Gymnothorax is a genus of fish in the family Muraenidae found in Atlantic, Indian, and Pacific Oceans. With more than 120 species, it the most speciose genus of moray eels.
Gymnothorax as currently recognized is polyphyletic, and comparative studies are needed before action is taken to resurrect generic synonyms of Gymnothorax.
Description
Species in this genus are slender (most skinny one being Gymnothorax gracilicauda) as are most eels. They grow to . On the opposite end of the spectrum, some species only grow to a maximum size of less than . Many species have complex patterns, but many others are quite bland without much coloration on their bodies. Moray eel species, inevitably applying to Gymnothorax owing to its high diversity, are often found in generally shallow-water habitats, but many species are also found in deep water, being more common in those environments than previously thought.
Like all moray eels, they have pharyngeal jaws.
Species
Both FishBase and World Register of Marine Species recognize a total of 128 valid species in the genus. However, each database includes three species that the other does not list, which are noted in the list below.
- Gymnothorax afer <small>Bloch, 1795</small> (Dark moray)
- Gymnothorax albimarginatus <small>(Temminck & Schlegel, 1846)</small> (White-margin moray)
- Gymnothorax andamanensis
- Gymnothorax angusticauda <small>(M. C. W. Weber & de Beaufort, 1916)</small> (Narrow-tail moray)
- Gymnothorax angusticeps <small>(Hildebrand & F. O. Barton, 1949)</small>
- Gymnothorax annasona <small>Whitley, 1937</small> (Lord Howe Island moray)
- Gymnothorax annulatus <small>D. G. Smith & E. B. Böhlke, 1997</small> (Ringed moray)
- Gymnothorax arabicus <small>D. G. Smith, Bogorodsky, Dandar & Zajonz, 2024</small>
- Gymnothorax atolli <small>(Pietschmann, 1935)</small> (Atoll moray)
- Gymnothorax aurocephalus <small>Nashad, Mohapatra, Varghese & Bineesh, 2020</small> (Goldenhead moray)
- Gymnothorax australicola <small>Lavenberg, 1992</small> (South Pacific moray)
- Gymnothorax austrinus <small>E. B. Böhlke & McCosker, 2001</small> (Southern moray)
- Gymnothorax bacalladoi <small>E. B. Böhlke & Brito, 1987</small> (Canary moray)
- Gymnothorax baranesi <small>D. G. Smith, Brokovich & Einbinder, 2008</small>
- Gymnothorax bathyphilus <small>J. E. Randall & McCosker, 1975</small> (Deep-dwelling moray)
- Gymnothorax berndti <small>Snyder, 1904</small> (Y-patterned moray)
- Gymnothorax breedeni <small>McCosker & J. E. Randall, 1977</small> (Black-cheek moray)
- Gymnothorax buroensis <small>(Bleeker, 1857)</small> (Vagrant moray)
- Gymnothorax castaneus <small>(D. S. Jordan & C. H. Gilbert, 1883)</small> (Panamic green moray)
- Gymnothorax castlei <small>E. B. Böhlke & J. E. Randall, 1999</small> (Castle's moray)
- Gymnothorax cephalospilus <small>E. B. Böhlke & McCosker, 2001</small> (Head-spot moray)
- Gymnothorax chilospilus <small>Bleeker, 1864</small> (Lip-spot moray)
- Gymnothorax chlamydatus <small>Snyder, 1908</small> (Banded mud moray)
- Gymnothorax cinerascens (Plain moray)
- Gymnothorax conspersus <small>Poey, 1867</small> (Saddled moray)
- Gymnothorax cribroris <small>Whitley, 1932</small> (Sieve-patterned moray)
- Gymnothorax davidsmithi <small>McCosker & J. E. Randall, 2008</small> (Flores mud moray)
- Gymnothorax dorsalis <small>Seale, 1917</small>
- Gymnothorax dovii <small>(Günther, 1870)</small> (Fine-spotted moray)
- Gymnothorax elaineheemstrae
- Gymnothorax elegans <small>Bliss, 1883</small> (Elegant moray)
- Gymnothorax emmae <small>Prokofiev, 2010</small>
- Gymnothorax enigmaticus <small>McCosker & J. E. Randall, 1982</small> (Enigmatic moray)
- Gymnothorax equatorialis <small>(Hildebrand, 1946)</small> (Spot-tail moray)
- Gymnothorax eurostus <small>(C. C. Abbott, 1860)</small> (Abbott's moray)
- Gymnothorax eurygnathos <small>E. B. Böhlke, 2001</small>
- Gymnothorax favagineus <small>Bloch & J. G. Schneider, 1801</small> (Laced moray)
- Gymnothorax fimbriatus <small>(E. T. Bennett, 1832)</small> (Fimbriated moray)
- Gymnothorax flavimarginatus <small>(Rüppell, 1830)</small> (Yellow-edged moray)
- Gymnothorax flavoculus <small>(E. B. Böhlke & J. E. Randall, 1996)</small> (Pale-nose moray)
- Gymnothorax formosus <small>Bleeker, 1864</small>
- Gymnothorax funebris <small>Ranzani, 1839</small> (Green moray)
- Gymnothorax fuscomaculatus <small>(L. P. Schultz, 1953)</small> (Brown-spotted moray)
- Gymnothorax gracilicauda <small>O. P. Jenkins, 1903</small> (Slender-tail moray)
- Gymnothorax griseus <small>(Lacépède, 1803)</small> (Geometric moray)
- Gymnothorax hansi <small>Heemstra, 2004</small>
- Gymnothorax hepaticus <small>(Rüppell, 1830)</small> (Liver-colored moray)
- Gymnothorax herrei <small>Beebe & Tee-Van, 1933</small> (Herre's moray)
- Gymnothorax hubbsi <small>J. E. Böhlke & E. B. Böhlke, 1977</small> (Lichen moray)
- Gymnothorax indicus <small>Mohapatra, D. Ray, D. G. Smith & Mishra, 2016</small> (Indian unpatterened moray)
- Gymnothorax intesi <small>(Fourmanoir & Rivaton, 1979)</small> (White-tip moray)
- Gymnothorax isingteena <small>(J. Richardson, 1845)</small> (Indo-Pacific spotted moray)
- Gymnothorax javanicus <small>(Bleeker, 1859)</small> (Giant moray)
- Gymnothorax johnsoni <small>(J. L. B. Smith, 1962)</small> (White-spotted moray)
- Gymnothorax kidako <small>(Temminck & Schlegel, 1846)</small> (Kidako moray)
- Gymnothorax kolpos <small>J. E. Böhlke & E. B. Böhlke, 1980</small> (Black-tail moray)
- Gymnothorax kontodontos <small>E. B. Böhlke, 2000</small> (Short-tooth moray)
- Gymnothorax longinquus <small>(Whitley, 1948)</small>
- Gymnothorax maderensis <small>(J. Y. Johnson, 1862)</small> (Shark-tooth moray)
- Gymnothorax mareei <small>Poll, 1953</small> (Spot-jaw moray)
- Gymnothorax margaritophorus <small>Bleeker, 1864</small> (Blotch-necked moray)
- Gymnothorax marshallensis <small>(L. P. Schultz, 1953)</small> (Marshall Islands moray)
- Gymnothorax mccoskeri <small>D. G. Smith & E. B. Böhlke, 1997</small>
- Gymnothorax megaspilus <small>E. B. Böhlke & J. E. Randall, 1995</small> (Oman moray)
- Gymnothorax melanosomatus <small>K. H. Loh, K. T. Shao & H. M. Chen, 2011</small> (Black-body moray)
- Gymnothorax melatremus <small>L. P. Schultz, 1953</small> (Dwarf moray)
- Gymnothorax meleagris <small>(G. Shaw, 1795)</small> (Turkey moray)
- Gymnothorax microspila <small>(Günther, 1870)</small>
- Gymnothorax microstictus <small>E. B. Böhlke, 2000</small> (Small-spot moray)
- Gymnothorax miliaris <small>(Kaup, 1856)</small> (Golden-tail moray)
- Gymnothorax minor <small>(Temminck & Schlegel, 1846)</small>
- Gymnothorax mishrai <small>D. Ray, Mohapatra & D. G. Smith, 2015</small> (Bengal moray)
- Gymnothorax moluccensis <small>(Bleeker, 1864)</small> (Moluccan moray)
- Gymnothorax monochrous <small>(Bleeker, 1856)</small> (Drab moray)
- Gymnothorax monostigma <small>(Regan, 1909)</small> (One-spot moray)
- Gymnothorax mordax <small>(Ayres, 1859)</small> (California moray)
- Gymnothorax moringa <small>(G. Cuvier, 1829)</small> (Spotted moray)
- Gymnothorax mucifer <small>Snyder, 1904</small>
- Gymnothorax nasuta <small>F. de Buen, 1961</small> (Easter Island moray)
- Gymnothorax neglectus <small>S. Tanaka (I), 1911</small>
- Gymnothorax nigromarginatus <small>(Girard, 1858)</small> (Black-edge moray)
- Gymnothorax niphostigmus <small>H. M. Chen, K. T. Shao & C. T. Chen, 1996</small> (Snowflake-patched moray)
- Gymnothorax nubilus <small>(J. Richardson, 1848)</small> (Grey moray)
- Gymnothorax nudivomer <small>(Günther, 1867)</small> (Yellow-mouth moray)
- Gymnothorax nuttingi <small>Snyder, 1904</small> (Nutting's moray)
- Gymnothorax obesus <small>(Whitley, 1932)</small> (Griffin's moray)
- Gymnothorax ocellatus <small>Agassiz, 1831</small> (Caribbean ocellated moray)
- Gymnothorax odishi <small>Mohapatra, Mohanty, Smith, Mishra, and Roy, 2018</small>
- Gymnothorax panamensis <small>(Steindachner, 1876)</small> (Masked moray)
- Gymnothorax parini <small>Collette, D. G. Smith & E. B. Böhlke, 1991</small>
- Gymnothorax phalarus <small>W. A. Bussing, 1998</small>
- Gymnothorax pharaonis <small>D. G. Smith, S. V. Bogordosky, A. O. Mal, and T. J. Alpermann, 2019</small> (Pharaoh's moray)
- Gymnothorax phasmatodes <small>(J. L. B. Smith], 1962)</small> (Ghost moray)
- Gymnothorax philippinus <small>D. S. Jordan & Seale, 1907</small> (Philippines moray)
- Gymnothorax pictus <small>(J. N. Ahl, 1789)</small> (Paint-spotted moray)
- Gymnothorax pikei <small>Bliss, 1883</small> (Pike's moray)
- Gymnothorax pindae <small>J. L. B. Smith, 1962</small> (Pinda moray)
- Gymnothorax poikilospilus <small>Chen & Huang, 2022</small>
- Gymnothorax polygonius <small>Poey, 1875</small> (Polygon moray)
- Gymnothorax polyspondylus <small>E. B. Böhlke & J. E. Randall, 2000</small>
- Gymnothorax polyuranodon <small>(Bleeker, 1854)</small> (Freshwater moray)
- Gymnothorax porphyreus <small>(Guichenot, 1848)</small> (Low-fin moray)
- Gymnothorax prasinus <small>(J. Richardson, 1848)</small> (Yellow moray)
- Gymnothorax prionodon <small>J. D. Ogilby, 1895</small> (Australian mottled moray)
- Gymnothorax prismodon <small>E. B. Böhlke & J. E. Randall, 2000</small>
- Gymnothorax prolatus <small>K. Sasaki & Amaoka, 1991</small>
- Gymnothorax pseudoherrei <small>E. B. Böhlke, 2000</small> (False brown moray)
- Gymnothorax pseudokidako <small>Huang, Loh & Liao, 2021</small>
- Gymnothorax pseudomelanosomatus <small>K. H. Loh, K. T. Shao & H. M. Chen, 2015</small> (False black-body moray)
- Gymnothorax pseudoprolatus (Short-tail Brown moray)
- Gymnothorax pseudothyrsoideus <small>(Bleeker, 1853)</small> (High-fin moray)
- Gymnothorax pseudotile <small>Mohapatra, Smith, Ray, Mishra & Mohanty, 2017</small> (Bengal low-fin moray)
- Gymnothorax punctatofasciatus <small>Bleeker, 1863</small> (Bars'n-spots moray)
- Gymnothorax punctatus <small>Bloch & J. G. Schneider, 1801</small> (Red Sea white-spotted moray)
- Gymnothorax randalli <small>D. G. Smith & E. B. Böhlke, 1997</small> (Randall's moray)
- Gymnothorax reevesii <small>(J. Richardson, 1845)</small> (Reeve's moray)
- Gymnothorax reticularis <small>Bloch, 1795</small>
- Gymnothorax richardsonii <small>(Bleeker, 1852)</small> (Richardson's moray)
- Gymnothorax robinsi <small>E. B. Böhlke, 1997</small> (Robins' moray)
- Gymnothorax rueppelliae <small>(McClelland, 1844)</small> (Banded moray)
- Gymnothorax ryukyuensis <small>Hatooka, 2003</small>
- Gymnothorax sagenodeta <small>(Richardson, 1848)</small> Gymnothorax sagmacephalus <small>E. B. Böhlke, 1997</small>
- Gymnothorax saxicola <small>D. S. Jordan & B. M. Davis, 1891</small> (Ocellated moray)
- Gymnothorax serratidens <small>(Hildebrand & F. O. Barton, 1949)</small>
- Gymnothorax shaoi <small>H. M. Chen & K. H. Loh, 2007</small>
- Gymnothorax sokotrensis <small>Kotthaus, 1968</small>
- Gymnothorax steindachneri <small>D. S. Jordan & Evermann, 1903</small> (Steindachner's moray)
- Gymnothorax taiwanensis <small>H. M. Chen, K. H. Loh & K. T. Shao, 2008</small> (Taiwanese moray)
- Gymnothorax tamilnaduensis <small>Kodeeswaran, Kantharajan, Mohapatra, Kumar & Sarkar 2023</small>
- Gymnothorax thyrsoideus <small>(J. Richardson, 1845)</small> (Grey-face moray)
- Gymnothorax tile <small>(F. Hamilton, 1822)</small> (Indian mud moray)
- Gymnothorax undulatus <small>(Lacépède, 1803)</small> (Undulated moray)
- Gymnothorax unicolor <small>(Delaroche, 1809)</small> (Brown moray)
- Gymnothorax vagrans <small>(Seale, 1917)</small>
- Gymnothorax verrilli <small>(D. S. Jordan & C. H. Gilbert, 1883)</small> (White-edged moray)
- Gymnothorax vicinus <small>(Castelnau, 1855)</small> (Purple-mouth moray)
- Gymnothorax vietnamensis (Vietnamese moray)
- Gymnothorax visakhaensis <small>Mohapatra, Smith, Mohanty, Mishra & Tudu, 2017</small>
- Gymnothorax walvisensis <small>Prokofiev, 2009</small>
- Gymnothorax woodwardi <small>McCulloch, 1912</small>
- Gymnothorax ypsilon <small>Hatooka & J. E. Randall, 1992</small> (Y-bar moray)
- Gymnothorax zonipectis <small>Seale, 1906</small> (Barred-fin moray)
Notes
References