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Cynipoidea

The Cynipoidea are a moderate-sized hymenopteran superfamily that presently includes seven extant families and three extinct families, though others have been recognized in the past. The most familiar members of the group are phytophagous, especially as gall-formers, though the actual majority of included species are parasitoids or hyperparasitoids. They are typically glossy, dark, smooth wasps with somewhat compressed bodies and somewhat reduced wing venation. It is common for various metasomal segments to be fused in various ways (often diagnostic for families or subfamilies), and the petiole is very short, when present .

With the exception of the Cynipidae (the gall wasps), it is a poorly known group as a whole, though there are nearly 3000 known species in total, and a great many species are still undescribed, mostly in the Figitidae. Each of the constituent families differs in biology, though life histories of one of the families (Liopteridae) are still largely unknown. In July 2020, an identification key for the superfamily was published, enabling identification to the family/tribe level. Several groups formerly included in Cynipidae were elevated to family status in 2023.

Classification

:: Family Protimaspidae <small>Liu & Engel, 2007</small>
:: Family Stolamissidae <small>Liu & Engel, 2007</small>
:: Family Gerocynipidae <small>Liu & Engel, 2007</small>
:: Family Austrocynipidae <small>Riek, 1971</small>
:: Family Paraulacidae <small>Nieves-Aldrey & Liljeblad, 2009</small>
:: Family Diplolepididae <small>Latreille, 1802</small>
:::: Subfamily Diplolepidinae <small>Latreille, 1802</small>
:::: Subfamily Pediaspinae <small>Ashmead, 1903</small>
:: Family Ibaliidae <small>Thomson, 1862</small>
:::: Subfamily Archaeibaliinae <small>Liu & Engel, 2010</small>
:::: Subfamily Ibaliinae <small>Thomson, 1862</small>
:: Family Liopteridae <small>Ashmead, 1895</small>
:::: Subfamily Proliopterinae <small>Liu & Engel, 2007</small>
:::: Subfamily Goeraniinae <small>Liu & Engel, 2007</small>
:::: Subfamily Mayrellinae <small>Hedicke, 1922</small>
:::: Subfamily Dallatorrellinae <small>Kieffer, 1911</small>
:::: Subfamily Oberthuerellinae <small>Hedicke, 1903</small>
:::: Subfamily Liopterinae <small>Ashmead, 1895</small>
:: Family Figitidae <small>Thomson, 1862</small>
:::: Subfamily Rasnicynipinae <small>Kozlov, 1996</small>
:::: Subfamily Palaeocynipinae <small>Kozlov, 1995</small>
:::: Subfamily Charipinae <small>Dalla Torre & Kieffer, 1910</small>
:::: Subfamily Emargininae <small>Kovalev, 1994</small>
:::: Subfamily Eucoilinae <small>Thomson, 1862</small>
::::: Tribe Diglyphosematini <small>Belizin, 1961</small>
::::: Tribe Eucoilini <small>Thomson, 1862</small>
::::: Tribe Ganaspini <small>Belizin, 1961</small>
::::: Tribe Kleidotomini <small>Hellén, 1960</small>
::::: Tribe Trichoplastini <small>Kovalev, 1989</small>
::::: Tribe Zaeucoilini <small>Buffington, 2009</small>
:::: Subfamily Figitinae <small>Thomson, 1862</small>
:::: Subfamily Mikeiinae <small>Paretas-Martinez & Pujade-Villar, 2011</small>
:::: Subfamily Aspicerinae <small>Dalla Torre & Kieffer, 1910</small>
:::: Subfamily Anacharitinae <small>Thomson, 1862</small>
:::: Subfamily Pycnostigminae <small>Cameron, 1905</small>
:::: Subfamily Thrasorinae <small>Kovalev, 1994</small>
:::: Subfamily Plectocynipinae <small>Ros-Farré & Pujade-Villar, 2007</small>
:::: Subfamily Euceroptresinae <small>Buffington & Liljeblad, 2008</small>
:::: Subfamily Parnipinae <small>Ronquist & Nieves-Aldrey, 2001</small>
:: Family Cynipidae <small>Latreille, 1802</small>
:::: Subfamily Hodiernocynipinae <small>Kovalev, 1994</small>
:::: Subfamily Cynipinae <small>Latreille, 1802</small>
::::: Tribe Aulacideini <small>Nieves-Aldrey & Ronquist, 2015</small>
::::: Tribe Aylacini <small>Ashmead, 1903</small>
::::: Tribe Ceroptresini <small>Nieves-Aldrey, Nylander, & Ronquist, 2015</small>
::::: Tribe Cynipini <small>Latreille, 1802</small>
::::: Tribe Diastrophini <small>Nieves-Aldrey, Nylander, & Ronquist, 2015</small>
::::: Tribe Eschatocerini <small>Ashmead, 1903</small>
::::: Tribe Phanacidini <small>Nieves-Aldrey, Nylander, & Ronquist, 2015</small>
::::: Tribe Qwaqwaiini <small>Liljeblad, Nieves-Aldrey, & Melika, 2011</small>
::::: Tribe Synergini <small>Ashmead, 1896</small>

References