Cyclophoroidea is a superfamily of land snails with an operculum, terrestrial gastropods within the order Architaenioglossa, that belongs to the subclass Caenogastropoda. Approximately 3,675 cyclophoroid species were known as of 2024, making them the second-most diverse clade of land snails after Stylommatophora.
These terrestrial gastropods have lost the ctenidium (comb-like respiratory apparatus) and osphradium, and the pallial cavity has been modified as a lung.
Cyclophoroids have detritivorous or herbivorous diets.
Cyclophoroids probably originated in the Late Jurassic or Early Cretaceous based on molecular clock estimates. Several fossil species are known from Burmese amber that dates to the mid-Cretaceous, approximately 99 million years ago.
Taxonomy
According to the Taxonomy of the Gastropoda (Bouchet & Rocroi, 2005), this superfamily consists of the following families:
- Family Aciculidae <small>Gray, 1850</small>
- Family Alycaeidae <small>W. T. Blanford, </small>
- Genus â Carinomphalus <small>W. Yü, 1974</small>
- Family Cochlostomatidae <small>Kobelt, 1902</small>
- Family Craspedopomatidae <small>Kobelt & Möllendorff, 1898</small>
- Family Cyclophoridae <small>Gray, 1847</small>
- Subfamily Cyclophorinae <small>Gray, 1847</small>
- Tribe Caspicyclotini <small>Wenz, 1938</small>
- Tribe Cyathopomatini <small>Kobelt & Möllendorff, 1897</small>
- Tribe Cyclophorini <small>Gray, 1847</small>
- Tribe Cyclotini <small>Pfeiffer, 1853</small>
- Tribe Pterocyclini <small>Kobelt & Möllendorff, 1897</small>
- Subfamily Spirostomatinae <small>Tielecke, 1940</small>
- Family Diplommatinidae <small>Pfeiffer, 1857</small>
- Subfamily Diplommatininae <small>Pfeiffer, 1857</small>
- Genus â Diplommoptychia <small>Maillard, 1884</small>
- â Family Ferussinidae <small>Wenz, 1923 (1915)</small>: accepted as â Ferussininae <small>Wenz, 1923 (1915)</small>
- Genus â Loriolina <small>Huckriede, </small>
- Genus â Maillardinus <small>, 1991 </small>
- Family Maizaniidae <small>Tielecke, 1940</small>
- Family Megalomastomatidae <small>Blanford, 1864</small>
- Family Neocyclotidae <small>Kobelt & Möllendorff, 1897</small>
- Subfamily Amphicyclotinae <small>Kobelt & Möllendorff, 1897</small>
- Family Pupinidae <small>Pfeiffer, 1853</small>
- Subfamily Liareinae <small>Powell, 1946</small>
- Subfamily Pupinellinae <small>Kobelt, 1902</small>
- Subfamily Pupininae <small>Pfeiffer, 1853</small>
Families brought into synonymy:
- Acmeidae <small>Pollonera, 1905</small>: synonym of Aciculidae <small>Gray, 1850</small>
- Alycaeidae <small>Blanford, 1864</small>: synonym of Cyclophoridae <small>Gray, 1847</small>
- Bolaniidae <small>Wenz, 1915</small>: synonym of Craspedopomatidae <small>Kobelt & Möllendorff, 1898</small>
- Dicristidae <small>Golikov & Starobogatov, 1975</small>: synonym of Neocyclotidae <small>Kobelt & Möllendorff, 1897</small>
- Lagocheilidae <small>Stoliczka, 1872</small>: synonym of Cyclophoroidea <small>Gray, 1847</small>
- â Strophostomatidae <small>Wenz, 1915</small>: synonym of â Ferussinidae <small>Wenz, 1923 (1915)</small>
References