Harpactea is a genus in the family Dysderidae (woodlouse hunting spiders). Harpactea is a replacement name published by W. S. Bristowe in 1939 for the unavailable name "Harpactes" published by R. Templeton in 1835, which had already been used for a genus of birds. They are non-web building predators that forage on the ground and on tree trunks at night, mainly in forests. During the day, they hide in silk retreats they build under rocks or bark.
Harpactea sadistica was found to use traumatic insemination, the arthropod behavior of directly inserting its sperm into the body cavity of females. It is the first time it has ever been observed in spiders.
Description
Like all woodlouse hunters, Harpactea have six eyes. The type species, H. hombergi, can grow up to a body length of . Males and females are similar, but the female has no epigyne.
Like the rest of their family, they are nocturnal. Unlike them, Harpactea do not specialize on hunting woodlice. H. rubicunda also hunts Drassodes and other spiders, but most Harpactea feed on insects in addition to woodlice.
Taxonomy
In 1835, the name "Harpactes" was published by R. Templeton for a taxon split off from the related genus Dysdera. Subsequent authors used this genus name for many years, but when published, Harpactes had already been used for a bird genus, so it was not available. Accordingly, in 1939, W. S. Bristowe published the replacement name Harpactea. The type species is Dysdera latreillei, synonym "Harpactes" latreillei, now accepted as a synonym of Harpactea hombergi.
Templeton did not explain his choice of genus name, but Greek , , means 'snatcher', 'plunderer', 'pillager' or 'thief'.
Species
Almost all species of this genus appear to be endemic to small regions of the Mediterranean. , the World Spider Catalog accepted 213 species from Europe and Northern Africa to Turkmenistan and Iran:
- Harpactea abantia <small>(Simon, 1885)</small> â Greece
- Harpactea abasgiana <small>Zamani & Marusik, 2024</small> â Georgia
- Harpactea achsuensis <small>Dunin, 1991</small> â Azerbaijan
- Harpactea acuta <small>Beladjal & Bosmans, 1997</small> â Algeria
- Harpactea adicensis <small>Ã
ÂezáÃÂ, 2023</small> â Portugal
- Harpactea aeoliensis <small>Alicata, 1973</small> â Italy
- Harpactea agnolettii <small>Brignoli, 1978</small> â Turkey
- Harpactea alanyana <small>ÃÂzkütük, Elverici, Marusik & Kunt, 2015</small> â Turkey
- Harpactea albanica <small>(Caporiacco, 1949)</small> â Albania
- Harpactea alexandrae <small>Lazarov, 2006</small> â Bulgaria, Romania, Ukraine, Russia (Caucasus)
- Harpactea algarvensis <small>Ferrández, 1990</small> â Portugal
- Harpactea alicatai <small>Brignoli, 1979</small> â Italy (Sardinia)
- Harpactea angustata <small>(Lucas, 1846)</small> â Algeria
- Harpactea antoni <small>Bosmans, 2009</small> â Greece (Lesbos)
- Harpactea apollinea <small>Brignoli, 1979</small> â Greece
- Harpactea arguta <small>(Simon, 1907)</small> â France, Italy
- Harpactea armenica <small>Dunin, 1989</small> â Armenia, Iran
- Harpactea arnedoi <small>Kunt, Elverici, ÃÂzkütük & YaÃÂmur, 2011</small> â Turkey
- Harpactea asparuhi <small>Lazarov, 2008</small> â Bulgaria
- Harpactea auresensis <small>Bosmans & Beladjal, 1991</small> â Algeria
- Harpactea auriga <small>(Simon, 1911)</small> â Algeria
- Harpactea aurigoides <small>Bosmans & Beladjal, 1991</small> â Algeria
- Harpactea azerbajdzhanica <small>Dunin, 1991</small> â Azerbaijan
- Harpactea azowensis <small>Charitonov, 1956</small> â Ukraine, Russia (Europe)
- Harpactea babori <small>(Nosek, 1905)</small> â Bulgaria, Greece, Turkey
- Harpactea ballarini <small>Kunt, ÃÂzkütük & Elverici, 2013</small> â Turkey
- Harpactea bilecenoglui <small>Kunt & ÃÂzkütük, 2023</small> â Turkey
- Harpactea bilgenur <small>Kunt & ÃÂzkütük, 2023</small> â Turkey
- Harpactea blasi <small>Ribera & Ferrández, 1986</small> â Spain
- Harpactea brachati <small>Wunderlich, 2020</small> â Turkey
- Harpactea buchari <small>Dunin, 1991</small> â Azerbaijan
- Harpactea bulgarica <small>Lazarov & Naumova, 2010</small> â North Macedonia, Bulgaria
- Harpactea caligata <small>Beladjal & Bosmans, 1997</small> â Algeria
- Harpactea carusoi <small>Alicata, 1974</small> â Italy, Tunisia
- Harpactea catholica <small>(Brignoli, 1984)</small> â Greece (Crete)
- Harpactea caucasia <small>(KulczyÃ
Âski, 1895)</small> â Russia (Caucasus), Georgia
- Harpactea cecconii <small>(KulczyÃ
Âski, 1908)</small> â Cyprus, Greece?
- Harpactea cesari <small>Van Keer, 2009</small> â Greece (Lesbos)
- Harpactea chaniaensis <small>Bosmans, 2023</small> â Greece (Crete)
- Harpactea chreensis <small>Bosmans & Beladjal, 1989</small> â Algeria
- Harpactea christae <small>Bosmans & Beladjal, 1991</small> â Algeria
- Harpactea christodeltshevi <small>Bayram, Kunt & YaÃÂmur, 2009</small> â Turkey
- Harpactea clementi <small>Bosmans, 2009</small> â Greece, Turkey
- Harpactea coccifera <small>Brignoli, 1984</small> â North Macedonia, Greece (Crete)
- Harpactea colchidis <small>Brignoli, 1978</small> â Turkey
- Harpactea complicata <small>Deltshev, 2011</small> â Serbia
- Harpactea corinthia <small>Brignoli, 1984</small> â Greece
- Harpactea corticalis <small>(Simon, 1882)</small> â France, Italy
- Harpactea crespoi <small>Ã
ÂezáÃÂ, 2023</small> â Portugal
- Harpactea cressa <small>Brignoli, 1984</small> â Greece (Crete)
- Harpactea cruriformis <small>Bosmans, 2011</small> â Greece (Chios)
- Harpactea damini <small>Pavlek & Arnedo, 2020</small> â Croatia
- Harpactea dashdamirovi <small>Dunin, 1993</small> â Azerbaijan
- Harpactea decebali <small>Nae, 2021</small> â Romania
- Harpactea deelemanae <small>Dunin, 1989</small> â Armenia
- Harpactea deltshevi <small>Dimitrov & Lazarov, 1999</small> â Bulgaria
- Harpactea dianae <small>Gavalas & Bosmans, 2023</small> â Greece (Iraklia)
- Harpactea digiovannii <small>Gasparo, 2014</small> â Greece
- Harpactea diraoi <small>Brignoli, 1978</small> â Turkey
- Harpactea dobati <small>Alicata, 1974</small> â Turkey
- Harpactea doblikae <small>(Thorell, 1875)</small> â Bulgaria, Ukraine
- Harpactea dolanskyi <small>Ã
ÂezáÃÂ, 2023</small> â Portugal
- Harpactea dufouri <small>(Thorell, 1873)</small> â Spain
- Harpactea dumonti <small>Bosmans & Beladjal, 1991</small> â Algeria
- Harpactea dunini <small>Zamani & Marusik, 2024</small> â Azerbaijan
- Harpactea elvericii <small>Kunt & ÃÂzkütük, 2023</small> â Turkey
- Harpactea eskovi <small>Dunin, 1989</small> â Georgia, Armenia
- Harpactea fageli <small>Brignoli, 1980</small> â Portugal, Spain
- Harpactea fontetareja <small>Wunderlich, 2024</small> â Portugal
- Harpactea forceps <small>Varol & DanñÃ
Âman, 2018</small> â Turkey
- Harpactea forcipifera <small>(Simon, 1911)</small> â Algeria
- Harpactea gaditana <small>Pesarini, 1988</small> â Portugal, Spain
- Harpactea galatica <small>Brignoli, 1978</small> â Turkey
- Harpactea gennargentu <small>Wunderlich, 1995</small> â Italy (Sardinia)
- Harpactea globifera <small>(Simon, 1911)</small> â Algeria
- Harpactea gobustanica <small>Nuruyeva & Huseynov, 2022</small> â Azerbaijan
- Harpactea golovatchi <small>Dunin, 1989</small> â Armenia
- Harpactea gridellii <small>(Caporiacco, 1951)</small> â Italy
- Harpactea grisea <small>(Canestrini, 1868)</small> â Switzerland, Austria, Italy, Slovenia
- Harpactea gunselorum <small>Gücel, Fuller, Göçmen & Kunt, 2018</small> â Cyprus
- Harpactea hamidae <small>Beladjal & Abrous, 2023</small> â Algeria
- Harpactea hauseri <small>Brignoli, 1976</small> â Greece
- Harpactea heizerensis <small>Bosmans & Beladjal, 1991</small> â Algeria
- Harpactea heliconia <small>Brignoli, 1984</small> â Greece
- Harpactea henriquesi <small>Ã
ÂezáÃÂ, 2023</small> â Portugal
- Harpactea henschi <small>(KulczyÃ
Âski, 1915)</small> â Bosnia and Herzegovina
- Harpactea herodis <small>Brignoli, 1978</small> â Israel
- Harpactea hispana <small>(Simon, 1882)</small> â Spain, France
- Harpactea hombergi <small>(Scopoli, 1763)</small> (type species) â Europe
- Harpactea hyrcanica <small>Dunin, 1991</small> â Azerbaijan
- Harpactea iasoni <small>Gavalas & Bosmans, 2023</small> â Greece (Iraklia)
- Harpactea ice <small>Komnenov & Chatzaki, 2016</small> â Greece
- Harpactea incerta <small>Brignoli, 1979</small> â Greece
- Harpactea incurvata <small>Bosmans & Beladjal, 1991</small> â Algeria
- Harpactea indistincta <small>Dunin, 1991</small> â Russia (Caucasus), Azerbaijan
- Harpactea innupta <small>Beladjal & Bosmans, 1997</small> â Algeria
- Harpactea isaurica <small>Brignoli, 1978</small> â Turkey
- Harpactea johannitica <small>Brignoli, 1976</small> â Greece
- Harpactea kalaensis <small>Beladjal & Bosmans, 1997</small> â Algeria
- Harpactea kalavachiana <small>Gücel, Charalambidou, Göçmen & Kunt, 2019</small> â Cyprus
- Harpactea kankilicorum <small>Kunt & ÃÂzkütük, 2023</small> â Turkey
- Harpactea karabachica <small>Dunin, 1991</small> â Azerbaijan
- Harpactea karaschkhan <small>Kunt, ÃÂzkütük, Elverici, Marusik & KarakaÃ
Â, 2016</small> â Turkey
- Harpactea kareli <small>Bosmans & Beladjal, 1991</small> â Algeria
- Harpactea kencei <small>Kunt, Elverici, ÃÂzkütük & YaÃÂmur, 2011</small> â Turkey
- Harpactea konradi <small>Lazarov, 2009</small> â Bulgaria
- Harpactea korenkoi <small>Ã
ÂezáÃÂ, 2023</small> â Portugal
- Harpactea korgei <small>Brignoli, 1979</small> â Turkey
- Harpactea krejcii <small>Ã
ÂezáÃÂ, 2023</small> â Portugal
- Harpactea krueperi <small>(Simon, 1885)</small> â Greece
- Harpactea krumi <small>Lazarov, 2010</small> â Bulgaria
- Harpactea kubrati <small>Lazarov, 2008</small> â Bulgaria
- Harpactea kulczynskii <small>Brignoli, 1976</small> â Albania, Greece
- Harpactea lazarovi <small>Deltshev, 2011</small> â Bulgaria
- Harpactea lazonum <small>Brignoli, 1978</small> â Turkey
- Harpactea lepida <small>(C. L. Koch, 1838)</small> â Europe
- Harpactea loebli <small>Brignoli, 1974</small> â Greece
- Harpactea logunovi <small>Dunin, 1992</small> â Russia (Caucasus), Georgia
- Harpactea longitarsa <small>Alicata, 1974</small> â Algeria, Tunisia
- Harpactea longobarda <small>Pesarini, 2001</small> â Spain (Balearic Is.), Italy, Ukraine
- Harpactea lucdevosi <small>Bosmans, 2023</small> â Greece (Iraklia)
- Harpactea maelfaiti <small>Beladjal & Bosmans, 1997</small> â Algeria
- Harpactea magnibulbi <small>Machado & Ferrández, 1991</small> â Portugal
- Harpactea major <small>(Simon, 1911)</small> â Algeria
- Harpactea mariae <small>Komnenov, 2014</small> â North Macedonia
- Harpactea martensi <small>Dunin, 1991</small> â Azerbaijan
- Harpactea mateparlovi <small>Pavlek & Arnedo, 2020</small> â Croatia
- Harpactea mcheidzeae <small>Dunin, 1992</small> â Georgia
- Harpactea medeae <small>Brignoli, 1978</small> â Turkey
- Harpactea mehennii <small>Bosmans & Beladjal, 1989</small> â Algeria
- Harpactea mentor <small>Lazarov & Naumova, 2010</small> â North Macedonia, Bulgaria
- Harpactea mertensi <small>Bosmans & Beladjal, 1991</small> â Algeria
- Harpactea minoccii <small>Ferrández, 1982</small> â Portugal, Spain
- Harpactea minuta <small>Alicata, 1974</small> â Tunisia
- Harpactea mithridatis <small>Brignoli, 1979</small> â Turkey, Georgia
- Harpactea mitidjae <small>Bosmans & Beladjal, 1991</small> â Algeria
- Harpactea modesta <small>Dunin, 1991</small> â Caucasus (Russia, Georgia, Azerbaijan)
- Harpactea monicae <small>Bosmans & Beladjal, 1991</small> â Algeria
- Harpactea mouzaiensis <small>Bosmans & Beladjal, 1989</small> â Algeria
- Harpactea muscicola <small>(Simon, 1882)</small> â France (Corsica)
- Harpactea nachitschevanica <small>Dunin, 1991</small> â Azerbaijan
- Harpactea nausicaae <small>Brignoli, 1976</small> â Albania, Kosovo, North Macedonia, Greece
- Harpactea nenilini <small>Dunin, 1989</small> â Armenia
- Harpactea nuragica <small>Alicata, 1966</small> â Italy (Sardinia)
- Harpactea oglasana <small>Gasparo, 1992</small> â Italy
- Harpactea oranensis <small>Bosmans & Beladjal, 1991</small> â Algeria
- Harpactea ortegai <small>Ribera & De Mas, 2003</small> â Spain
- Harpactea osellai <small>Brignoli, 1978</small> â Turkey
- Harpactea ouarsenensis <small>Bosmans & Beladjal, 1991</small> â Algeria
- Harpactea ovata <small>Beladjal & Bosmans, 1997</small> â Algeria
- Harpactea paradoxa <small>Dunin, 1992</small> â Georgia
- Harpactea parthica <small>Brignoli, 1980</small> â Iran, Turkmenistan?
- Harpactea pekari <small>Ã
ÂezáÃÂ, 2023</small> â Portugal
- Harpactea persephone <small>Gasparo, 2011</small> â Greece (Crete)
- Harpactea petrovi <small>Lazarov & Dimitrov, 2018</small> â Bulgaria
- Harpactea piligera <small>(Thorell, 1875)</small> â Italy
- Harpactea pisidica <small>Brignoli, 1978</small> â Turkey
- Harpactea planembolus <small>Wunderlich, 2024</small> â Portugal
- Harpactea popovi <small>Dimitrov, Deltshev & Lazarov, 2019</small> â Bulgaria
- Harpactea proxima <small>Ferrández, 1990</small> â Portugal
- Harpactea pugio <small>Varol & Akpñnar, 2016</small> â Turkey
- Harpactea punica <small>Alicata, 1974</small> â Algeria, Tunisia
- Harpactea reniformis <small>Beladjal & Bosmans, 1997</small> â Algeria
- Harpactea rubicunda <small>(C. L. Koch, 1838)</small> â Europe, Caucasus (Russia, Georgia)
- Harpactea rucnerorum <small>Polenec & Thaler, 1975</small> â Croatia
- Harpactea ruffoi <small>Alicata, 1974</small> â Tunisia
- Harpactea rugichelis <small>Denis, 1955</small> â Lebanon
- Harpactea sadistica <small>Ã
ÂezáÃÂ, 2008</small> â Israel
- Harpactea saeva <small>(Herman, 1879)</small> â Slovakia, Hungary, south-eastern Europe to Ukraine
- Harpactea salvatorei <small>Platania & Arnedo, 2020</small> â Italy
- Harpactea samuili <small>Lazarov, 2006</small> â North Macedonia, Bulgaria, Greece
- Harpactea sanctaeinsulae <small>Brignoli, 1978</small> â Turkey
- Harpactea sanctidomini <small>Gasparo, 1997</small> â Italy
- Harpactea sardoa <small>Alicata, 1966</small> â Italy
- Harpactea sbordonii <small>Brignoli, 1978</small> â Turkey
- Harpactea sciakyi <small>Pesarini, 1988</small> â Spain
- Harpactea secunda <small>Dunin, 1989</small> â Armenia
- Harpactea senalbensis <small>Beladjal & Bosmans, 1997</small> â Algeria
- Harpactea serena <small>(Simon, 1907)</small> â Spain, France
- Harpactea sicula <small>Alicata, 1966</small> â Italy (Sicily), Malta
- Harpactea simovi <small>Deltshev & Lazarov, 2018</small> â Bulgaria
- Harpactea sinuata <small>Beladjal & Bosmans, 1997</small> â Algeria
- Harpactea spasskyi <small>Dunin, 1992</small> â Ukraine (Crimea), Caucasus (Russia, Georgia)
- Harpactea spilioioannis <small>Bosmans & Gavalas, 2023</small> â Greece (Iraklia)
- Harpactea spirembolus <small>Russell-Smith, 2011</small> â Greece (Chios)
- Harpactea srednagora <small>Dimitrov & Lazarov, 1999</small> â Albania, North Macedonia, Bulgaria
- Harpactea stalitoides <small>Ribera, 1993</small> â Portugal
- Harpactea stoevi <small>Deltshev & Lazarov, 2018</small> â Bulgaria
- Harpactea strandi <small>(Caporiacco, 1939)</small> â Italy
- Harpactea strandjica <small>Dimitrov, 1997</small> â Bulgaria, Turkey
- Harpactea strinatii <small>Brignoli, 1979</small> â Greece
- Harpactea sturanyi <small>(Nosek, 1905)</small> â Greece, Turkey, Georgia
- Harpactea subiasi <small>Ferrández, 1990</small> â Portugal
- Harpactea talyschica <small>Dunin, 1991</small> â Azerbaijan
- Harpactea tavirensis <small>Wunderlich, 2020</small> â Portugal
- Harpactea tenuiemboli <small>Deltshev, 2011</small> â Serbia
- Harpactea tergestina <small>Gasparo, 2014</small> â Italy
- Harpactea terveli <small>Lazarov, 2009</small> â Bulgaria, Turkey
- Harpactea thaleri <small>Alicata, 1966</small> â Switzerland, Italy
- Harpactea umay <small>Kunt, YaÃÂmur & ÃÂzkütük, 2023</small> â Turkey
- Harpactea undosa <small>Beladjal & Bosmans, 1997</small> â Algeria
- Harpactea vagabunda <small>Dunin, 1991</small> â Azerbaijan
- Harpactea vignai <small>Brignoli, 1978</small> â Turkey
- Harpactea villehardouini <small>Brignoli, 1979</small> â Greece
- Harpactea walterdebucki <small>Bosmans, 2023</small> â Greece (Iraklia)
- Harpactea wolfgangi <small>Komnenov & Chatzaki, 2016</small> â Greece
- Harpactea yakourensis <small>Beladjal & Bosmans, 1997</small> â Algeria
- Harpactea yanardagi <small>Kunt & ÃÂzkütük, 2023</small> â Turkey
- Harpactea zagros <small>Zamani & Marusik, 2023</small> â Iran
- Harpactea zaitzevi <small>Charitonov, 1956</small> â Georgia, Azerbaijan, Armenia
- Harpactea zannonensis <small>Alicata, 1966</small> â Italy
- Harpactea zjuzini <small>Dunin, 1991</small> â Azerbaijan
- Harpactea zoiai <small>Gasparo, 1999</small> â Greece
References
Further reading
External links