Heliocopris (large dung beetles) is a genus of Scarabaeidae or scarab beetles in the superfamily Scarabaeoidea. Forty-seven of the fifty-two known species are found in Africa, but a few are found in southern and southeast Asia.
Heliocopris includes some of the world's largest dung beetles (their size only rivalled by certain Catharsius and Coprophanaeus), with H. anderson, H. colossus, H. dilloni, H. dominus, H. gigas and H. tyrannus reaching up to about long, although a more typical size for the members of this genus is . They are generally black or dark brown beetles where males tend to have horns on their head or pronotum, which females lack. Some are associated with the dung of elephants, rhinos or hippos, but there are large variations among the species in the genus and dung from other animals, like domestic cattle or monkeys, is also used. The female lays her eggs in a tunnel she digs under the excrement.
Heliocopris has been placed in the tribe Dichotomiini, but Montreuil places them in Coprini.
Species
The genus Heliocopris includes the following species:
- Heliocopris alatus <small>Felsche, 1910</small>
- Heliocopris anadematus <small>Gillet, 1908</small>
- Heliocopris andersoni <small>Bates, 1868</small>
- Heliocopris anguliceps <small>Janssens, 1943</small>
- Heliocopris antenor <small>(Olivier, 1789)</small>
- Heliocopris ares <small>Philippe, Minetti, 2022</small>
- Heliocopris atropos <small>Boheman, 1860</small>
- Heliocopris beccarii <small>Harold, 1871</small>
- Heliocopris biimpressus <small>Kolbe, 1893</small>
- Heliocopris bucephalus <small>(Fabricius, 1775)</small>
- Heliocopris camerunus <small>Pokorny & Zidek, 2009</small>
- Heliocopris colossus <small>Bates, 1868</small>
- Heliocopris corniculatus <small>Janssens, 1939</small>
- Heliocopris coronatus <small>Felsche, 1901</small>
- Heliocopris cuneifer <small>Lesne, 1906</small>
- Heliocopris densissa <small>Roth, 1851</small>
- Heliocopris dianae <small>Hope, 1842</small>
- Heliocopris dilloni <small>Guérin-Méneville, 1847</small>
- Heliocopris dolosus <small>Janssens, 1939</small>
- Heliocopris dominus <small>Bates, 1868</small>
- Heliocopris erycoides <small>Felsche, 1907</small>
- Heliocopris eryx <small>(Fabricius, 1801)</small>
- Heliocopris faunus <small>Boheman, 1857</small>
- Heliocopris felschei <small>Kolbe, 1904</small>
- Heliocopris fonsecai <small>Ferreira, 1967</small>
- Heliocopris furcithorax <small>Müller, 1941</small>
- Heliocopris gigas <small>(Linnaeus, 1758)</small>
- Heliocopris hamadryas <small>(Fabricius, 1775)</small>
- Heliocopris hamifer <small>Harold, 1878</small>
- Heliocopris haroldi <small>Kolbe, 1893</small>
- Heliocopris helleri <small>Felsche, 1907</small>
- Heliocopris hermes <small>Gillet, 1911</small>
- Heliocopris hunteri <small>Waterhouse, 1891</small>
- Heliocopris japetus <small>Klug, 1855</small>
- Heliocopris kolbei <small>Felsche, 1901</small>
- Heliocopris marshalli <small>Péringuey, 1901</small>
- Heliocopris midas <small>(Fabricius, 1775)</small>
- Heliocopris mimus <small>Janssens, 1939</small>
- Heliocopris minos <small>Gillet, 1907</small>
- Heliocopris mutabilis <small>Kolbe, 1893</small>
- Heliocopris myrmidon <small>Kolbe, 1893</small>
- Heliocopris neptuniformis <small>Felsche, 1907</small>
- Heliocopris neptunoides <small>Janssens, 1939</small>
- Heliocopris neptunus <small>Boheman, 1857</small>
- Heliocopris pauliani <small>Janssens, 1939</small>
- Heliocopris pirmal <small>(Fabricius, 1798)</small>
- Heliocopris quinqueangulatus <small>Janssens, 1939</small>
- Heliocopris samson <small>Harold, 1878</small>
- Heliocopris sirius <small>Gillet, 1925</small>
- Heliocopris solitarius <small>Kolbe, 1893</small>
- Heliocopris staudingeri <small>Kolbe, 1893</small>
- Heliocopris sylvanus <small>Gillet, 1925</small>
- Heliocopris tyrannus <small>Thomson, 1859</small>
Notes
Further reading
- Pokorný, Svatopluk; ZÃÂdek, JiÃ
Âàand Werner, Karl (2009) Giant dung beetles of the genus Heliocopris (Scarabaeidae) Taita Publishers, Hradec Králové, Czech Republic,