Ixodes is a genus of hard-bodied ticks (family Ixodidae). It includes important disease vectors of animals and humans (tick-borne disease), and some species (notably Ixodes holocyclus) inject toxins that can cause paralysis. Some ticks in this genus may transmit the pathogenic bacterium Borrelia burgdorferi responsible for causing Lyme disease. Additional organisms that may be transmitted by Ixodes are parasites from the genus Babesia, which cause babesiosis, and bacteria from the related genus Anaplasma, which cause anaplasmosis.
Ecology and distribution
Ixodes species have a near-cosmopolitan distribution and occur in all major biogeographic realms, including Antarctica, where they are associated with seabirds and penguin rookeries. The genus parasitizes a wide range of vertebrate hosts worldwide, including mammals, birds, and reptiles. Rodents and passerine birds are among the most commonly recorded hosts, especially in the Americas.
Description and systematics
Ixodes is the sole representative of the Prostriata clade of the Ixodidae. Prostriate ticks are distinguished from the Metastriata with reference to the position of the anal groove. In Ixodes species, the groove loops anterior to the anus, whereas in Ambloymma, for example, the groove is positioned posterior to the anus. Ixodes species are small to medium-sized ticks, with a usually pyriform (pear-shaped) or ovate body profile. Mouthparts are anterior in both sexes, but usually long and slender in females, and short in males. Males have sclerotised adanal plates. Ixodes species are eyeless.
Taxonomy
Ixodes contains 274 species. Classification, recognition and phylogenetic resolution of the Ixodes subgenera is ongoing. As many as 24 subgenera have been recognised by different authors. All are included here for completeness.
Subgenera
- Afrixodes <small>Morel, 1966</small>
- Alloixodes <small>ÃÂerný, 1969</small>
- Amerixodes <small>Morel, 1998</small>
- Australixodes <small>Barker & Barker et al., 2023</small>
- Ceratixodes <small>Neumann, 1902</small>
- Coxixodes <small>Schulze, 1941</small>'
- Endopalpiger <small>Schulze, 1935</small>
- Eschatocephalus <small>Frauenfeld, 1853</small>
- Exopalpiger <small>Schulze, 1935</small>
- Filippoviella <small>Apanaskevich, Greiman, Fedorov, Ahmed & Barker</small>
- Haemixodes <small>Kohls & Clifford, 1967</small>
- Indixodes <small>Morel, 1998</small>
- Ixodes <small>Latreille, 1795</small>
- Ixodiopsis <small>Filippova (1957)</small>
- Lepidixodes <small>Schulze, 1935</small>
- Monoindex <small>Emelyanova & Kozlovskaya, 1968</small>
- Multidentatus <small>Neumann, 1904</small>
- Partipalpiger <small>Hoogstraal et al., 1973</small>
- Pholeoixodes <small>Schulze, 1942</small>
- Pomerantzevella <small>Feider, 1965</small>
- Scaphixodes <small>Schulze, 1941</small>
- Sternalixodes <small>Schulze, 1935</small>
- Trichotoixodes <small>Reznik, 1961</small>
- Xiphixodes <small>Schulze, 1941</small>
Species
- Ixodes abrocomae <small>Lahille, 1916</small>
- Ixodes (Ixodes) abramovi <small>Apanaskevich, 2024</small>
- Ixodes acer <small>Apanaskevich & Schenk, 2020</small>
- Ixodes acuminatus <small>Neumann, 1901</small>
- Ixodes (Indixodes) acutitarsus <small>(Karsch, 1880)</small>
- Ixodes affinis <small>Neumann, 1899</small>
- Ixodes albignaci <small>Uilenberg and Hoogstraal, 1969</small>
- Ixodes (Ixodes) algericus <small>Keskin, Aftisse and Apanaskevich</small><small>, 2024</small>
- Ixodes alluaudi <small>Neumann, 1913</small>
- Ixodes amarali <small>Fonseca, 1935b</small>
- Ixodes (Afrixodes) ambohitantelensis <small>Englert, Goodman & Apanaskevich, 2023</small>
- Ixodes amersoni <small>Kohls, 1966</small>
- Ixodes (Australixodes) anatis <small>Chilton, 1904</small>
- Ixodes andinus <small>Kohls, 1956a</small>
- Ixodes (Pholeoixodes) angustus <small>Neumann, 1899</small>
- Ixodes antechini <small>Roberts, 1960</small>
- Ixodes apronophorus <small>Schulze, 1924</small>
- Ixodes arabukiensis <small>Arthur, 1959</small>
- Ixodes arboricola <small>Schulze and Schlottke, 1929</small>
- Ixodes arebiensis <small>Arthur, 1956c</small>
- Ixodes ariadnae <small>Hornok, 2014</small>
- Ixodes asanumai <small>Kitaoka, 1973</small>
- Ixodes aulacodi <small>Arthur, 1956c</small>
- Ixodes auriculaelongae <small>Arthur, 1958 </small>
- Ixodes auritulus <small>Neumann, 1904</small>
- Ixodes (Endopalpiger) australiensis <small>Neumann, 1904</small>
- Ixodes baergi <small>Cooley and Kohls, 1942</small>
- Ixodes bakeri <small>Arthur and Clifford, 1961</small>
- Ixodes banksi <small>Bishopp, 1911</small>
- Ixodes (Endopalpiger) barkeri <small>Barker, 2019</small>
- Ixodes bedfordi <small>Arthur, 1959</small>
- Ixodes bequaerti <small>Cooley and Kohls, 1945</small>
- Ixodes berlesei <small>Birula, 1895</small>
- Ixodes bivari <small>Santos Dias, 1990</small>
- Ixodes bocatorensis <small>Apanaskevich and Bermúdez, 2017</small>
- Ixodes boliviensis <small>Neumann, 1904</small>
- Ixodes brewsterae <small>Keirans, Clifford and Walker, 1982</small>
- Ixodes browningi <small>Arthur, 1956</small>
- Ixodes brumpti <small>Morel, 1965</small>
- Ixodes brunneus <small>Koch, 1844</small>
- Ixodes calcarhebes <small>Arthur and Zulu, 1980</small>
- Ixodes caledonicus <small>Nuttall, 1910</small>
- Ixodes canisuga <small>Johnston, 1849</small>
- Ixodes capromydis <small>ÃÂerný, 1966</small>
- Ixodes catherinei <small>Keirans, Clifford and Walker, 1982</small>
- Ixodes cavipalpus <small>Nuttall and Warburton, 1908</small>
- Ixodes ceylonensis <small>Kohls, 1950</small>
- Ixodes chilensis <small>Kohls, 1956</small>
- Ixodes colasbelcouri <small>Arthur, 1957</small>
- Ixodes collaris <small>Hornok, 2016</small>
- Ixodes collocaliae <small>Schulze, 1937</small>
- Ixodes columnae <small>Takada and Fujita, 1992</small>
- Ixodes conepati <small>Cooley and Kohls, 1943</small>
- Ixodes (Sternalixodes) confusus <small>Roberts, 1960</small>
- Ixodes cookei <small>Packard, 1869</small>
- Ixodes cooleyi <small>Aragão and Fonseca, 1951</small>
- Ixodes copei <small>Wilson, 1980</small>
- Ixodes cordifer <small>Neumann, 1908</small>
- Ixodes cornuae <small>Arthur, 1960c</small>
- Ixodes (Sternalixodes) cornuatus <small>Roberts, 1960</small>
- Ixodes cornutus <small>Lotozky, 1956</small>
- Ixodes corwini <small>Keirans, Clifford and Walker, 1982</small>
- Ixodes crenulatus <small>Koch, 1844</small>
- Ixodes cuernavacensis <small>Kohls and Clifford, 1966</small>
- Ixodes cumulatimpunctatus <small>Schulze, 1943</small>
- Ixodes dampfi <small>Cooley, 1943</small>
- Ixodes (Trichotoixodes) daveyi <small>Nuttall, 1913</small>
- Ixodes dawesi <small>Arthur, 1956</small>
- Ixodes dendrolagi <small>Wilson, 1967</small>
- Ixodes dentatus <small>Marx, 1899</small>
- Ixodes dicei <small>Keirans and Ajohda, 2003</small>
- Ixodes diomedeae <small>Arthur, 1958</small>
- Ixodes diversifossus <small>Neumann, 1899</small>
- Ixodes djaronensis <small>Neumann, 1907</small>
- Ixodes domerguei <small>Uilenberg and Hoogstraal, 1965</small>
- Ixodes downsi <small>Kohls, 1957</small>
- Ixodes drakensbergensis <small>Clifford, Theiler and Baker, 1975b</small>
- Ixodes eadsi <small>Kohls and Clifford, 1964</small>
- Ixodes eastoni <small>Keirans and Clifford, 1983</small>
- Ixodes eichhorni <small>Nuttall, 1916</small>
- Ixodes eldaricus <small>Dzhaparidze, 1950a</small>
- Ixodes elongatus <small>Bedford, 1929</small>
- Ixodes eudyptidis <small>Maskell, 1885</small>
- Ixodes euplecti <small>Arthur, 1958</small>
- Ixodes evansi <small>Arthur, 1956</small>
- Ixodes (Exopalpiger) fecialis <small>Warburton and Nuttall, 1909</small>
- Ixodes (Ixodes) festai <small>Arthur, 1961</small>
- Ixodes filippovae <small>ÃÂerný, 1961</small>
- Ixodes fossulatus <small>Neumann, 1899</small>
- Ixodes (Trichotoixodes) frontalis <small>(Panzer, 1798)</small>
- Ixodes fuscipes <small>Koch, 1844a</small>
- Ixodes fynbosensis <small>Apanaskevich, Horak, Matthee and Matthee, 2011</small>
- Ixodes galapagoensis <small>Clifford and Hoogstraal, 1980</small>
- Ixodes (Filippoviella) ghilarovi <small>Filippova and Panova, 1988b</small>
- Ixodes gibbosus <small>Nuttall, 1916</small>
- Ixodes giluwensis <small>Apanaskevich & Schenk, 2020</small>
- Ixodes goliath <small>Apanaskevich and Lemon, 2018</small>
- Ixodes granulatus <small>Supino, 1897</small>
- Ixodes gregsoni <small>Lindquist, Wu and Redner, 1999</small>
- Ixodes guatemalensis <small>Kohls, 1956</small>
- Ixodes hearlei <small>Gregson, 1941</small>
- Ixodes heathi <small>Kwak, 2018</small>
- Ixodes heinrichi <small>Arthur, 1962</small>
- Ixodes (Pholeoixodes) hexagonus <small>Leach, 1815</small>
- Ixodes himalayensis <small>Dhanda and Kulkarni, 1969</small>
- Ixodes (Sternalixodes) hirsti <small>Hassall, 1931</small>
- Ixodes (Sternalixodes) holocyclus <small>Neumann, 1899</small>
- Ixodes hoogstraali <small>Arthur, 1955</small>
- Ixodes howelli <small>Cooley and Kohls, 1938</small>
- Ixodes hyatti <small>Clifford, Hoogstraal and Kohls, 1971</small>
- Ixodes hydromyidis <small>Swan, 1931</small>
- Ixodes (Pholeoixodes) hunanensis <small>Apanaskevich &</small> <small>Duan, 2022</small>
- Ixodes (Afrixodes) hyracis <small>Apanaskevich, Drew & Pienaar, 2025</small>
- Ixodes inopinatus <small>Estrada-Peña, Nava and Petney, 2014</small>
- Ixodes jacksoni <small>Hoogstraal, 1967</small>
- Ixodes jellisoni <small>Cooley and Kohls, 1938</small>
- Ixodes jonesae <small>Kohls, Sonenshine and Clifford, 1969</small>
- Ixodes kaiseri <small>Arthur, 1957</small>
- Ixodes kandingensis <small>Guo, Sun, Xu and Durden, 2017</small>
- Ixodes kashmiricus <small>Pomerantzev, 1948</small>
- Ixodes kazakstani <small>Olenev and Sorokoumov, 1934</small>
- Ixodes kerguelenensis <small>André and Colas-Belcour, 1942</small>
- Ixodes kingi <small>Bishopp, 1911</small>
- Ixodes (Scaphixodes) kohlsi <small>Arthur, 1955b</small>
- Ixodes kopsteini <small>(Oudemans, 1926)</small>
- Ixodes kuntzi <small>Hoogstraal and Kohls, 1965a</small>
- Ixodes laguri <small>Olenev, 1929</small>
- Ixodes (Pholeoixodes) lanigeri <small>Hornok, 2024</small>
- Ixodes laridis <small>Heath and Palma, 2017</small>
- Ixodes lasallei <small>Méndez Arocha and Ortiz, 1958</small>
- Ixodes latus <small>Arthur, 1958</small>
- Ixodes laysanensis <small>Wilson, 1964a</small>
- Ixodes lemuris <small>Arthur, 1958</small>
- Ixodes lewisi <small>Arthur, 1965</small>
- Ixodes lividus <small>Koch, 1844</small>
- Ixodes (Haemixodes) longiscutatus <small>Boero, 1944</small>
- Ixodes (Amerixodes) loricatus <small>Neumann, 1899</small>
- Ixodes loveridgei <small>Arthur, 1958</small>
- Ixodes luciae <small>Sénevet, 1940</small>
- Ixodes lunatus <small>Neumann, 1907</small>
- Ixodes luxuriosus <small>Schulze, 1935</small>
- Ixodes macfarlanei <small>Keirans, Clifford and Walker, 1982</small>
- Ixodes malayensis <small>Kohls, 1962</small>
- Ixodes marmotae <small>Cooley and Kohls, 1938</small>
- Ixodes marxi <small>Banks, 1908</small>
- Ixodes maslovi <small>Emel'yanova and Kozlovskaya, 1967</small>
- Ixodes matopi <small>Spickett, Keirans, Norval and Clifford, 1981</small>
- Ixodes mexicanus <small>Cooley and Kohls, 1942</small>
- Ixodes microgalei <small>Apanaskevich, Soarimalala and Goodman, 2013</small>
- Ixodes minor <small>Neumann, 1902</small>
- Ixodes minutae <small>Arthur, 1959</small>
- Ixodes mitchelli <small>Kohls, Clifford and Hoogstraal, 1970</small>
- Ixodes mirzai <small>Apanaskevich & Schenk, 2020</small>
- Ixodes (Ixodes) mojavensis <small>Backus & Beati</small><small>, 2022</small>
- Ixodes monospinosus <small>Saito, 1967</small>
- Ixodes montoyanus <small>Cooley, 1944</small>
- Ixodes moreli <small>Arthur, 1957</small>
- Ixodes moscharius <small>Teng, 1982</small>
- Ixodes moschiferi <small>Nemenz, 1968</small>
- Ixodes muniensis <small>Arthur and Burrow, 1957</small>
- Ixodes muris <small>Bishopp and Smith, 1937</small>
- Ixodes murreleti <small>Cooley and Kohls, 1945</small>
- Ixodes myospalacis <small>Teng, 1986</small>
- Ixodes myotomys <small>Clifford and Hoogstraal, 1970</small>
- Ixodes (Sternalixodes) myremecobii <small>Roberts, 1962</small>
- Ixodes nairobiensis <small>Nuttall, 1916</small>
- Ixodes nchisiensis <small>Arthur, 1958</small>
- Ixodes nectomys <small>Kohls, 1956a</small>
- Ixodes neitzi <small>Clifford, Walker and Keirans, 1977</small>
- Ixodes nesomys <small>Uilenberg and Hoogstraal, 1969</small>
- Ixodes neuquenensis <small>Ringuelet, 1947</small>
- Ixodes nicolasi <small>Santos Dias, 1982</small>
- Ixodes nipponensis <small>Kitaoka and Saito, 1967</small>
- Ixodes nitens <small>Neumann, 1904</small>
- Ixodes nuttalli <small>Lahille, 1913</small>
- Ixodes nuttallianus <small>Schulze, 1930</small>
- Ixodes occultus <small>Pomerantzev, 1946</small>
- Ixodes ochotonae <small>Gregson, 1941</small>
- Ixodes okapiae <small>Arthur, 1956</small>
- Ixodes oldi <small>Nuttall, 1913</small>
- Ixodes (Coxixodes) ornithorhynchi <small>Lucas, 1846</small>
- Ixodes (Partipalpiger) ovatus <small>Neumann, 1899</small>
- Ixodes pacificus <small>Cooley and Kohls, 1943</small>
- Ixodes (Ixodes) paragibbosus <small>Hornok and Kontschán, 2025</small>
- Ixodes paranaensis <small>Barros-Battesti, Arzua, Pichorim and Keirans, 2003</small>
- Ixodes pararicinus <small>Keirans and Clifford, 1985</small>
- Ixodes pavlovskyi <small>Pomerantzev, 1946</small>
- Ixodes percavatus <small>Neumann, 1906</small>
- Ixodes peromysci <small>Auguston, 1940</small>
- Ixodes persulcatus <small>Schulze, 1930</small>
- Ixodes petauristae <small>Warburton, 1933</small>
- Ixodes (Scaphixodes) philipi <small>Keirans and Kohls, 1970</small>
- Ixodes pilosus <small>Koch, 1844</small>
- Ixodes planiscutatus <small>Apanaskevich & Schenk, 2020</small>
- Ixodes pomerantzevi <small>Serdjukova, 1941</small>
- Ixodes pomerantzi <small>Kohls, 1956</small>
- Ixodes priscicollaris <small>Schulze, 1932</small>
- Ixodes procaviae <small>Arthur and Burrow, 1957</small>
- Ixodes prokopjevi <small>(Emel'yanova, 1979)</small>
- Ixodes radfordi <small>Kohls, 1948</small>
- Ixodes rageaui <small>Arthur, 1958</small>
- Ixodes randrianasoloi <small>Uilenberg and Hoogstraal, 1969b</small>
- Ixodes rasus <small>Neumann, 1899</small>
- Ixodes redikorzevi <small>Olenev, 1927</small>
- Ixodes rhabdomysae <small>Arthur, 1959</small>
- Ixodes ricinus <small>(Linnaeus, 1758)</small>
- Ixodes rothschildi <small>Nuttall and Warburton, 1911</small>
- Ixodes rotundatus <small>Arthur, 1958</small>
- Ixodes (Afrixodes) rubicundus <small>Neumann, 1904</small>
- Ixodes rubidus <small>Neumann, 1901</small>
- Ixodes rugicollis <small>Schulze and Schlottke, 1929b</small>
- Ixodes rugosus <small>Bishopp, 1911</small>
- Ixodes sachalinensis <small>Filippova, 1971</small>
- Ixodes scapularis <small>Say, 1821</small>
- Ixodes schillingsi <small>Neumann, 1901</small>
- Ixodes schulzei <small>Aragão and Fonseca, 1951</small>
- Ixodes sculptus <small>Neumann, 1904</small>
- Ixodes semenovi <small>Olenev, 1929</small>
- Ixodes shahi <small>Clifford, Hoogstraal and Kohls, 1971</small>
- Ixodes siamensis <small>Kitaoka and Suzuki, 1983</small>
- Ixodes sigelos <small>Keirans, Clifford and Corwin, 1976</small>
- Ixodes signatus <small>Birula, 1895</small>
- Ixodes (Trichotoixodes) silvanus <small>Saracho-Bottero, Beati, Venzal, Guglielmone</small> <small>& Nava, 2021</small>
- Ixodes (Pomerantzevella) simplex <small>Neumann, 1906</small>
- Ixodes sinaloa <small>Kohls and Clifford, 1966</small>
- Ixodes sinensis <small>Teng, 1977</small>
- Ixodes (Afrixodes) soarimalalae <small>Apanaskevich & Goodman, 2020</small>
- Ixodes soricis <small>Gregson, 1942</small>
- Ixodes spinae <small>Arthur, 1958</small>
- Ixodes spinicoxalis <small>Neumann, 1899</small>
- Ixodes spinipalpis <small>Hadwen and Nuttall, 1916</small>
- Ixodes spinosus <small>Neumann, 1899</small>
- Ixodes steini <small>Schulze, 1935</small>
- Ixodes stellae <small>Apanaskevich & Schenk, 2020</small>
- Ixodes stilesi <small>Neumann, 1911</small>
- Ixodes stromi <small>Filippova, 1957</small>
- Ixodes subterraneus <small>Filippova, 1961</small>
- Ixodes succineus <small>Weidner, 1964</small>
- Ixodes taglei <small>Kohls, 1969</small>
- Ixodes tamaulipas <small>Kohls and Clifford, 1966</small>
- Ixodes tancitarius <small>Cooley and Kohls, 1942</small>
- Ixodes tanuki <small>Saito, 1964</small>
- Ixodes tapirus <small>Kohls, 1956</small>
- Ixodes (Ixodes) tatei <small>Arthur, 1959</small>
- Ixodes (Endopalpiger) tasmani <small>Neumann, 1899</small>
- Ixodes tecpanensis <small>Kohls, 1956</small>
- Ixodes texanus <small>Banks, 1909</small>
- Ixodes (Trichotoixodes) theilerae <small>Arthur, 1953</small>
- Ixodes thomasae <small>Arthur and Burrow, 1957</small>
- Ixodes tiptoni <small>Kohls and Clifford, 1962</small>
- Ixodes tovari <small>Cooley, 1945</small>
- Ixodes transvaalensis <small>Clifford and Hoogstraal, 1966</small>
- Ixodes (Filippoviella) trianguliceps <small>Birula, 1895</small>
- Ixodes (Sternalixodes) trichosuri <small>Roberts, 1960</small>
- Ixodes tropicalis <small>Kohls, 1956a</small>
- Ixodes (Trichotoixodes) turdus <small>Nakatsudi, 1942</small>
- Ixodes ugandanus <small>Neumann, 1906</small>
- Ixodes (Afrixodes) uilenbergi <small>Apanaskevich & Goodman, 2020</small>
- Ixodes (Afrixodes) uncus <small>Apanaskevich & Goodman, 2020</small>
- Ixodes unicavatus <small>Neumann, 1908</small>
- Ixodes (Ceratixodes) uriae <small>White, 1852</small>
- Ixodes vanidicus <small>Schulze, 1943</small>
- Ixodes venezuelensis <small>Kohls, 1953</small>
- Ixodes ventalloi <small>Gill Collado, 1936</small>
- Ixodes (Eschatocephalus) vespertilionis <small>Koch, 1844</small>
- Ixodes vestitus <small>Neumann, 1908</small>
- Ixodes victoriensis <small>Nuttall, 1916</small>
- Ixodes walkerae <small>Clifford, Kohls and Hoogstraal, 1968</small>
- Ixodes werneri <small>Kohls, 1950</small>
- Ixodes woodi <small>Bishopp, 1911</small>
- Ixodes (Endopalpiger) woyliei <small>Ash, Elliot, Godfrey, Burmej, Abdad, Northover, Wayne, Morris, Clode, Lymbery and Thompson, 2017</small>
- Ixodes zaglossi <small>Kohls, 1960</small>
- Ixodes (Ixodes) zacateco <small>Apanaskevich & Bunn, 2025</small>
- Ixodes zairensis <small>Keirans, Clifford and Walker, 1982</small>
Fossil Species
Three fossil species of Ixodes are known from amber deposits ranging from the Cretaceous to the Eocene:
- â Ixodes antiquorum <small>Chitimia-Dobler, Mans and Dunlop, 2022</small> (Burmese amber, Cenomanian, ~ 99 Ma)
- â Ixodes succineus <small>Weidner, 1964</small> (Baltic amber, Eocene, ~ 49âÂÂ44 Ma)
- â Ixodes tertiarius <small>Scudder, 1885</small> (Green River Formation, Eocene, 50 Ma)
Notes
Additional Readings
Refer to the following external links for recent changes to Ixodes taxonomy:
- Guglielmone, A. A., Robbins, R. G., Apanaskevich, D. A., Petney, T. N., Estrada-Peña, A., & Horak, I. G. (2014a). The Hard Ticks of the World: (Acari: Ixodida: Ixodidae). Springer Netherlands. https://doi.org/10.1007/978-94-007-7497-1
- Guglielmone, A.A., Petney, T.N. & Robbins, R.G. (2020) Ixodidae (Acari: Ixodoidea): descriptions and redescriptions of all known species from 1758 to December 31, 2019. Zootaxa, 4871 (1), 1âÂÂ322. https://doi.org/10.11646/zootaxa.4871.1.1
References
External links