Emarginula is a genus of small keyhole limpets, marine gastropod molluscs in the family Fissurellidae.
Description
The oval shell is obliquely conical with a backward-pointing recurved apex. There is a deep incision in the anterior margin. A distinct anal fasciole extends upwards from this incision. The surface is latticed, sculptured with radial ribs. There is no septum inside.
Radula
The radula has a dental formula of [many].1.4.1.4.1.[many], with a wide, rectangular, acuspate rachidian tooth. The various genera have a few dozen rows of teeth.
Species
According to the World Register of Marine Species (WoRMS), species within the genus Emarginula include:
- Emarginula adamsiana <small>Sowerby in Adams & Sowerby</small>, 1863
- Emarginula adriatica <small>Costa O.G., 1829</small>
- Emarginula agulhasensis <small>Thiele, 1925</small>
- Emarginula alba <small>Watson, 1897</small>
- Emarginula amyda <small>Shikama, 1962</small>
- Emarginula angusta <small>McLean, 1970</small>
- Emarginula annielangleitae <small>Poppe & Tagaro, 2020</small>
- Emarginula apolonia <small>Pacaud, 2015</small>
- â Emarginula auversiensis <small>Deshayes, 1861</small>
- Emarginula bajula <small>Hedley, 1913</small>
- Emarginula balicasagensis <small>Poppe & Tagaro, 2020</small>
- Emarginula bellula <small>A. Adams, 1852</small>
- Emarginula bicancellata <small>Montrouzier, 1860</small>
- Emarginula bonfittoi <small>Smriglio & Mariottini, 2001</small>
- Emarginula boucheti <small>Poppe & Tagaro, 2020</small>
- â Emarginula brebioni <small>Ceulemans, Van Dingenen & Landau, 2016</small>
- Emarginula camilla <small>Melvill & Standen, 1903</small>
- Emarginula candida <small>Adams, 1852</small>
- Emarginula capuloidea <small>G. Nevill & H. Nevill, 1869</small>
- â Emarginula carezi <small>de Laubrière, 1881</small>
- Emarginula choristes <small>Dall, 1925</small>
- Emarginula christiaensi <small>Piani, 1985</small>
- Emarginula circumalbum P<small>oppe & Tagaro, 2020</small>
- â Emarginula clathrata <small>Deshayes, 1824</small>
- â Emarginula clathrataeformis <small>Eichwald, 1830</small>
- Emarginula colipanoae <small>Poppe & Tagaro, 2020</small>
- Emarginula compta <small>Habe, 1953</small>
- Emarginula concinna <small>Adams, 1852</small>
- Emarginula connectens <small>Thiele, 1915</small>
- Emarginula connelli <small>Kilburn, 1978</small>
- Emarginula convexa <small>Hedley, 1907</small>
- â Emarginula costata <small>Lamarck, 1803</small>
- Emarginula costulata <small>Deshayes, 1863</small>
- Emarginula crassa <small>Sowerby, 1812</small>
- Emarginula crassicostata <small>Sowerby in Adams & Sowerby, 1863</small>
- Emarginula cucullata <small>A. Adams, 1852</small>
- Emarginula curvamen <small>Iredale, 1925</small>
- Emarginula curvata <small>Schepman, 1908</small>
- Emarginula cuvieri <small>Audouin, 1827</small> (species inquirenda)
- Emarginula decorata <small>Deshayes, 1863</small>
- â Emarginula delicatissima <small>Chapman & Gabriel, 1923</small>
- Emarginula delonguevilleae <small>Poppe & Tagaro, 2020</small>
- â Emarginula dennanti <small>Chapman & Gabriel, 1923</small>
- Emarginula devota <small>Thiele, 1915</small>
- Emarginula dictya <small>McLean, 1970</small>
- â Emarginula didactica <small>Ludbrook, 1956</small>
- â Emarginula dilatoria <small>Ludbrook, 1956</small>
- Emarginula dilecta <small>Adams, 1852</small>
- Emarginula divae<small> (van Aartsen & Carrozza, 1995)</small>
- Emarginula dubia <small>Schepman, 1908</small> (accepted > unreplaced junior homonym, non Emarginula dubia <small>Defrance, 1819</small>)
- Emarginula dujardini <small>Dollfus & Dautzenberg, 1886</small> âÂÂ
- â Emarginula espibosensis <small>Lozouet, 1999</small>
- â Emarginula exigua <small>Trautschold, 1866</small>
- Emarginula fenestrella <small>Deshayes, 1863</small>
- Emarginula fissura<small> (Linnaeus, 1758)</small> - slit limpet
- â Emarginula fossorensis <small>Lozouet, 1999</small>
- Emarginula foveolata <small>Schepman, 1908</small>
- Emarginula foveolata fugitai <small>Habe, 1953</small>
- â Emarginula foveolata <small>Gerasimov, 1955</small> (accepted > unreplaced junior homonym)
- Emarginula fragilis <small>Yokoyama, 1920</small>
- Emarginula fuliginea <small>Adams, 1851</small>
- Emarginula gabensis <small>(Gabriel, 1961)</small>
- Emarginula galericulata <small>A. Adams, 1852</small>
- â Emarginula galeriformis <small>Marwick, 1928</small>
- â Emarginula gigantea<small> Poppe, 2008</small>
- â Emarginula guerangeri <small>A. d'Orbigny, 1843</small>
- Emarginula gustavi <small>Poppe & Tagaro, 2020</small>
- Emarginula hataii <small>Habe, 1953</small>
- Emarginula hawaiiensis <small>Dall, 1895</small>
- â Emarginula haweraensis <small>Powell, 1931</small>
- Emarginula hosoyai <small>Habe, 1953</small>
- Emarginula huzardii <small>(Payraudeau, 1826)</small>
- Emarginula icosisculpta <small>Simone & Cunha, 2014</small>
- Emarginula imaizumi <small>Dall, 1926</small>
- Emarginula imella <small>Dall, 1926</small>
- Emarginula janae <small>Cossignani, 2020</small>
- Emarginula japonica <small>Sowerby in Adams & Sowerby, 1863</small>
- Emarginula jungcheni <small>K.-Y. Lai, 2019</small>
- â Emarginula kaiparica <small>Laws, 1939</small>
- â Emarginula kaiwarensis <small>Bandel, Gründel & P. A. Maxwell, 2000</small> âÂÂ
- â Emarginula karuna <small>Das, Bardhan & Lahiri, 1999</small>
- Emarginula kashimaensis <small>Shikama, 1962</small>
- â Emarginula komitica <small>Laws, 1939</small>
- Emarginula koon <small>Kilburn, 1978</small>
- â Emarginula ladowae <small>Eichman, 1955</small>
- Emarginula lata <small>J.R.C. Quoy & J.P. Gaimard, 1834</small> ((uncertain, status in recent literature not researched)
- Emarginula lentiginosa <small>D. G. Herbert, 2024</small>
- Emarginula liuzzii <small>Poppe & Tagaro, 2020</small>
- Emarginula longifissa <small>Sowerby, 1866</small>
- Emarginula lorenzoi <small>Giusti & Micali, 2019</small>
- Emarginula macclurgi <small>Kilburn, 1978</small>
- Emarginula mactanensis <small>Poppe & Tagaro, 2020</small>
- Emarginula maculata <small>Adams in Adams & Sowerby, 1863</small>
- Emarginula margarita <small>Poppe & Tagaro, 2020</small>
- â Emarginula mariae <small>Cossmann, 1888</small>
- â Emarginula maudensis <small>Chapman & Gabriel, 1923</small>
- Emarginula micans <small>Adams, 1852</small>
- Emarginula montrouzieri <small>Souverbie, 1872</small>
- Emarginula multisquamosa <small>Schepman, 1908</small>
- Emarginula multistriata <small>Jeffreys, 1882</small>
- Emarginula natalensis <small>Barnard, 1963</small>
- Emarginula nigromaculata <small>(Thiele, 1930)</small>
- Emarginula nodulicostata <small>D. G. Herbert, 2024</small>
- Emarginula nordica <small>Pérez Farfante, 1947</small> - northern emarginula
- â Emarginula oblonga <small>Sandberger, 1859</small>
- Emarginula obovata <small>Adams, 1852</small>
- Emarginula octaviana <small>Coen, 1939</small>
- Emarginula oppressa <small>Barnard, 1963</small>
- Emarginula paivana <small>(Crosse, 1867)</small>
- Emarginula papilionacea <small>G. Nevill & H. Nevill, 1869</small>
- Emarginula patula <small>Cotton, 1930</small>
- â Emarginula paucicostata <small>Laws, 1936</small>
- Emarginula paucipunctata <small>Schepman, 1908</small>
- â Emarginula pelagica <small>Passy, 1832</small>
- Emarginula philippinensis <small>Poppe & Tagaro, 2020</small>
- Emarginula phrixodes <small>Dall, 1927</small> - ruffled emarginula
- Emarginula pittensis <small>Marwick, 1928</small>
- Emarginula poppeorum <small>Romani & Crocetta, 2017</small>
- â Emarginula praesicula <small>Lozouet, 1999</small>
- Emarginula puncticulata <small>Adams, 1852</small>
- Emarginula punctulum <small>Piani, 1980</small>
- â Emarginula punctura <small>S. Wood, 1848</small>
- Emarginula pustula <small>Thiele in Küster, 1913</small>
- Emarginula quadrata <small>Poppe & Tagaro, 2020</small>
- â Emarginula radiocostata <small>Kiel & Bandel, 2000</small>
- Emarginula regia <small>Habe, 1953</small>
- Emarginula retecosa <small>Adams, 1851</small>
- Emarginula retrogyra <small>D. G. Herbert, 2024</small>
- Emarginula rollandi <small>P. Fischer, 1857</small> (uncertain: status in recent literature not researched)
- Emarginula rosea <small>Bell, 1824</small>
- Emarginula salebrosa <small>D. G. Herbert, 2024</small>
- â Emarginula sanctaecaatharine <small>Passy, 1832</small>
- Emarginula scabricostata <small>A. Adams, 1852</small>
- Emarginula scabriuscula <small>Adams, 1852</small>
- Emarginula sicula <small>Gray, 1825</small> - dagger emarginula
- Emarginula sinica <small>Lu, 1986</small>
- Emarginula solidula <small>Costa, 1829</small>
- Emarginula souverbiana <small>Pilsbry, 1890</small>
- Emarginula spinosa <small>Deshayes, 1863</small>
- â Emarginula squammata <small>Grateloup, 1836</small>
- â Emarginula striatissima <small>Traub, 1979</small>
- Emarginula striatula <small>Quoy and Gaimard, 1834</small> - dagger emarginula
- Emarginula sublaevis <small>Schepman, 1908</small>
- Emarginula subtilitexta <small>Verco, 1908</small>
- Emarginula superba <small>Hedley, 1906</small>
- Emarginula suspira <small>Simone & Cunha, 2014</small>
- Emarginula tenera <small>Locard, 1892</small>
- â Emarginula teneraeformis <small>Lozouet, 1999</small>
- Emarginula tenuicostata <small>Sowerby, 1863</small>
- Emarginula teramachii <small>Habe, 1953</small>
- Emarginula textilis <small>Gould, 1859 in 1859-61</small>
- Emarginula thomasi<small> Crosse, 1864</small>
- Emarginula thorektes <small>Kilburn, 1978</small>
- Emarginula tosaensis <small>Habe, 1953</small>
- â Emarginula transenna <small>Tenison Woods, 1877</small>
- Emarginula tuberculosa <small>Libassi, 1859</small> - tuberculate emarginula
- Emarginula undulata <small>Melvill & Standen, 1903</small>
- Emarginula velascoensis <small>Shasky, 1961</small>
- Emarginula velascoi <small>Rehder, 1980</small>
- Emarginula viridibrunneis <small>Poppe & Tagaro, 2020</small>
- Emarginula viridicana <small>Herbert & Kilburn, 1985</small>
- Emarginula viridis <small>Poppe & Tagaro, 2020</small>
- â Emarginula wannonensis <small>G. F. Harris, 1897</small>
- â Emarginula washingtoniana <small>Squires & Goedert, 1994</small>
- Emarginula xishaensis <small>Lu, 1986</small>
- Emarginula yangeorum <small>Poppe & Tagaro, 2020</small>
Synonyms
- Emarginula altilis Gould, 1859: synonym of Variemarginula punctata (A. Adams, 1852)
- Emarginula amitina Iredale, 1925: synonym of Emarginula dilecta A. Adams, 1852
- Emarginula angustata Thiele, 1915: synonym of Tugali decussata A. Adams, 1852
- Emarginula arabica A. Adams, 1852: synonym of Octomarginula arabica (A. Adams, 1852)
- Emarginula arconatii Issel, 1869: synonym of Montfortista panhi (Quoy & Gaimard, 1834)
- Emarginula australis Quoy & Gaimard, 1834: synonym of Montfortia subemarginata (Blainville, 1819)
- Emarginula bella Gabb, 1865: synonym of Scelidotoma bella (Gabb, 1865)
- Emarginula biangulata Sowerby, 1901: synonym of Emarginella biangulata (G. B. Sowerby III, 1901)
- â Emarginula blainvillii <small>Defrance, 1825</small>: synonym of â Rimula blainvillii <small>(Defrance, 1825)</small>
- Emarginula brevirimata Deshayes, 1862: synonym of Montfortula brevirimata (Deshayes, 1862)
- Emarginula clathrata A. Adams & Reeve, 1850: synonym of Montfortista panhi (Quoy & Gaimard, 1834)
- Emarginula clathrata Pease, 1863: synonym of Emarginula dilecta A. Adams, 1852
- Emarginula conica Lamarck, 1801: synonym of Emarginula fissura (Linnaeus, 1758)
- Emarginula conica Schumacher, 1817 non Lamarck, 1801: synonym of Emarginula rosea Bell, 1824
- Emarginula dentigera Heilprin, 1889: synonym of Hemimarginula dentigera (Heilprin, 1889)
- Emarginula elata <small>Locard, 1898</small>: synonym of Emarginula christiaensi <small>Piani, 1985</small>
- Emarginula elongata O. G. Costa, 1829: synonym of Emarginula octaviana Coen, 1939
- Emarginula eximia <small>A. Adams, 1852</small>: synonym of Emarginella eximia <small>(A. Adams, 1852)</small>
- Emarginula flindersi Cotton, 1930: synonym of Emarginula convexa Hedley, 1907
- Emarginula fuliginea Kuroda, 1941: synonym of Variemarginula variegata (A. Adams, 1852)
- Emarginula guernei Dautzenberg & Fischer, 1896: synonym of Emarginula tuberculosa Libassi, 1859
- Emarginula hedleyi Thiele, 1915: synonym of Emarginula candida A. Adams, 1852
- Emarginula incisula(sic): synonym of Emarginella incisura <small>(A. Adams, 1852)</small>
- Emarginula incisura <small>A. Adams, 1852</small>: synonym of Emarginella incisura <small>(A. Adams, 1852)</small>
- Emarginula kimberti Cotton, 1930: synonym of Laeviemarginula kimberti (Cotton, 1930)
- Emarginula modesta H. Adams, 1872: synonym of Hemimarginula modesta (H. Adams, 1872)
- Emarginula muelleri Forbes & Hanley, 1849: synonym of Emarginula fissura (Linnaeus, 1758)
- Emarginula ostheimerae Abbott, 1958: synonym of Octomarginula ostheimerae (Abbott, 1958)
- Emarginula panhi Quoy & Gaimard, 1834: synonym of Montfortista panhi (Quoy & Gaimard, 1834)
- Emarginula papillosa <small>Risso 1826</small>: synonym of Emarginula huzardii <small>Payraudeau, 1826</small>
- Emarginula parmophoidea Quoy & Gaimard, 1834: synonym of Tugali elegans Gray, 1843
- Emarginula peasei Thiele, 1915: synonym of Emarginula dilecta A. Adams, 1852
- Emarginula phrygium <small>Herbert & Kilburn, 1985</small>: synonym of Agariste phrygium <small>(Herbert & Kilburn, 1986)</small> (original combination)
- Emarginula picta Dunker, 1860: synonym of Montfortula picta (Dunker, 1860)
- Emarginula pileata Gould, 1859: synonym of Variemarginula punctata (A. Adams, 1852)
- Emarginula pileum Heilprin, 1889: synonym of Hemimarginula pileum (Heilprin, 1889)
- Emarginula planulata <small>A. Adams, 1852</small>: synonym of Emarginella planulata <small>(A. Adams, 1852)</small>
- Emarginula polygonalis A. Adams, 1852: synonym of Hemitoma polygonalis (A. Adams, 1852)
- Emarginula pulchreclathrata Tomlin, 1932: synonym of Emarginula agulhasensis Thiele, 1925
- Emarginula pumila A. Adams, 1852: synonym of Hemimarginula pumila (A. Adams, 1852)
- Emarginula punctata A. Adams, 1852: synonym of Variemarginula punctata (A. Adams, 1852)
- Emarginula reticulata Sowerby, 1813: synonym of Emarginula fissura (Linnaeus, 1758)
- Emarginula rugosa Quoy & Gaimard, 1834: synonym of Montfortula rugosa (Quoy & Gaimard, 1834)
- Emarginula scutellata Deshayes, 1863: synonym of Octomarginula scutellata (Deshayes, 1863)
- Emarginula sibogae Schepman, 1908: synonym of Emarginella sibogae (Schepman, 1908)
- Emarginula simpla Christiaens, 1987: synonym of Hemimarginula simpla (Christiaens, 1987)
- Emarginula stellata A. Adams, 1852: synonym of Montfortula rugosa (Quoy & Gaimard, 1834)
- Emarginula subclathrata Pilsbry, 1890: synonym of Emarginula dilecta A. Adams, 1852
- Emarginula subemarginata Blainville, 1819: synonym of Montfortia subemarginata (Blainville, 1819)
- Emarginula subrugosa Thiele, 1916: synonym of Hemimarginula subrugosa (Thiele, 1916)
- Emarginula sulcifera A. Adams, 1852: synonym of Montfortulana sulcifera (A. Adams, 1852)
- Emarginula tricarinata Pilsbry, 1890: synonym of Montfortista panhi (Quoy & Gaimard, 1834)
- Emarginula tricostata G.B. Sowerby I, 1823: synonym of Hemitoma octoradiata (Gmelin, 1791)
- Emarginula vadososinuata Yokoyama, 1922: synonym of Tugalina vadososinuata (Yokoyama, 1922)
- Emarginula vadum Barnard, 1963: synonym of Emarginula undulata Melvill & Standen, 1903
- Emarginula variegata A. Adams, 1852: synonym of Variemarginula variegata (A. Adams, 1852)
References
- Powell A W B, New Zealand Mollusca, William Collins Publishers Ltd, Auckland, New Zealand 1979
- Piani P. (1985). Revisione del genere Emarginula Lamarck, 1801 in Mediterraneo. Lavori, SocietÃÂ Italiana di Malacologia 21: 193-238
- Gofas, S.; Le Renard, J.; Bouchet, P. (2001). Mollusca, in: Costello, M.J. et al. (Ed.) (2001). European register of marine species: a check-list of the marine species in Europe and a bibliography of guides to their identification. Collection Patrimoines Naturels, 50: pp. 180âÂÂ213
- Rolán E., 2005. Malacological Fauna From The Cape Verde Archipelago. Part 1, Polyplacophora and Gastropoda.
- Spencer, H.; Marshall. B. (2009). All Mollusca except Opisthobranchia. In: Gordon, D. (Ed.) (2009). New Zealand Inventory of Biodiversity. Volume One: Kingdom Animalia. 584 pp