 1988</small>
- Leptoconops chilensis <small>Forattini, 1958</small>
- Leptoconops chinensis <small>Sun, 1968</small>
- Leptoconops conulus <small>Yu and Liu, 1990</small>
- Leptoconops copiosus <small>Borkent, 1996</small>
- Leptoconops curvachelus <small>Borkent, 1996</small>
- â Leptoconops daugeroni <small>Choufani, Azar and Nel, 2011</small>
- Leptoconops demeilloni <small>Clastrier and Nevill, 1984</small>
- Leptoconops dissimilis <small>Clastrier, 1975</small>
- Leptoconops dixi <small>de Meillon, 1936</small>
- Leptoconops doyeni <small>Spinelli and Ronderos, 1993</small>
- â Leptoconops ellenbergeri <small>Szadziewski, 2015</small>
- Leptoconops endialis <small>Smee, 1966</small>
- Leptoconops exspectator <small>Clastrier, 1975</small>
- Leptoconops flaviventris <small>Kieffer, 1918</small>
- Leptoconops floridensis <small>Wirth, 1951</small>
- Leptoconops foleyi <small>Clastrier, 1975</small>
- Leptoconops fortipalpus <small>Mazumdar, Saha & Chaudhuri, 2010</small>
- Leptoconops foulki <small>Clastrier and Wirth, 1978</small>
- Leptoconops freeborni <small>Wirth, 1952</small>
- Leptoconops fretus <small>Yu and Zhan, 1990</small>
- Leptoconops fuscipennis <small>Clastrier, Rioux, and Descous, 1961</small>
- Leptoconops gallicus <small>Clastrier, 1973</small>
- Leptoconops golanensis <small>Clastrier, 1981</small>
- Leptoconops grandis <small>Carter, 1921</small>
- â Leptoconops gravesi <Small>Choufani et al., 2014</small>
- Leptoconops halophilus <small>Smee, 1966</small>
- Leptoconops hamariensis <small>Herzi and Sabatini, 1983</small>
- Leptoconops harrisoni <small>de Meillon and Hardy, 1953</small>
- Leptoconops helobius <small>Ma and Yu, 1990</small>
- Leptoconops hutsoni <small>Clastrier, 1974</small>
- Leptoconops hyalinipennis <small>Kieffer, 1918</small>
- Leptoconops indicus <small>(Kieffer), 1918</small>
- Leptoconops interruptus <small>(Enderlein), 1908</small>
- Leptoconops irritans <small>(Noè), 1905</small>
- Leptoconops kerteszi <small>Kieffer, 1908</small>
- Leptoconops kinmenensis <small>Lien, Lin, Weng and Chin, 1996</small>
- Leptoconops knowltoni <small>Clastrier and Wirth, 1978</small>
- Leptoconops lacteipennis <small>Kieffer, 1918</small>
- Leptoconops laosensis <small>Clastrier, 1974</small>
- Leptoconops latibulorum <small>Gutsevich, 1973</small>
- Leptoconops laurae <small>(Weiss), 1912</small>
- Leptoconops linleyi <small>Wirth and Atchley, 1973</small>
- Leptoconops lisbonnei <small>Harant and Galan, 1944</small>
- Leptoconops longicauda <small>Yu, 1997</small>
- Leptoconops longicornis <small>Carter, 1921</small>
- Leptoconops lucidus <small>Gutsevich, 1964</small>
- Leptoconops mackerrassae <small>Smee, 1966</small>
- Leptoconops macfiei <small>Clastrier, 1975</small>
- Leptoconops melanderi <small>Wirth and Atchley, 1973</small>
- Leptoconops mellori <small>Clastrier and Boorman, 1987</small>
- Leptoconops mesopotamiensis <small>(Patton), 1920</small>
- Leptoconops minutus <small>Gutsevich, 1973</small>
- Leptoconops mohavensis <small>Wirth and Atchley, 1973</small>
- Leptoconops montanus <small>Konurbajev, 1965</small>
- Leptoconops montigenus <small>Clastrier, 1981</small>
- Leptoconops mooloolabaensis <small>(Smee), 1966</small>
- Leptoconops myersi <small>(Tonnoir), 1924</small>
- â Leptoconops myanmaricus <Small>Szadziewski, 2004</small>
- Leptoconops nachitschevanicus <small>Dzhafarov, 1961</small>
- Leptoconops nevilli <small>Clastrier, 1981</small>
- Leptoconops nicolayi <small>de Meillon, 1937</small>
- Leptoconops nigripes <small>Dzhafarov, 1961</small>
- Leptoconops nipponensis <small>Tokunaga, 1937</small>
- Leptoconops nivalis <small>Smee, 1966</small>
- Leptoconops noei <small>Clastrier and Coluzzi, 1973</small>
- â Leptoconops nosopheris <small>Poinar, 2008</small>
- Leptoconops obscurus <small>Smee, 1966</small>
- Leptoconops panamensis <small>Ronderos and Spinelli, 1993</small>
- Leptoconops parvichelus <small>Chanthawanich and Delfindao, 1967</small>
- Leptoconops patagoniensis <small>Ronderos, 1990</small>
- Leptoconops pavlovskyi <small>Dzhafarov, 1961</small>
- Leptoconops peneti <small>(Langeron), 1913</small>
- Leptoconops petrocchiae <small>Shannon and Del Ponte, 1927</small>
- Leptoconops primaevus <small>Borkent, 1995</small>
- Leptoconops pseudosetosifrons <small>(Smee), 1966</small>
- Leptoconops pugnax <small>Clastrier, 1973</small>
- Leptoconops reesi <small>Clastrier and Wirth, 1978</small>
- Leptoconops rhodesiensis <small>Carter, 1921</small>
- Leptoconops ricardoi <small>Ronderos and Spinelli, 1992</small>
- Leptoconops riverinaensis <small>Smee, 1966</small>
- â Leptoconops rossi <Small>Szadziewski, 2004</small>
- Leptoconops rufiventris <small>(Kieffer), 1923</small>
- Leptoconops setosifrons <small>(Smee), 1966</small>
- Leptoconops shangweni <small>Xu and Yu, 1989</small>
- Leptoconops siamensis <small>Carter, 1921</small>
- Leptoconops sibericus <small>Szadziewski, 1996</small>
- Leptoconops smeei <small>Wirth and Atchley, 1973</small>
- Leptoconops spinosifrons <small>(Carter), 1921</small>
- Leptoconops stygius <small>Skuse, 1889</small>
- Leptoconops sublettei <small>Clastrier and Wirth, 1978</small>
- Leptoconops succineus <small>Szadziewski, 1988</small>
- Leptoconops tarimensis <small>Yu, 1982</small>
- Leptoconops tenebrostigmatus <small>Mazumdar, Saha & Chaudhuri, 2010</small>
- Leptoconops tibetensis <small>Lee, 1978</small>
- Leptoconops torrens <small>(Townsend), 1893</small>
- Leptoconops transversalis <small>(Kieffer), 1921</small>
- Leptoconops turkmenicus <small>Molotova, 1967</small>
- Leptoconops umbellifer <small>Clastrier, 1981</small>
- Leptoconops vargasi <small>Clastrier and Wirth, 1978</small>
- Leptoconops venezuelensis <small>Ortiz, 1952</small>
- Leptoconops vexans <small>(Kieffer), 1921</small>
- Leptoconops wehaiensis <small>Yu and Xue, 1988</small>
- Leptoconops werneri <small>Wirth and Atchley, 1973</small>
- Leptoconops whitseli <small>Clastrier and Wirth, 1978</small>
- Leptoconops woodhilli <small>Lee, 1948</small>
- Leptoconops xuthosceles <small>Chanthawanich and Delfinado, 1967</small>
- Leptoconops yalongensis <small>Yu and Wang, 1988</small>
- Leptoconops yunhsienensis <small>Yu, 1963</small>
- â Leptoconops zherikhini <small>Szadziewski & Arillo, 2003</small>
References