Harpalus (beetle)

Harpalus is a genus of ground beetle first described by Pierre André Latreille in 1802.

Species

Harpalus contains the following 430 species:

Subgenus Cephalophonus <small>Ganglbauer, 1891</small>

Harpalus cephalotes <small>Fairmaire & Laboulbène, 1854</small>

Subgenus Cryptophonus <small>Brandmayr & Zetto Brandmayr, 1982</small>

Harpalus agnatus <small>Reiche, 1850</small>

Harpalus cyrenaicus <small>Koch, 1939</small>

Harpalus fulvus <small>Dejean, 1829</small>

Harpalus grilli <small>Kataev, 2002</small>

Harpalus idiotus <small>Bates, 1889</small>

Harpalus janinae <small>Jeanne, 1984</small>

Harpalus litigiosus <small>Dejean, 1829</small>

Harpalus melancholicus <small>Dejean, 1829</small>

Harpalus schaumii <small>Wollaston, 1864</small>

Harpalus tenebrosus <small>Dejean, 1829</small>

Subgenus Glanodes <small>Casey, 1914</small>

Harpalus cohni <small>Ball, 1972</small>

Harpalus corpulentus <small>(Casey, 1914)</small>

Harpalus huachuca <small>Ball, 1972</small>

Harpalus obliquus <small>G.Horn, 1880</small>

Harpalus puncticeps <small>(Casey, 1914)</small>

Harpalus stephani <small>Ball, 1972</small>

Subgenus Harpalus <small>Latreille, 1802</small>

Harpalus acupalpoides <small>Reitter, 1900</small>

Harpalus aeneipennis <small>(Faldermann, 1836)</small>

Harpalus aequicollis <small>Motschulsky, 1844</small>

Harpalus affinis <small>(Schrank, 1781)</small>

Harpalus agakhaniantzi <small>(Mikhailov, 1972)</small>

Harpalus akbensis <small>Jedlicka, 1958</small>

Harpalus akinini <small>Tschitscherine, 1895</small>

Harpalus alajensis <small>Tschitscherine, 1898</small>

Harpalus albanicus <small>Reitter, 1900</small>

Harpalus alexandrae <small>Kataev, 1990</small>

Harpalus alexeevi <small>Kataev, 1990</small>

Harpalus alpivagus <small>Tschitscherine, 1899</small>

Harpalus amarellus <small>Bates, 1891</small>

Harpalus amariformis <small>Motschulsky, 1844</small>

Harpalus amplicollis <small>Ménétriés, 1848</small>

Harpalus amputatus <small>Say, 1830</small>

Harpalus anatolicus <small>Tschitscherine, 1898</small>

Harpalus angulatus <small>Putzeys, 1878</small>

Harpalus angustipennis <small>Boheman, 1848</small>

Harpalus angustitarsis <small>Reitter, 1887</small>

Harpalus animosus <small>Casey, 1924</small>

Harpalus anisodactyliformis <small>Solsky, 1874</small>

Harpalus antonowi <small>(Tschitscherine, 1898)</small>

Harpalus anxioides <small>Kataev, 1991</small>

Harpalus anxius <small>(Duftschmid, 1812)</small>

Harpalus araraticus <small>Mlynar, 1979</small>

Harpalus arcuatus <small>Tschitscherine, 1898</small>

Harpalus arnoldii <small>Kataev, 1988</small>

Harpalus ascetes <small>Kataev & Wrase, 1997</small>

Harpalus asemus <small>Basilewsky, 1946</small>

Harpalus aterrimus <small>Casey, 1914</small>

Harpalus atratus <small>Latreille, 1804</small>

Harpalus atrichatus <small>Hatch, 1949</small>

Harpalus atripes <small>Casey, 1914</small>

Harpalus attenuatus <small>Stephens, 1828</small>

Harpalus australasiae <small>Dejean, 1829</small>

Harpalus autumnalis <small>(Duftschmid, 1812)</small>

Harpalus badakschanus <small>Jedlicka, 1956</small>

Harpalus baleensis <small>Clarke, 1973</small>

Harpalus balli <small>Noonan, 1991</small>

Harpalus basanicus <small>J.Sahlberg, 1913</small>

Harpalus basilewskyi <small>Facchini, 2003</small>

Harpalus basuto <small>Basilewsky, 1958</small>

Harpalus bellieri <small>Reiche, 1861</small>

Harpalus beneshi <small>Kataev & Wrase, 1997</small>

Harpalus brachypterus <small>Tschitscherine, 1898</small>

Harpalus brevicornis <small>Germar, 1823</small>

Harpalus brevis <small>Motschulsky, 1844</small>

Harpalus breviusculus <small>Chaudoir, 1846</small>

Harpalus bucharicus <small>Tschitscherine, 1898</small>

Harpalus bulirschi <small>(Facchini, 2012)</small>

Harpalus bungii <small>Chaudoir, 1844</small>

Harpalus caeruleatus <small>Bates, 1878</small>

Harpalus caiphus <small>Reiche & Saulcy, 1855</small>

Harpalus calathoides <small>Motschulsky, 1844</small>

Harpalus capicola <small>Dejean, 1829</small>

Harpalus cardoni <small>Antoine, 1922</small>

Harpalus caspius <small>(Steven, 1806)</small>

Harpalus cautus <small>Dejean, 1829</small>

Harpalus chalcentus <small>Bates, 1873</small>

Harpalus chasanensis <small>Lafer, 1989</small>

Harpalus chobautianus <small>Lutshnik, 1922</small>

Harpalus chrysopus <small>Reitter, 1887</small>

Harpalus circumpunctatus <small>Chaudoir, 1846</small>

Harpalus cisteloides <small>Motschulsky, 1844</small>

Harpalus clarkei <small>Kataev & J.Schmidt, 2020</small>

Harpalus compressus <small>Motschulsky, 1844</small>

Harpalus contemptus <small>Dejean, 1829</small>

Harpalus convexus <small>Faldermann, 1836</small>

Harpalus corporosus <small>(Motschulsky, 1862)</small>

Harpalus corrugatus <small>Basilewsky, 1958</small>

Harpalus crates <small>Bates, 1883</small>

Harpalus cupreus <small>Dejean, 1829</small>

Harpalus cursorius <small>Péringuey, 1896</small>

Harpalus cyanopterus <small>(Tschitscherine, 1897)</small>

Harpalus cyclogonus <small>Chaudoir, 1844</small>

Harpalus danieli <small>Reitter, 1900</small>

Harpalus davidianus <small>Tschitscherine in Jakobson, 1903</small>

Harpalus decipiens <small>Dejean, 1829</small>

Harpalus defector <small>Péringuey, 1896</small>

Harpalus desertus <small>LeConte, 1859</small>

Harpalus diligens <small>Tschitscherine, 1898</small>

Harpalus dimidiatus <small>(P.Rossi, 1790)</small>

Harpalus dispar <small>Dejean, 1829</small>

Harpalus dissitus <small>Antoine, 1931</small>

Harpalus distinguendus <small>(Duftschmid, 1812)</small>

Harpalus diversicollis <small>Basilewsky, 1958</small>

Harpalus dubius <small>Boheman, 1848</small>

Harpalus dudkoi <small>Kataev, 2011</small>

Harpalus durangoensis <small>Bates, 1891</small>

Harpalus ebeninus <small>Heyden, 1870</small>

Harpalus egenus <small>Dejean, 1829</small>

Harpalus egorovi <small>Lafer, 1989</small>

Harpalus ellipsis <small>LeConte, 1847</small>

Harpalus elliptipennis <small>Facchini, 2003</small>

Harpalus erosus <small>Mannerheim, 1825</small>

Harpalus exiguus <small>Boheman, 1848</small>

Harpalus famelicus <small>Tschitscherine, 1898</small>

Harpalus farkaci <small>Kataev & Wrase, 1995</small>

Harpalus fimetarius <small>Dejean, 1829</small>

Harpalus flavescens <small>(Piller & Mitterpacher, 1783)</small>

Harpalus flavicornis <small>Dejean, 1829</small>

Harpalus foveiger <small>Tschitscherine, 1895</small>

Harpalus franzi <small>Mateu, 1954</small>

Harpalus frater <small>Chaudoir, 1876</small>

Harpalus fraternus <small>LeConte, 1852</small>

Harpalus froelichii <small>Sturm, 1818</small>

Harpalus fulvicornis <small>Thunberg, 1806</small>

Harpalus fulvilabris <small>Mannerheim, 1853</small>

Harpalus fulvipennis <small>Chaudoir, 1843</small>

Harpalus fuscicornis <small>Ménétriés, 1832</small>

Harpalus fuscipalpis <small>Sturm, 1818</small>

Harpalus fuscipennis <small>Wiedemann, 1825</small>

Harpalus fuscoaeneus <small>Dejean, 1829</small>

Harpalus ganssuensis <small>Semenov, 1889</small>

Harpalus giacomazzoi <small>Kataev & Wrase, 1996</small>

Harpalus gilgil <small>Basilewsky, 1946</small>

Harpalus gisellae <small>Csiki, 1932</small>

Harpalus glasunovi <small>Kataev, 1987</small>

Harpalus gravis <small>LeConte, 1858</small>

Harpalus gregoryi <small>Alluaud, 1917</small>

Harpalus hartmanni <small>Kataev, 2002</small>

Harpalus herbivagus <small>Say, 1823</small>

Harpalus heyrovskyi <small>Jedlicka, 1928</small>

Harpalus hiekei <small>Kataev & Wrase, 2010</small>

Harpalus hirtipes <small>(Panzer, 1796)</small>

Harpalus honestus <small>(Duftschmid, 1812)</small>

Harpalus hospes <small>Sturm, 1818</small>

Harpalus hybridus <small>Boheman, 1848</small>

Harpalus impressus <small>Roth, 1851</small>

Harpalus inconcinnus <small>Chaudoir, 1876</small>

Harpalus indianus <small>Csiki, 1932</small>

Harpalus indicola <small>Bates, 1878</small>

Harpalus indigens <small>Casey, 1924</small>

Harpalus inexspectatus <small>Kataev, 1989</small>

Harpalus ingenuus <small>Tschitscherine, 1898</small>

Harpalus innocuus <small>LeConte, 1863</small>

Harpalus italus <small>Schaum, 1860</small>

Harpalus jeanneli <small>Basilewsky, 1946</small>

Harpalus jordanus <small>Jedlicka, 1964</small>

Harpalus kabakianus <small>Kataev, 1988</small>

Harpalus kabakovi <small>Kataev, 1987</small>

Harpalus kadleci <small>Kataev & Wrase, 1995</small>

Harpalus kadyrbekovi <small>Kataev, 1988</small>

Harpalus kagyzmanicus <small>Kataev, 1984</small>

Harpalus kandaharensis <small>Jedlicka, 1956</small>

Harpalus karakorum <small>Jedlicka, 1958</small>

Harpalus karamani <small>Apfelbeck, 1902</small>

Harpalus kazanensis <small>Jedlicka, 1958</small>

Harpalus kaznakovi <small>Kataev & Wrase, 1997</small>

Harpalus kibonoti <small>Alluaud, 1927</small>

Harpalus kirgisicus <small>(Motschulsky, 1844)</small>

Harpalus kiritshenkoi <small>Kataev, 1990</small>

Harpalus klapperichi <small>Jedlicka, 1956</small>

Harpalus kmecoi <small>Facchini, 2003</small>

Harpalus kozlovi <small>Kataev, 1993</small>

Harpalus kryzhanovskii <small>Kataev, 1988</small>

Harpalus kunarensis <small>Kataev, 1993</small>

Harpalus laetus <small>Reiche, 1843</small>

Harpalus laevipes <small>Zetterstedt, 1828</small>

Harpalus lama <small>Kataev & Wrase, 1997</small>

Harpalus lateralis <small>Dejean, 1829</small>

Harpalus laticeps <small>LeConte, 1850</small>

Harpalus latus <small>(Linnaeus, 1758)</small>

Harpalus lederi <small>Tschitscherine, 1899</small>

Harpalus lethierryi <small>Reiche, 1860</small>

Harpalus lewisii <small>LeConte, 1865</small>

Harpalus longipalmatus <small>Mordkovitsh, 1969</small>

Harpalus lopezi <small>J.Serrano & Lencina, 2009</small>

Harpalus lugubris <small>Boheman, 1848</small>

Harpalus lumbaris <small>Mannerheim, 1825</small>

Harpalus luteicornis <small>(Duftschmid, 1812)</small>

Harpalus lutshniki <small>Schauberger, 1932</small>

Harpalus macronotus <small>Tschitscherine, 1893</small>

Harpalus mairei <small>Peyerimhoff, 1928</small>

Harpalus major <small>(Motschulsky, 1850)</small>

Harpalus makhekensis <small>Basilewsky, 1958</small>

Harpalus manas <small>Kataev, 1990</small>

Harpalus marginellus <small>Gyllenhal, 1827</small>

Harpalus martini <small>Van Dyke, 1926</small>

Harpalus masoreoides <small>Bates, 1878</small>

Harpalus massarti <small>Burgeon, 1936</small>

Harpalus mauritanicus <small>Gaubil, 1844</small>

Harpalus medvedevi <small>Kataev, 2006</small>

Harpalus megacephalus <small>LeConte, 1847</small>

Harpalus melaneus <small>Bates, 1878</small>

Harpalus metallinus <small>Ménétriés, 1836</small>

Harpalus metarsius <small>Andrewes, 1930</small>

Harpalus meteorus <small>Basilewsky, 1946</small>

Harpalus michaili <small>Kataev, 1990</small>

Harpalus michailovi <small>Kataev, 1987</small>

Harpalus microthorax <small>(Motschulsky, 1849)</small>

Harpalus miles <small>Péringuey, 1896</small>

Harpalus minutissimus <small>Facchini, 2015</small>

Harpalus minutulus <small>Kataev & Liang, 2004</small>

Harpalus mitridati <small>Pliginskiy, 1915</small>

Harpalus mlynari <small>Kataev, 1990</small>

Harpalus modestus <small>Dejean, 1829</small>

Harpalus morvani <small>Kataev, 2002</small>

Harpalus murzini <small>Kataev, 2008</small>

Harpalus nanniscus <small>Péringuey, 1896</small>

Harpalus natalensis <small>Boheman, 1848</small>

Harpalus natalicus <small>Péringuey, 1896</small>

Harpalus neglectus <small>Audinet-Serville, 1821</small>

Harpalus nevadensis <small>K. & J.Daniel, 1898</small>

Harpalus nigrans <small>A.Morawitz, 1862</small>

Harpalus nigritarsis <small>C.R.Sahlberg, 1827</small>

Harpalus numidicus <small>Bedel, 1893</small>

Harpalus nyassicus <small>Basilewsky, 1946</small>

Harpalus oblitus <small>Dejean, 1829</small>

Harpalus obnixus <small>Casey, 1924</small>

Harpalus ochropus <small>Kirby, 1837</small>

Harpalus oodioides <small>Dejean, 1829</small>

Harpalus opacipennis <small>(Haldeman, 1843)</small>

Harpalus optabilis <small>Dejean, 1829</small>

Harpalus ovtshinnikovi <small>Kataev, 1990</small>

Harpalus pallidipennis <small>A.Morawitz, 1862</small>

Harpalus pallipes <small>Chaudoir, 1837</small>

Harpalus parallelocollis <small>Facchini, 2003</small>

Harpalus parasinuatus <small>Kataev & Liang, 2007</small>

Harpalus parvulus <small>Dejean, 1829</small>

Harpalus petri <small>Tschitscherine, 1902</small>

Harpalus picipennis <small>(Duftschmid, 1812)</small>

Harpalus plancyi <small>Tschitscherine, 1897</small>

Harpalus plenalis <small>Casey, 1914</small>

Harpalus politus <small>Dejean, 1829</small>

Harpalus polyglyptus <small>Schaum, 1862</small>

Harpalus potanini <small>Tschitscherine, 1906</small>

Harpalus praecurrens <small>Schauberger, 1934</small>

Harpalus procognatus <small>Lorenz, 1998</small>

Harpalus progrediens <small>Schauberger, 1922</small>

Harpalus prosperus <small>Basilewsky, 1972</small>

Harpalus providens <small>Casey, 1914</small>

Harpalus pseudoserripes <small>Reitter, 1900</small>

Harpalus pterostichus <small>(Reitter, 1900)</small>

Harpalus puetzi <small>Kataev & Wrase, 1997</small>

Harpalus pulchrinulus <small>Reitter, 1900</small>

Harpalus pulvinatus <small>Ménétriés, 1848</small>

Harpalus pumilus <small>Sturm, 1818</small>

Harpalus punctatostriatus <small>Dejean, 1829</small>

Harpalus punctipennis <small>Mulsant, 1852</small>

Harpalus pusillus <small>(Motschulsky, 1850)</small>

Harpalus pygmaeus <small>Dejean, 1829</small>

Harpalus quadratus <small>Chaudoir, 1846</small>

Harpalus raphaili <small>Kataev, 1997</small>

Harpalus reflexus <small>Putzeys in Schneider & Leder, 1878</small>

Harpalus reitteri <small>Wrase & Kataev, 2011</small>

Harpalus remboides <small>Solsky, 1874</small>

Harpalus reversus <small>Casey, 1924</small>

Harpalus rhemboides <small>Solsky, 1874</small>

Harpalus rivalsi <small>Jeannel, 1948</small>

Harpalus rotundus <small>Facchini, 2003</small>

Harpalus rougemonti <small>Clarke, 1973</small>

Harpalus rubripes <small>(Duftschmid, 1812)</small>

Harpalus rufipalpis <small>Sturm, 1818</small>

Harpalus rufiscapus <small>Gebler, 1833</small>

Harpalus rufitarsoides <small>Schauberger, 1934</small>

Harpalus rufomarginatus <small>(Boheman, 1848)</small>

Harpalus rumelicus <small>Apfelbeck, 1904</small>

Harpalus salinulus <small>Reitter, 1900</small>

Harpalus salinus <small>Dejean, 1829</small>

Harpalus sanctaehelenae <small>Basilewsky, 1972</small>

Harpalus saxicola <small>Dejean, 1829</small>

Harpalus semenowi <small>Tschitscherine, 1901</small>

Harpalus semipunctatus <small>Dejean, 1829</small>

Harpalus serripes <small>(Quensel, 1806)</small>

Harpalus servus <small>(Duftschmid, 1812)</small>

Harpalus setiporus <small>(Reitter, 1894)</small>

Harpalus siculus <small>Dejean, 1829</small>

Harpalus silipes <small>Dejean, 1831</small>

Harpalus sinuaticollis <small>Facchini, 2015</small>

Harpalus sinuatus <small>Tschitscherine, 1893</small>

Harpalus smaragdinus <small>(Duftschmid, 1812)</small>

Harpalus smyrnensis <small>Heyden, 1888</small>

Harpalus solitaris <small>Dejean, 1829</small>

Harpalus somereni <small>Basilewsky, 1946</small>

Harpalus somnulentus <small>Dejean, 1829</small>

Harpalus spadiceus <small>Dejean, 1829</small>

Harpalus spretus <small>Péringuey, 1896</small>

Harpalus spurius <small>Péringuey, 1896</small>

Harpalus stevenii <small>Dejean, 1829</small>

Harpalus stevensi <small>Kataev, 2011</small>

Harpalus stoetznerianus <small>Schauberger, 1932</small>

Harpalus strenuus <small>Tschitscherine, 1898</small>

Harpalus subaeneus <small>Boheman, 1848</small>

Harpalus subcylindricus <small>Dejean, 1829</small>

Harpalus subtruncatus <small>Chaudoir, 1846</small>

Harpalus sulphuripes <small>Germar, 1823</small>

Harpalus sundaicus <small>Schauberger, 1933</small>

Harpalus sushenicus <small>Kataev, 1990</small>

Harpalus suturangulus <small>Reitter, 1887</small>

Harpalus szalliesi <small>Kataev & Wrase, 1995</small>

Harpalus taciturnus <small>Dejean, 1829</small>

Harpalus tangutorum <small>Kataev, 1993</small>

Harpalus tardus <small>(Panzer, 1796)</small>

Harpalus tarsalis <small>Mannerheim, 1825</small>

Harpalus terrestris <small>(Motschulsky, 1844)</small>

Harpalus tibeticus <small>Andrewes, 1930</small>

Harpalus tichonis <small>Jakobson, 1907</small>

Harpalus tiridates <small>Reitter, 1900</small>

Harpalus tithonus <small>Reitter, 1900</small>

Harpalus tjanschanicus <small>Semenov, 1889</small>

Harpalus torosensis <small>Jedlicka, 1961</small>

Harpalus torridoides <small>Reitter, 1900</small>

Harpalus trichophorus <small>Tschitscherine, 1897</small>

Harpalus triseriatus <small>A.Fleischer, 1897</small>

Harpalus turcicus <small>Jedlicka, 1958</small>

Harpalus turmalinus <small>Erichson, 1847</small>

Harpalus udege <small>Lafer, 1989</small>

Harpalus uniformis <small>(Motschulsky, 1844)</small>

Harpalus ussuricus <small>Mlynar, 1979</small>

Harpalus vanemdeni <small>Schauberger, 1932</small>

Harpalus venator <small>Boheman, 1848</small>

Harpalus ventralis <small>LeConte, 1847</small>

Harpalus vereschaginae <small>Kataev, 1988</small>

Harpalus vernicosus <small>Kataev & Liang, 2007</small>

Harpalus viridanus <small>(Motschulsky, 1844)</small>

Harpalus viridicupreus <small>Reiche, 1843</small>

Harpalus vittatus <small>Gebler, 1833</small>

Harpalus wadiensis <small>Jedlicka, 1964</small>

Harpalus wohlberedti <small>Emden, 1932</small>

Harpalus xanthopus <small>Gemminger & Harold, 1868</small>

Harpalus zabroides <small>Dejean, 1829</small>

Harpalus zhdankoi <small>Kataev, 1990</small>

Subgenus Loboharpalus <small>Schauberger, 1932</small>

Harpalus platynotus <small>Bates, 1873</small>

Harpalus rubefactus <small>Bates, 1873</small>

Subgenus Opadius <small>Casey, 1914</small>

Harpalus apache <small>Kataev, 2010</small>

Harpalus cordatus <small>(LeConte, 1853)</small>

Harpalus cordifer <small>Notman, 1919</small>

Subgenus Plectralidus <small>Casey, 1914</small>

Harpalus erraticus <small>Say, 1823</small>

Harpalus retractus <small>LeConte, 1863</small>

Subgenus Pseudoophonus <small>Motschulsky, 1844</small>

Harpalus actiosus <small>Casey, 1914</small>

Harpalus aenigma <small>(Tschitscherine, 1897)</small>

Harpalus alienus <small>Bates, 1878</small>

Harpalus aogashimensis <small>(Habu, 1957)</small>

Harpalus azumai <small>Habu, 1968</small>

Harpalus babai <small>Habu, 1973</small>

Harpalus calceatus <small>(Duftschmid, 1812)</small>

Harpalus caliginosus <small>(Fabricius, 1775)</small>

Harpalus capito <small>A.Morawitz, 1862</small>

Harpalus compar <small>LeConte, 1847</small>

Harpalus coreanus <small>(Tschitscherine, 1895)</small>

Harpalus davidi <small>(Tschitscherine, 1897)</small>

Harpalus disimuciulus <small>Huang; Lei; Yan & Hu, 1996</small>

Harpalus eous <small>Tschitscherine, 1901</small>

Harpalus erythropus <small>Dejean, 1829</small>

Harpalus faunus <small>Say, 1823</small>

Harpalus fokienensis <small>Schauberger, 1930</small>

Harpalus griseus <small>(Panzer, 1796)</small>

Harpalus hatchi <small>Ball & Anderson, 1962</small>

Harpalus hauserianus <small>Schauberger, 1929</small>

Harpalus indicus <small>Bates, 1891</small>

Harpalus jureceki <small>(Jedlicka, 1928)</small>

Harpalus katiae <small>F.Battoni, 1985</small>

Harpalus liobasis <small>Chaudoir, 1868</small>

Harpalus meghalayensis <small>Kataev, 2001</small>

Harpalus meridianus <small>Andrewes, 1923</small>

Harpalus paratus <small>Casey, 1924</small>

Harpalus pastor <small>Motschulsky, 1844</small>

Harpalus pensylvanicus <small>(DeGeer, 1774)</small>

Harpalus poncei <small>Will, 2002</small>

Harpalus protractus <small>Casey, 1914</small>

Harpalus pseudohauserianus <small>Kataev, 2001</small>

Harpalus pseudophonoides <small>Schauberger, 1930</small>

Harpalus roninus <small>Bates, 1873</small>

Harpalus rufipes <small>(DeGeer, 1774)</small>

Harpalus sericatus <small>Tschitscherine, 1906</small>

Harpalus simplicidens <small>Schauberger, 1929</small>

Harpalus singularis <small>Tschitscherine, 1906</small>

Harpalus sinicus <small>Hope, 1845</small>

Harpalus suensoni <small>Kataev, 1997</small>

Harpalus texanus <small>Casey, 1914</small>

Harpalus tridens <small>A.Morawitz, 1862</small>

Harpalus ussuriensis <small>Chaudoir, 1863</small>

Harpalus vagans <small>LeConte, 1865</small>

Harpalus yinchuanensis <small>Huang, 1993</small>

Subgenus Semiophonus <small>Schauberger, 1933</small>

Harpalus signaticornis <small>(Duftschmid, 1812)</small>

Subgenus Zangoharpalus <small>Huang, 1998</small>

Harpalus mariae <small>Kataev, 1997</small>

Harpalus microdemas <small>Schauberger, 1932</small>

Harpalus praticola <small>Bates, 1891</small>

Harpalus pseudotinctulus <small>Schauberger, 1932</small>

Harpalus tinctulus <small>Bates, 1873</small>

Extinct, not assigned to subgenus

†Harpalus abolitus <small>C. & L.Heyden, 1866</small>

†Harpalus anactus <small>Giebel, 1856</small>

†Harpalus bruckmanni <small>Heer, 1862</small>

†Harpalus burmeisteri <small>Giebel, 1856</small>

†Harpalus conditus <small>Scudder, 1900</small>

†Harpalus constrictus <small>Heer, 1862</small>

†Harpalus deletus <small>Oustalet, 1864</small>

†Harpalus dilatatus <small>Theobald, 1937</small>

†Harpalus ewaldi <small>Giebel, 1856</small>

†Harpalus excavatus <small>Forster, 1891</small>

†Harpalus heeri <small>Giebel, 1856</small>

†Harpalus knorri <small>Giebel, 1856</small>

†Harpalus lheritieri <small>Piton, 1940</small>

†Harpalus liasinus <small>Giebel, 1856</small>

†Harpalus maceratus <small>Wickham, 1911</small>

†Harpalus nero <small>Oustalet, 1864</small>

†Harpalus nitens <small>Birket-Smith, 1977</small>

†Harpalus nuperus <small>Scudder, 1900</small>

†Harpalus obtusus <small>Forster, 1891</small>

†Harpalus oustaleti <small>Meunier, 1915</small>

†Harpalus pleistocenicus <small>Lomnicki, 1894</small>

†Harpalus redivivus <small>Wickham, 1917</small>

†Harpalus schlotheimi <small>Giebel, 1856</small>

†Harpalus sinis <small>Heer, 1847</small>

†Harpalus stierlini <small>Heer, 1862</small>

†Harpalus stygius <small>Heer, 1862</small>

†Harpalus tabidus <small>Heer, 1847</small>

†Harpalus tardigradus <small>Heer, 1862</small>

†Harpalus ulomaeformis <small>Wickham, 1917</small>

†Harpalus veterum <small>Cockerell, 1920</small>

†Harpalus whitfieldi <small>Scudder, 1900</small>

References