Holothyrida

The Holothyrida are a small order of mites in the superorder Parasitiformes. No fossils are known. With body lengths of more than they are relatively large mites, with a heavily sclerotized body. It is divided into three families, Allothyridae, Holothyridae, and Neothyridae. In a 1998 experimental study, members of the family Allothyridae were found to ignore living animals but readily fed on the body fluids of dead arthropods, making them scavengers.

The order has a distribution largely confined to former Gondwanan landmasses. They are the sister group to Ixodida (ticks).

Systematics

Allothyridae

Allothyridae <small>van der Hammen, 1972</small> — Australia, New Zealand

• Allothyrus <small>van der Hammen, 1961</small>

* Allothyrus australasiae <small>(Womersley, 1935)</small>

* Allothyrus constrictus <small>(Domrow, 1955)</small>

• Australothyrus <small>van der Hammen, 1983</small>

* Australothyrus ocellatus <small>van der Hammen, 1983</small>

Holothyridae

Holothyridae <small>Thorell, 1882</small> Sri Lanka, Indian Ocean islands, New Guinea, New Caledonia

• Sternothyrus <small>Lehtinen, 1995</small>
• Sternothyrus braueri <small>(Thon, 1905)</small> — Seychelles

• Lindothyrus <small>Lehtinen, 1995</small>
• Lindothyrus elongatus <small>Lehtinen, 1995</small> — Lord Howe Island
• Lindothyrus rubellus <small>Lehtinen, 1995</small> — New Caledonia

• Indothyrus <small>Lehtinen, 1995</small>
• Indothyrus greeni <small>Lehtinen, 1995</small> — Sri Lanka

• Haplothyrus <small>Lehtinen, 1995</small>
• Haplothyrus expolitissimus <small>(Berlese, 1924)</small> — New Caledonia
• Haplothyrus hyatti <small>Lehtinen, 1995</small> — unknown locality

• Holothyrus <small>Gervais, 1842</small> — Mauritius
• Holothyrus coccinella <small>Gervais, 1842</small>
• Holothyrus legendrei <small>Hammen, 1983</small>

• Hammenius <small>Lehtinen, 1981</small>
• Hammenius armatus <small>(Canestrini, 1897)</small> — Tamara Island (Aitape): New Guinea
• Hammenius berlesei <small>(Lehtinen, 1995)</small> — New Guinea
• Hammenius braueri <small>(Thon, 1906)</small>
• Hammenius fujuge <small>Lehtinen, 1981</small> — New Guinea (Central District, Oro Province)
• Hammenius grandjeani <small>(Hammen, 1961)</small> — Mount Bosavi: New Guinea
• Hammenius holthuisi <small>van der Hammen, 1983</small>
• Hammenius ingii <small>Lehtinen, 1981</small> — New Guinea
• Hammenius insularis <small>Lehtinen, 1995</small> — Louisiade Archipelago: New Guinea
• Hammenius longipes <small>(Thorell, 1882)</small> — Fly River, New Guinea (?)
• Hammenius mendi <small>(Lehtinen, 1995)</small> — Strickland River: New Guinea
• Hammenius montanus <small>Hammen, 1983</small> — Irian Jaya
• Hammenius niger <small>(Thon, 1906)</small>

Neothyridae

Neothyridae <small>Lehtinen, 1981</small> Northern South America and the Caribbean

• Diplothyrus <small>Lehtinen, 1999</small> Brazil, French Guiana
• Diplothyrus schubarti <small>Lehtinen, 1999</small>
• Diplothyrus lecorrei <small>Klompen 2010</small>
• Diplothyrus lehtineni <small>Vázquez & de Araújo & Feres 2016</small>
• Neothyrus <small>Lehtinen, 1981</small> Peru
• Neothyrus ana <small>Lehtinen, 1981</small>
• Caribothyrus <small>Kontschán & Mahunka 2004</small> Dominican Republic
• Caribothyrus barbatus <small>Kontschán & Mahunka 2004</small>

Footnotes

References

• Joel Hallan's Biology Catalog: Holothyrida
• Bruce Halliday: Order Holothyrida
• Lehtinen, Pekka T. (1995): Revision of the old world Holothyridae (Arachnida : Anactinotrichida : Holothyrina). Invertebrate Taxonomy 9(4): 767-826.