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Dactylioceras

Dactylioceras was a widespread genus of ammonites from the Lower Jurassic period, approximately 180 million years ago (mya).Like many other ammonites, the genus Dactylioceras is extremely important in biostratigraphy, being a key index fossil for identifying their region of the Jurassic. It had a nearly cosmopolitan distribution during the time it existed.

Etymology

The name Dactylioceras comes from the Greek dactyl, meaning “finger”, and keras, meaning "horn" and refers to the shell's branching ribs.

Description

Dactylioceras are generally small, averaging in diameter. They have a strong, ribbed shell. The ribs are slightly inclined forward, running over the outer edge, and either simple or forking at outer end. Though they eventually died out 180 mya, their style of ribbing was copied by numerous subsequent ammonite genera until the whole group became extinct 66 million years ago with the dinosaurs.

Ecology

Dactylioceras probably lived by scavenging on the sea floor. Mass mortality specimens of Dactylioceras are common, and perhaps suggest that these ammonites may often have died shortly after spawning. The dead shells were probably gently washed up into a shell bank on the margins of the Lower Jurassic seas. Flow tank experiments show that Dactylioceras was probably a slow swimmer. It was likely prey for larger marine animals at the time including several belemnites.

Distribution

Dactylioceras has been collected from almost every continent, and was one of the most successful ammonite lineages ever. They are abundant throughout Europe, with exceptionally fine specimens found in England and Germany.

Subgenera

Currently, 3 or 4 subgenera are considered to be valid. Number differs on authors opinions.

  • D. (Dactylioceras) <small>Hyatt, 1867</small>: type species: Ammonites communis <small>Sowerby, 1815</small>
  • D. (Orthodactylites) <small>Buckman, 1926</small>: type species: Dactylioceras directus <small>Buckman, 1926</small>
  • D. (Iranodactylites) <small>Repin, 2000</small>: type species: Dactylioceras (Iranodactylites) ketevanae <small>Repin, 2000</small>
  • D. (Eodactylites) <small>Schmidt-Effing, 1972</small>: type species: Dactylioceras pseudocommune <small>Fucini, 1935</small>. This subgenus is not recognized by some authors, while some other prefer to maintain it.

Species

Species within the genus Dactylioceras include:

  • D. (O.) aequistriatum <small>Zieten, 1830</small>
  • D. (D.) alpestre <small>Wiedenmayer, 1980</small>
  • D. (D.) amplum <small>Dagis, 1968</small>
  • D. (O.) andaluciensis <small>Jiménes & Rivas, 1991</small>
  • D. (O.) anguiforme <small>Buckman, 1928</small>
  • D. (O.) anguinum <small>Reinecke, 1818</small>
  • D. (D.) annuliferum <small>Simpson, 1855</small>
  • D. (D.) arcus <small>Buckman, 1926</small>
  • D. (D.) athleticum <small>Simpson, 1855</small>
  • D. (D.) attenuatum <small>Simpson, 1855</small>
  • D. (O.) chilense <small> Hillebrandt and Schmidt-Effing, 1981</small>
  • D. (O.) clevelandicum <small>Howarth, 1973</small>
  • D. (D.) commune <small>Sowerby, 1815</small>
  • D. (D.) comptum <small>Dagis, 1968</small>
  • D. (D.) consimile <small>Buckman, 1926</small>
  • D. (D.) crassescens <small>Simpson, 1855</small>
  • D. (O.) crassifactum <small>Simpson, 1855</small>
  • D. (O.) crassiusculosum <small>Buckman, 1912</small>
  • D. (D.) crassiusculum <small>Simpson, 1855</small>
  • D. (D.) crassulum <small>Buckman, 1921</small>
  • D. (D.) crosbeyi <small>Simpson, 1843</small>
  • D. (O.) directum <small>Buckman, 1926</small>
  • D. (O.) ernsti <small>Lehmann, 1968</small>
  • D. (D.) gracile <small>Simpson, 1843</small>
  • D. (O.) helianthoides <small>Yokoyama, 1904</small>
  • D. (O.) hispansum <small>Schmidt-Effing, 1972</small>
  • D. (O.) hoelderi <small>Hillebrandt & Schmidt-Effing, 1981</small>
  • D. (D.) holandrei <small>d'Orbigny, 1845</small>
  • D. (O.) ketevanae <small>Repin, 2000</small>
  • D. (O.) kanense <small>McLearn, 1930</small>
  • D. (I.) ketevanae <small>Repin, 2000</small>
  • D. (D.) laticostatum <small>Bardin et al., 2014</small>
  • D. (O.) marioni <small>Lissajous, 1906</small>
  • D. (E.) mirabile <small>Fucini, 1935</small>
  • D. (D.) mite <small>Buckman, 1927 </small>
  • D. (D.) peloritanum <small>Fucini, 1935</small>
  • D. (D.) percostatum <small>Fucini, 1935</small>
  • D. (D.) perplicatum <small>Fucini, 1935</small>
  • D. (E.) polymorphum <small>Fucini, 1935</small>
  • D. (D.) praepositum <small>Buckman, 1927</small>
  • D. (E.) pseudocommune <small>Fucini, 1935</small>
  • D. (D.) pseudocrassoides <small>Maubeuge, 1957</small>
  • D. (O.) sapunovi <small>Repin, 2000</small>
  • D. (O.) semiannulatum <small>Howarth, 1978</small>
  • D. (O.) semicelatum <small>Simpson, 1843</small>
  • D. (E.) simplex <small>Fucini, 1935</small>
  • D. (D.) stresherense <small>Sapunov, 1963</small>
  • D. (D.) subholandrei <small>Fucini, 1935</small>
  • D. (D.) suntarense <small>Krimholz, 1957</small>
  • D. (D.) tardosensis <small>Kovács, 2014</small>
  • D. (D.) tauromenense <small>Fucini, 1935</small>
  • D. (D.) temperatum <small>Buckman, 1927</small>
  • D. (O.) tenuicostatum <small>Young & Bird, 1822</small>
  • D. (O.) toxophorum <small>Buckman, 1926</small>
  • D. (D.) triangulum <small>Fischer, 1966</small>
  • D. (D.) vermis <small>Simpson, 1855</small>
  • D. (O.) wunnenbergi <small>Hoffmann, 1968</small>

Dactylioceras commune has shells reaching a diameter of . Usually the average diameter reach about , while the average width is .

In these small but strong shells ribs run straight or are slightly convex across the venter. These ribs are quite coarse on the outer whorls and finer on the inner whorls. The whorl section is as round as a circle.

Fossils of this species have been found in Lower Jurassic, Toarcian age of Canada, France, Italy, Russia, Serbia and Montenegro, United Kingdom and United States.

References