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Hypselostomatidae

Hypselostomatidae is a family of small, air-breathing land snails, terrestrial pulmonate gastropod molluscs in the superfamily Pupilloidea.

General characteristics

The shell of a typical hypselostomatid ranges from conical or conical-ovoid to depressed-conical. However, the body whorl frequently becomes detached from the penultimate whorl (Hoekstra and Schilthuizen 2011; Chen et al. 2022). This separation—known as a solute or uncoiled condition—drastically alters the overall shell profile, often resulting in a trumpet-like or distorted appearance.

Distribution

Distributed mainly in Southeastern Asia,but also in India to Pakistan, China, Japan and Australia; mostly on limestone habitats.

Taxonomy

The Hypselostomatidae were originally established as a subfamily of Chondrinidae (Zilch 1959). In that same work, Zilch erected the subfamily Aulacospirinae, which was later synonymized by Schileyko (1998). Systematic placement has since remained a subject of debate: while Bouchet et al. (2017) treated the group as a subfamily of Gastrocoptidae, other authors have classified these representatives under Vertiginidae (Panha 1998; Tanmuangpak and Dumrongrojwattana 2022) or Pupillidae (Burch and Panha 2002). More recently, several authors have advocated for full family status as the Hypselostomatidae (Gojšina et al. 2022; Páll-Gergely 2023a, b).

Genera

  • Acinolaemus <small>Thompson & Upatham, 1997</small>
  • Anauchen <small>Pilsbry, 1917</small>
  • Angustopila <small>Jochum, Slapnik & Páll-Gergely, 2014</small>
  • Antroapiculus <small>Panha & Burch, 1999</small>: synonym of Antroapiculus <small>Panha & J. B. Burch, 2002</small> accepted as Hypselostoma <small>W. H. Benson, 1856</small> (incorrect subsequent spelling)
  • Aulacospira <small>Möllendorff, 1890</small>
  • Bensonella <small>Pilsbry & Vanatta, 1900</small>
  • Boysia <small>L. Pfeiffer, 1849</small>
  • Boysidia <small>Ancey, 1881</small>
  • Clostophis <small>Benson, 1860</small>
  • Dentisphaera <small>Páll-Gergely & Jochum, 2017</small> - endemic to the caves in Northern Vietnam
  • Gyliotrachela <small>Tomlin, 1930</small>: synonym of Hypselostoma <small>W. H. Benson, 1856</small> (junior subjective synonym)
  • Hypselostoma <small>Benson, 1856</small> - the type genus of the family
  • Krobylos <small>Panha & Burch, 1999</small>
  • Montapiculus <small>Panha & Burch, 1999</small>: synonym of Clostophis <small>W. H. Benson, 1860</small> (junior synonym)
  • Pseudostreptaxis <small>Möllendorff, 1890</small>
  • Tonkinospira <small>Jochum, Slapnik & Páll-Gergely, 2014</small>
  • Paraboysidia <small>Pilsbry, 1917</small>: synonym of Bensonella <small>Pilsbry & Vanatta, 1900</small> (junior subjective synonym)

References

  • Zilch, A. (1959-1960). Gastropoda. Teil 2. Euthyneura. In: Schindewolf, O. H. (Ed.), Handbuch der Paläozoologie, Band 6. Borntraeger, Berlin. Lfg. 1: pp. 1-200 (17 July 1959); Lfg. 2: pp. 201-400 (25 November 1959); Lfg. 3: pp. 401-600
  • Schileyko, A. A. (1998). Treatise on Recent terrestrial pulmonate molluscs. Part 2. Gastrocoptidae, Hypselostomatidae, Vertiginidae, Truncatellinidae, Pachnodidae, Enidae, Sagdidae. Ruthenica. Supplement 2: 129-261. Moskva
  • Panha S (1998a) [“1997”] Three new species of microsnails from southern Thailand (Pulmonata: Vertiginidae; Prosobranchia: Diplommatinidae). Malacological Review 30(1): 53–59.
  • Tanmuangpak K, Dumrongrojwattana P (2022) Gyliotrachela cultura, a new species of terrestrial microsnail (Gastropoda: Eupulmonata: Vertiginidae) from Thailand. Raffles Bulletin of Zoology, 70: 407–416.
  • Burch JB, Panha S (2002) [“2000”] The pupillid genus Anauchen in Thailand (Pulmonata: Stylommatophora). Walkerana 11(26): 239–248.

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