Geophagus is a genus of cichlids that mainly live in South America as far south as Argentina and Uruguay, but a single species, G. crassilabris is from Panama. They are found in a wide range of freshwater habitats. They are part of a group popularly known as eartheaters and mostly feed by picking up mouthfuls of sediment to sift out food items such as invertebrates, plant material and detritus. The largest species reach up to in standard length. They are mostly kept in aquariums.
As an invasive species
Geophagus âÂÂsurinamensisâ was an invasive species in Malaysia, recently found in Putrajaya, the populations are now controlled by giant snakeheads.
Taxonomy
Some cichlids previously included in this genus have been reallocated to Biotodoma, Gymnogeophagus or Satanoperca. Even with these as separate genera, Geophagus is currently polyphyletic and in need of further taxonomic revision. There are three main groups:
- Geophagus sensu stricto are mostly relatively peaceful, often have long fin extensions and are native to the Amazon, Orinoco and ParnaÃÂba basins, as well as rivers of the Guianas. This group can be divided into two subgroups: The first is the G. surinamensis complex, which includes most species (fish in the aquarium trade often are identified as G. surinamensis itself, but they are typically other members of this complex.) The second subgroup contains G. argyrostictus, G. gottwaldi, G. grammepareius, G. harreri and G. taeniopareius, which are somewhat less peaceful and can be separated from the G. surinamensis complex by their dark stripe below the eye (however, this feature is shared with the next group).
- G. brasiliensis complex (including G. diamantinensis, G. iporangensis, G. itapicuruensis, G. multiocellus, G. obscurus, G. rufomarginatus and G. santosi) are more robust and aggressive species found in river basins of eastern and southeastern Brazil, Uruguay and northeastern Argentina.
- G. steindachneri complex (including G. crassilabris and G. pellegrini and undescribed species entering the aquarium trade from Colombia) found west of the Andes in northern and western Colombia, northwestern Venezuela and Panama where adult males develop a distinct, bulbous red forehead.
Species
There are currently 33 recognized species in this genus.
- Geophagus abalios <small>López-Fernández & Taphorn, 2004</small>
- Geophagus altifrons <small>Heckel, 1840</small>
- Geophagus argyrostictus <small>S. O. Kullander, 1991</small>
- Geophagus brachybranchus <small>S. O. Kullander & Nijssen, 1989</small>
- Geophagus brasiliensis <small>(Quoy & Gaimard, 1824)</small> (Pearl cichlid)
- Geophagus brokopondo <small>S. O. Kullander & Nijssen, 1989</small>
- Geophagus camopiensis <small>Pellegrin, 1903</small> (Oyapock eartheater)
- Geophagus crassilabris <small>Steindachner, 1876</small> (Panamanian eartheater)
- Geophagus crocatus <small>Hauser & López-Fernández, 2013</small>
- Geophagus diamantinensis <small>Mattos, W. J. E. M. Costa & A. C. A. Santos, 2015</small>
- Geophagus dicrozoster <small>López-Fernández & Taphorn, 2004</small>
- Geophagus gottwaldi <small>I. Schindler & Staeck, 2006</small>
- Geophagus grammepareius <small>S. O. Kullander & Taphorn, 1992</small>
- Geophagus harreri <small>J. P. Gosse, 1976</small> (Maroni eartheater)
- Geophagus iporangensis <small>Haseman, 1911</small>
- Geophagus itapicuruensis <small>Haseman, 1911</small>
- Geophagus megasema <small>Heckel, 1840</small>
- Geophagus mirabilis <small>Deprá, S. O. Kullander, Pavanelli & da Graça, 2014</small>
- Geophagus multiocellus <small>Mattos & W. J. E. M. Costa, 2018</small>
- Geophagus neambi <small>P. H. L. Lucinda, C. A. S. de Lucena & Assis, 2010</small>
- Geophagus obscurus <small>(Castelnau, 1855)</small>
- Geophagus parnaibae <small>Staeck & I. Schindler, 2006</small>
- Geophagus pellegrini <small>Regan, 1912</small> (Yellowhump eartheater)
- Geophagus pirangaensis <small>Assis, Dergam, Cunha, Machado, Hrbek, Vicente, Queiroz et Henschel, 2026</small>
- Geophagus proximus <small>(Castelnau, 1855)</small>
- Geophagus pyrineusi <small>Deprá, Ohara & Silva, 2022</small>
- Geophagus pyrocephalus <small>Chuctaya et al., 2022</small>
- Geophagus rufomarginatus <small>Mattos & W. J. E. M. Costa, 2018</small>
- Geophagus santosi <small>Mattos & W. J. E. M. Costa, 2018</small>
- Geophagus steindachneri <small>C. H. Eigenmann & Hildebrand, 1922</small> (Redhump eartheater)
- Geophagus surinamensis <small>(Bloch, 1791)</small> (Red-striped eartheater)
- Geophagus sveni <small>P. H. F. Lucinda, C. A. S. de Lucena & Assis, 2010</small>
- Geophagus taeniopareius <small>S. O. Kullander & Royero-L., 1992</small>
- Geophagus winemilleri <small>López-Fernández & Taphorn, 2004</small>
References