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Helius (fly)

Helius is a genus of crane fly in the family Limoniidae, distributed worldwide with most species in Australia and East Asia.

Species

*H. abnormalis <small>(Brunetti, 1918)</small>
*H. ata <small>Alexander, 1931</small>
*H. atroapicalis <small>Alexander, 1978</small>
*H. auranticolor <small>Alexander, 1936</small>
*H. connectus <small>Edwards, 1928</small>
*H. diacanthus <small>Alexander, 1931</small>
*H. fuscofemoratus <small>Alexander, 1931</small>
*H. glabristylatus <small>Alexander, 1931</small>
*H. indivisus <small>Alexander, 1931</small>
*H. inelegans <small>(Alexander, 1928)</small>
*H. invenustipes <small>(Alexander, 1930)</small>
*H. melanosoma <small>Alexander, 1936</small>
*H. mimicans <small>Edwards, 1933</small>
*H. mirus <small>Edwards, 1926</small>
*H. monticola <small>(Edwards, 1919)</small>
*H. mouensis <small>Alexander, 1948</small>
*H. nigrofemoratus <small>(Alexander, 1930)</small>
*H. niveitarsis <small>(Skuse, 1890)</small>
*H. pallens <small>Edwards, 1933</small>
*H. perelegans <small>Alexander, 1930</small>
*H. perlongatus <small>Alexander, 1978</small>
*H. scabiosus <small>Alexander, 1932</small>
*H. vitiensis <small>Alexander, 1956</small>
  • Subgenus Helius <small>Lepeletier & Serville, 1828</small>
*H. acanthostyla <small>Alexander, 1944</small>
*H. aciferus <small>Alexander, 1937</small>
*H. aka <small>Alexander, 1972</small>
*H. albitarsis <small>(Osten Sacken, 1888)</small>
*H. albogeniculatus <small>Alexander, 1936</small>
*H. amplus <small>Edwards, 1933</small>
*H. anaemicus <small>Alexander, 1932</small>
*H. anamalaiensis <small>Alexander, 1967</small>
*H. angustalbus <small>Alexander, 1953</small>
*H. aphrophilus <small>Alexander, 1948</small>
*H. apicalis <small>(Alexander, 1915)</small>
*H. apoensis <small>Alexander, 1931</small>
*H. apophysalis <small>Alexander, 1967</small>
*H. araucariae <small>Alexander, 1945</small>
*H. arcuarius <small>Alexander, 1929</small>
*H. argyrosternus <small>Alexander, 1930</small>
*H. arunachalus <small>Alexander, 1975</small>
*H. attenuatus <small>Alexander, 1929</small>
*H. barbatus <small>Edwards, 1921</small>
*H. bicolor <small>Edwards, 1933</small>
*H. bifurcus <small>Alexander, 1956</small>
*H. bitergatus <small>Alexander, 1950</small>
*H. boops <small>Alexander, 1942</small>
*H. brachyphallus <small>Alexander, 1956</small>
*H. brevioricornis <small>(Alexander, 1920)</small>
*H. brevisector <small>Alexander, 1956</small>
*H. brunneus <small>Byers, 1981</small>
*H. cacoxenus <small>(Alexander, 1920)</small>
*H. calviensis <small>Edwards, 1928</small>
*H. capensis <small>(Alexander, 1917)</small>
*H. capniopterus <small>Alexander, 1945</small>
*H. catreus <small>Alexander, 1967</small>
*H. cavernicolus <small>Alexander, 1961</small>
*H. chikurinensis <small>Alexander, 1930</small>
*H. chintoo <small>Theischinger, 1994</small>
*H. communis <small>(Skuse, 1890)</small>
*H. comoreanus <small>Alexander, 1959</small>
*H. compactus <small>Alexander, 1967</small>
*H. copiosus <small>Alexander, 1935</small>
*H. corniger <small>Savchenko, 1983</small>
*H. costofimbriatus <small>Alexander, 1930</small>
*H. costosetosus <small>Alexander, 1932</small>
*H. creper <small>Alexander, 1926</small>
*H. ctenonycha <small>Alexander, 1938</small>
*H. dafla <small>Alexander, 1972</small>
*H. darlingtonae <small>Welch & Gelhaus, 1994</small>
*H. destitutus <small>Alexander, 1971</small>
*H. devinctus <small>Alexander, 1932</small>
*H. distinervis <small>Alexander, 1940</small>
*H. dolichorhynchus <small>Edwards, 1933</small>
*H. eremnophallus <small>Alexander, 1978</small>
*H. euryphallus <small>Alexander, 1960</small>
*H. fasciventris <small>Edwards, 1926</small>
*H. ferruginosus <small>(Brunetti, 1912)</small>
*H. flavidibasis <small>Alexander, 1975</small>
*H. flavipes <small>(Macquart, 1855)</small>
*H. flavitarsis <small>(Alexander, 1920)</small>
*H. flavus <small>(Walker, 1856)</small>
*H. fragosus <small>Alexander, 1937</small>
*H. franckianus <small>Alexander, 1940</small>
*H. fratellus <small>(Brunetti, 1918)</small>
*H. fulvithorax <small>(Skuse, 1890)</small>
*H. fumicosta <small>Edwards, 1928</small>
*H. fuscoangustus <small>Alexander, 1967</small>
*H. garcianus <small>Alexander, 1972</small>
*H. gibbifer <small>Savchenko, 1981</small>
*H. gorokanus <small>Alexander, 1962</small>
*H. gracillimus <small>Alexander, 1938</small>
*H. graphipterus <small>Alexander, 1954</small>
*H. haemorrhoidalis <small>Alexander, 1937</small>
*H. harrisi <small>(Alexander, 1923)</small>
*H. hatschbachi <small>Alexander, 1954</small>
*H. hispanicus <small>Lackschewitz, 1928</small>
*H. hova <small>Alexander, 1955</small>
*H. impensus <small>Alexander, 1967</small>
*H. imperfectus <small>(Alexander, 1920)</small>
*H. inconspicuus <small>(Brunetti, 1912)</small>
*H. ineptus <small>Alexander, 1938</small>
*H. infirmus <small>Alexander, 1932</small>
*H. invariegatus <small>Alexander, 1938</small>
*H. iris <small>(Alexander, 1920)</small>
*H. kambangani <small>(de Meijere, 1913)</small>
*H. kambanganoides <small>Alexander, 1931</small>
*H. larotypa <small>Alexander, 1926</small>
*H. lectus <small>Alexander, 1936</small>
*H. leucoplaca <small>Alexander, 1936</small>
*H. lienpingensis <small>Alexander, 1945</small>
*H. liguliferus <small>Alexander, 1978</small>
*H. liliputanus <small>Alexander, 1929</small>
*H. lobuliferus <small>Alexander, 1938</small>
*H. longinervis <small>Edwards, 1928</small>
*H. longirostris <small>(Meigen, 1818)</small>
*H. luachimoensis <small>Alexander, 1963</small>
*H. luniger <small>(Riedel, 1914)</small>
*H. mainensis <small>(Alexander, 1916)</small>
*H. malagasicus <small>Alexander, 1955</small>
*H. manueli <small>Alexander, 1972</small>
*H. medleri <small>Alexander, 1976</small>
*H. melanophallus <small>Alexander, 1938</small>
*H. mesorhynchus <small>Alexander, 1933</small>
*H. micracanthus <small>Alexander, 1945</small>
*H. mirabilis <small>(Alexander, 1914)</small>
*H. mirandus <small>(Alexander, 1921)</small>
*H. morosus <small>(Alexander, 1920)</small>
*H. multivolus <small>Alexander, 1945</small>
*H. murreensis <small>Alexander, 1960</small>
*H. myersiellus <small>Alexander, 1935</small>
*H. nawaianus <small>Alexander, 1929</small>
*H. neocaledonicus <small>Alexander, 1945</small>
*H. nigricapella <small>Alexander, 1938</small>
*H. nigriceps <small>(Edwards, 1916)</small>
*H. nipponensis <small>(Alexander, 1913)</small>
*H. numenius <small>Alexander, 1967</small>
*H. obliteratus <small>(Alexander, 1920)</small>
*H. obsoletus <small>(Alexander, 1920)</small>
*H. oxystylus <small>Alexander, 1967</small>
*H. pallidipes <small>Alexander, 1926</small>
*H. pallidissimus <small>Alexander, 1930</small>
*H. pallirostris <small>Edwards, 1921</small>
*H. panamensis <small>Alexander, 1934</small>
*H. paramorosus <small>Alexander, 1949</small>
*H. parvidens <small>Alexander, 1944</small>
*H. patens <small>Edwards, 1933</small>
*H. pauperculus <small>(Alexander, 1921)</small>
*H. pavoninus <small>Alexander, 1938</small>
*H. perflavens <small>Alexander, 1964</small>
*H. perpallidus <small>Alexander, 1942</small>
*H. pervenustus <small>Alexander, 1953</small>
*H. phasmatis <small>Alexander, 1945</small>
*H. pictus <small>Edwards, 1926</small>
*H. plebeius <small>Alexander, 1946</small>
*H. pluto <small>Alexander, 1932</small>
*H. polionotus <small>Alexander, 1938</small>
*H. procerus <small>Alexander, 1931</small>
*H. productellus <small>Alexander, 1944</small>
*H. protumidus <small>Alexander, 1978</small>
*H. quadrifidus <small>Alexander, 1926</small>
*H. quadrivena <small>Alexander, 1971</small>
*H. rectispinus <small>Alexander, 1947</small>
*H. rectus <small>Alexander, 1945</small>
*H. regius <small>Alexander, 1944</small>
*H. rostratus <small>Alexander, 1956</small>
*H. rubicundus <small>(Alexander, 1922)</small>
*H. rufescens <small>(Edwards, 1916)</small>
*H. rufithorax <small>Alexander, 1928</small>
*H. sanguinolentus <small>(Alexander, 1921)</small>
*H. schildi <small>Alexander, 1945</small>
*H. selectus <small>Alexander, 1936</small>
*H. serenus <small>Alexander, 1967</small>
*H. setigerus <small>Alexander, 1980</small>
*H. sigillatus <small>Alexander, 1964</small>
*H. stenorhynchus <small>Alexander, 1954</small>
*H. stolidus <small>Alexander, 1948</small>
*H. subanaemicus <small>Alexander, 1964</small>
*H. subarcuarius <small>Alexander, 1934</small>
*H. subfasciatus <small>Alexander, 1924</small>
*H. submorosus <small>(Alexander, 1921)</small>
*H. subobsoletus <small>(Alexander, 1921)</small>
*H. subpauperculus <small>Alexander, 1975</small>
*H. tantalus <small>Alexander, 1967</small>
*H. tanyrhinus <small>Alexander, 1964</small>
*H. taos <small>Alexander, 1967</small>
*H. tenuirostris <small>Alexander, 1924</small>
*H. tenuistylus <small>Alexander, 1929</small>
*H. tetracradus <small>Alexander, 1967</small>
*H. tienmuanus <small>Alexander, 1940</small>
*H. tonaleah <small>Theischinger, 1994</small>
*H. trianguliferus <small>Alexander, 1931</small>
*H. tshinta <small>Theischinger, 1994</small>
*H. unicolor <small>(Brunetti, 1912)</small>
*H. uniformis <small>(Alexander, 1914)</small>
*H. venustus <small>(Skuse, 1890)</small>
*H. verticillatus <small>Alexander, 1967</small>
*H. yindi <small>Theischinger, 1994</small>
*H. melanolithus <small>Alexander, 1953</small>
*H. mirificus <small>Alexander, 1953</small>
*H. pentaneura <small>Alexander, 1948</small>
*H. allunga <small>Theischinger, 1994</small>
*H. dugaldi <small>Alexander, 1945</small>
*H. edwardsianus <small>Alexander, 1956</small>
*H. fulani <small>Alexander, 1975</small>
*H. camerounensis <small>(Alexander, 1920)</small>
*H. alluaudi <small>(Riedel, 1914)</small>
*H. venustissimus <small>(Alexander, 1920)</small>
*H. bigeminatus <small>Alexander, 1962</small>
*H. brevinasus <small>Alexander, 1934</small>
*H. deentoo <small>Theischinger, 1994</small>
*H. diffusus <small>Alexander, 1941</small>
*H. fenestratus <small>Alexander, 1973</small>
*H. gracilirostris <small>Alexander, 1962</small>
*H. guttulinus <small>Alexander, 1960</small>
*H. japenensis <small>Alexander, 1948</small>
*H. matilei <small>Hynes, 1993</small>
*H. mesolineatus <small>Alexander, 1962</small>
*H. nimbus <small>Alexander, 1941</small>
*H. papuanus <small>Alexander, 1934</small>
*H. reticularis <small>(Alexander, 1915)</small>
*H. simulator <small>Alexander, 1962</small>
*H. spinteris <small>Alexander, 1960</small>
*H. subreticulatus <small>Alexander, 1962</small>
*H. dicroneurus <small>Alexander, 1964</small>

References