Cryptinae

Cryptinae is a subfamily of wasps in the family Ichneumonidae. The family has also been called Gelinae, Hemitelinae, and Phygadeuontinae by various authorities, though the Phygadeuontinae have since been elevated to a separate subfamily.

Description

Species are distributed worldwide. A 5-sided areolet, a short sternaulus, and a puffy face with a convex clypeus are diagnostic characters for most members of this large subfamily.

Taxonomy

Gelinae was the name used by H. K. Townes in 1969. Fitton and Gauld's review of ichneumonid families synonymized Gelinae under Hemitelinae in 1976 and later as Phygadeuontinae in 1978. However, due to the decision of International Code of Zoological Nomenclature's Opinion 1715 in 1994 to conserve the status of the genus Cryptus <small>Fabricius, 1804</small>, both Gelinae and Phygadeuontinae became junior synonyms for Cryptinae. Santos later restricted the family in 2017 to only include the tribes Aptesini and Cryptini, forming the separate families Claseinae and Phygadeuontinae for the remaining tribes of Claseini (1 genus) and Phygadeuontini (123 genera), respectively.

There are about 273 genera in Cryptinae between the tribes of Aptesini and Cryptini.

Classification

The following classification of Phygadeuontini is based on Wahl (2014):

Tribe Cryptini

• Subtribe Agrothereutina
• Agrothereutes <small>Förster, 1850 </small> — Holarctic
• Amauromorpha <small>Ashmead, 1905</small> — Oriental
• Apsilops <small>Förster, 1869</small> — Holarctic
• Aritranis <small>Förster, 1869</small> — Holarctic
• Gambrus <small>Förster, 1869</small> — Holarctic
• Hidryta <small>Förster, 1869</small> — Holarctic
• Idiolispa <small>Förster, 1869</small> — Holarctic
• Mesostenidea <small>Viereck, 1914</small> — Palearctic
• Pycnocryptodes <small>Aubert, 1971</small> — Palearctic
• Pycnocryptus <small>Thomson, 1873</small> — Holarctic
• Thrybius <small>Townes, 1965</small> — Palearctic
• Trychosis <small>Förster, 1869</small> — Holarctic, Oriental
• Subtribe Ateleutina
• Ateleute <small>Förster, 1869</small> — worldwide
• Tamaulipeca <small>Kasparyan & Hernandez, 2001</small> — Neotropical
• Subtribe Baryceratina
• Baryceros <small>Gravenhorst, 1829</small> — Nearctic, Neotropical
• Brussinocryptus <small>Pagliano & Scarramozino, 1990</small> — Oriental, Palearctic
• Buysmania <small>Cheesman, 1941</small> — Oriental
• Calaminus <small>Townes, 1965</small> — Palearctic
• Ceratomansa <small>Cushman, 1922</small> — Australian
• Chlorocryptus <small>Cameron, 1903</small> — Oriental, Palearctic
• Coccygodes <small>Saussure, 1892</small> — Ethiopian
• Jonathania <small>Gupta, 1987</small> — Oriental
• Lamprocryptidea <small>Viereck, 1913</small> — Neotropical
• Paragambrus <small>Uchida, 1936</small> — Palearctic
• Prionacis <small>Townes, 1970a</small> — Oriental
• Whymperia <small>Cameron, 1903</small> — Nearctic, Neotropical
• Subtribe Ceratocryptina
• Acromia <small>Townes, 1961b</small> — Oriental
• Afretha <small>Seyrig, 1952</small> — Ethiopian
• Aprix <small>Townes, 1961b</small> — Australian
• Ceratocryptus <small>Cameron, 1903</small> — Ethiopian, Oriental
• Ceratodolius <small>Seyrig, 1952</small> — Ethiopian
• Chamula <small>Townes, 1962</small> — Nearctic, Neotropical
• Cheesmanella <small>Ozdikmen, 2005</small> — Oriental
• Cremnocryptus <small>Cushman, 1945</small> — Australian, Oriental
• Ecaepomia <small>Wang, 2001</small> — Oriental
• Fitatsia <small>Seyrig, 1952</small> — Ethiopian, Oriental
• Lipoprion <small>Townes, 1970a</small> — Oriental
• Lorio <small>Cheesman, 1936</small> — Oriental
• Malaycromia <small>Gupta, 1999</small> — Oriental
• Neaprix <small>Gauld, 1984</small> — Australian
• Nematocryptus <small>Roman, 1910</small> — Ethiopian, Oriental
• Piasites <small>Seyrig, 1952</small> — Ethiopian
• Rambites <small>Seyrig, 1952</small> — Ethiopian
• Silsila <small>Cameron, 1903</small> — Oriental
• Thelodon <small>Townes, 1961b</small> — Australian, Oriental
• Trachyglutus <small>Townes, 1970a</small> — Neotropical
• Wuda <small>Cheesman, 1936</small> — Oriental
• Subtribe Coesulina
• Coesula <small>Cameron, 1905</small> — Oriental
• Subtribe Cryptina

• Aeglocryptus <small>Porter, 1987</small> — Neotropical
• Aeliopotes <small>Porter, 1985</small> — Neotropical
• Aglaodina <small>Porter, 1987</small> — Neotropical
• Araucacis <small>Porter, 1987</small> — Neotropical
• Biconus <small>Townes, 1970a</small> — Neotropical
• Buathra <small>Cameron, 1903</small> — Holarctic, Oriental
• Caenocryptoides <small>Uchida, 1936</small> — Oriental, Palearctic
• Caenocryptus <small>Thomson, 1873</small> — Holarctic
• Caenopelte <small>Porter, 1967</small> — Neotropical
• Camera <small>Townes, 1962</small> — Nearctic, Neotropical
• Chilecryptus <small>Porter, 1987</small> — Neotropical
• Chromocryptus <small>Ashmead, 1900</small> — Neotropical
• Compsocryptus <small>Ashmead, 1900</small> — Nearctic, Neotropical
• Cosmiocryptus <small>Cameron, 1902</small> — Neotropical (Porter, 1987)
• Cryptopteryx <small>Ashmead, 1900</small> — Neotropical
• Cryptus <small>Fabricius, 1804</small> — Ethiopian, Holarctic, Oriental
• Cyanodolius <small>Seyrig, 1952</small> — Ethiopian
• Cyanopelor <small>Townes, 1973</small> — Ethiopian
• Cyclaulus <small>Townes, 1970a</small> — Neotropical
• Dihelus <small>Townes, 1970a</small> — Oriental, Palearctic
• Diplohimas <small>Townes, 1970a</small> — Neotropical
• Distictus <small>Townes, 1966</small> — Neotropical
• Dochmidium <small>Porter, 1967</small> — Neotropical (Porter, 1987)
• Dotocryptus <small>Brèthes, (1918) 1919</small> — Neotropical
• Enclisis <small>Townes, 1970a</small> — Palearctic
• Etha <small>Cameron, 1903</small> — Oriental
• Gessia <small>Townes, 1973</small> — Ethiopian
• Glabridorsum <small>Townes, 1970a</small> — Australian, Oriental, Palearctic
• Gyropyga <small>Townes, 1970a</small> — Oriental, Palearctic
• Hedycryptus <small>Cameron, 1903</small> — Oriental, Palearctic
• Hypsanacis <small>Porter, 1987</small> — Neotropical
• Ischnus <small>Gravenhorst, 1829</small> — worldwide
• Itamuton <small>Porter, 1987</small> — Neotropical
• Joppidium <small>Cresson, 1872</small> — Nearctic, Neotropical
• Lanugo <small>Townes, 1962</small> — Nearctic, Neotropical
• Leptarthron <small>Townes, 1970a</small> — Neotropical
• Meringopus <small>Förster, 1869</small> — Holarctic
• Mesophragis <small>Seyrig, 1952</small> — Ethiopian
• Monothela <small>Townes, 1970a</small> — Neotropical
• Myrmecacis <small>Porter, 1987</small> — Neotropical
• Myrmeleonostenus <small>Uchida, 1936</small> — Australian, Oriental, Palearctic
• Nebostenus <small>Gauld, 1984</small> — Australian
• Nelophia <small>Porter, 1967</small> — Neotropical
• Neocryptopteryx <small>Blanchard, 1947</small> — Neotropical (Porter, 1987)
• Neodontocryptus <small>Uchida, 1940</small> — Oriental
• Nippocryptus <small>Uchida, 1936</small> — Oriental, Palearctic
• Nothischnus <small>Porter, 1967</small> — Neotropical (Porter, 1987)
• Odontocryptus <small>Saussure, 1892</small> — Ethiopian
• Oecetiplex <small>Porter, 1987</small> — Neotropica
• Palmerella <small>Cameron, 1908</small> — Oriental, Palearctic
• Periplasma <small>Porter, 1967</small> — Neotropical (Porter, 1987)
• Phycitiplex <small>Porter, 1987</small> — Neotropical
• Picrocryptoides <small>Porter, 1965</small> — Neotropical
• Reptatrix <small>Townes, 1962</small> — Nearctic
• Rhynchocryptus <small>Cameron, 1905</small> — Ethiopian (Townes & Townes, 1973)
• Sciocryptus <small>Porter, 1987</small> — Neotropical
• Synechocryptus <small>Schmiedeknecht, 1904</small> — Ethiopian, Palearctic
• Trachysphyrus <small>Haliday, 1836</small> — Neotropical
• Tricentrum <small>Townes, 1970a</small> — Neotropical
• Trihapsis <small>Townes, 1970a</small> — Neotropical
• Xiphonychidion <small>Porter, 1967</small> — Neotropical (Porter, 1987)
• Xylacis <small>Porter, 1987</small> — Neotropical
• Xylophrurus <small>Förster, 1869</small> — Holarctic
• Xylostenus <small>Gauld, 1984</small> — Australian
• Zonocryptus <small>Ashmead, 1900</small> — Ethiopian
• Subtribe Gabuniina

• Agonocryptus <small>Cushman, 1929</small> — Nearctic, Neotropical
• Ahilya <small>Gupta & Gupta, 1985</small> — Oriental
• Amrapalia <small>Gupta & Jonathan, 1970</small> — Oriental
• Anepomias <small>Seyrig, 1952</small> — Ethiopian
• Apocryptus <small>Uchida, 1932</small> — Oriental
• Arhytis <small>Townes, 1970a</small> — Australian, Oriental
• Cestrus <small>Townes, 1966</small> — Neotropical
• Cryptohelcostizus <small>Cushman, 1919</small> — Nearctic
• Dagathia <small>Cameron, 1903</small> — Oriental
• Digonocryptus <small>Viereck, 1913</small> — Neotropical
• Eurycryptus <small>Cameron, 1901</small> — Australian, Ethiopian, Oriental
• Fenixia <small>Aguiar, 2005</small> — Neotropical
• Fortipalpa <small>Kasparyan & Ruiz, 2007</small> — Neotropical
• Gabunia <small>Kriechbaumer, 1895</small> — Ethiopian
• Gerdius <small>Townes, 1970a</small> — Ethiopian
• Hackerocryptus <small>Gauld, 1984</small> — Australian
• Hadrocryptus <small>Cameron, 1903</small> — Oriental
• Kemalia <small>Koçak, 2009</small> — Oriental
• Kriegeria <small>Ashmead, 1905</small> — Oriental (Gupta, 1987)
• Lagarosoma <small>Gupta & Gupta, 1984</small> — Neotropical
• Lophoglutus <small>Gauld, 1984</small> — Australian
• Microstenus <small>Szépligeti, 1916</small> — Oriental
• Nesolinoceras <small>Ashmead, 1906</small> — Neotropical
• Pharzites <small>Cameron, 1905</small> — Oriental
• Prosthoporus <small>Porter, 1976</small> — Neotropical
• Pterocryptus <small>Szépligeti, 1916</small> — Ethiopian, Palearctic
• Schreineria <small>Schreiner, 1905</small> — Ethiopian, Oriental, Palearctic
• Spathacantha <small>Townes, 1970a</small> — Ethiopian
• Tanepomidos <small>Gupta & Jonathan, 1971</small> — Oriental
• Torbda <small>Cameron, 1902</small> — Oriental
• Trypha <small>Townes, 1970a</small> — Neotropical
• Xanthocryptus <small>Cameron, 1901</small> — Australian, Oriental
• Xoridesopus <small>Cameron, 1907</small> — Oriental
• Subtribe Glodianina
• Dicamixus <small>Szépligeti, 1916</small> — Neotropical
• Glodianus <small>Cameron, 1902</small> — Neotropical
• Lamprocryptus <small>Schmiedeknecht, 1904</small> — Neotropical
• Subtribe Goryphina
• Allophatnus <small>Cameron, 1905</small> — Australian, Ethiopian, Oriental (Townes, 1971)
• Baltazaria <small>Townes, 1961b</small> — Neotropical, Oriental
• Bozakites <small>Seyrig, 1952</small> — Ethiopian
• Buodias <small>Cameron, 1902</small> — Ethiopian, Oriental (Gupta, 1987)
• Hoeocryptus <small>Habermehl, 1902</small>
• Takastenus <small>Uchida, 1931 (Gauld, 1984</small> — Gupta, 1987)
• Soratsia <small>Seyrig, 1952</small> (Townes & Townes, 1973)
• Ceratella <small>Seyrig, 1952</small>
• Calosphyrum <small>Townes, 1970a</small> — Oriental
• Ceratophenax <small>Seyrig, 1952</small> — Ethiopian
• Colaulus <small>Townes, 1970a</small> — Oriental
• Costifrons <small>Townes, 1970a</small> — Ethiopian
• Debilos <small>Townes, 1966</small> — Neotropical
• Diapetimorpha <small>Viereck, 1913</small> — Nearctic, Neotropical
• Euchalinus <small>Townes, 1961b</small> — Australian, Oriental
• Formostenus <small>Uchida, 1931</small> (Jonathan, 1980) — Oriental
• Fotsiforia <small>Seyrig, 1952</small> — Ethiopian, Oriental (Jonathan, 1980)
• Friona <small>Cameron, 1902</small> — Oriental
• Gambroides <small>Betrem, 1941</small> — Australian, Ethiopian, Oriental (Jonathan, 1980)
• Goryphus <small>Holmgren, 1868</small> — Australian, Ethiopian, Oriental
• Hemisphragia <small>Seyrig, 1952</small> — Ethiopian
• Hoeocryptus <small>Habermehl, 1902</small> — Ethiopian
• Hylophasma <small>Townes, 1970a</small> — Holarctic, Neotropical
• Isotima <small>Förster, 1869</small> — Oriental, Ethiopian
• Larpelites <small>Cameron, 1904</small> — Ethiopian
• Lavinifia <small>Seyrig, 1952</small> — Ethiopian
• Listrognathus <small>Tschek, 1870</small> — Australian, Ethiopian, Holarctic, Oriental
• Loxopus <small>Townes, 1970a</small> — Neotropical
• Madagascesa <small>Kocak & Kemal, 2008 - formerly Perinetia Seyrig, 1952</small> — Ethiopian
• Madastenus <small>Seyrig, 1952</small> — Ethiopian
• Melcha <small>Cameron, 1902</small> — Oriental
• Menaforia <small>Seyrig, 1952</small> — Ethiopian, Oriental
• Necolio <small>Cheesman, 1936</small> — Australian, Ethiopian, Oriental, Palearctic
• Neobuodias <small>Rao & Nikam, 1984</small> — Oriental
• Owenus <small>Townes, 1970a</small> — Ethiopian
• Perjiva <small>Jonathan & Gupta, 1973</small> — Oriental
• Piambia <small>Seyrig, 1952</small> — Ethiopian
• Pseudotricapus <small>Jonathan, 1987</small> — Oriental
• Skeatia <small>Cameron, 1901</small> — Ethiopian, Oriental (Jonathan & Gupta, 1973)
• Syntrips <small>Gauld, 1984</small> — Australian
• Tanyloncha <small>Townes, 1970a</small> — Ethiopian
• Tolonus <small>Seyrig, 1952</small> — Ethiopian
• Trafana <small>Seyrig, 1952</small> — Ethiopian
• Tricapus <small>Townes, 1970a</small> — Ethiopian
• Tsirambia <small>Seyrig, 1952</small> — Ethiopian
• Subtribe Lymeonina

• Acerastes <small>Cushman, 1929</small> — Nearctic, Neotropical
• Basileucus <small>Townes, 1970a</small> — Neotropical
• Bathyzonus <small>Townes, 1970a</small> — Neotropical
• Bicryptella <small>Strand, 1917</small> — Neotropical
• Dismodix <small>Townes, 1966</small> — Neotropical
• Golbachiella <small>Townes, 1970a</small> — Neotropical
• Latosculum <small>Townes, 1966</small> — Neotropical
• Lymeon <small>Förster, 1869</small> — Nearctic, Neotropical
• Mallochia <small>Viereck, 1912</small> — Nearctic, Neotropical
• Pachysomoides <small>Strand, 1917</small> — Nearctic, Neotropical
• Petila <small>Tedesco & Aguiar, 2009</small> — Neotropic
• Polycyrtidea <small>Viereck, 1913</small> — Nearctic, Neotropical
• Polyphrix <small>Townes, 1970a</small> — Neotropical
• Priotomis <small>Townes, 1970a</small> — Neotropical
• Rhinium <small>Townes, 1966</small> — Neotropical
• Savolia <small>Seyrig, 1952</small> — Ethiopian
• Strabotes <small>Townes, 1970a</small> — Neotropical
• Toechorychus <small>Townes, 1946</small> — Neotropical
• Subtribe Melanocryptina
• Melanocryptus <small>Cameron, 1902</small> — Neotropical
• Subtribe Mesostenina

• Acorystus <small>Townes, 1970a</small> — Neotropical
• Anacis <small>Porter, 1967</small> — Australian, Neotropical
• Anupama <small>Jonathan, 1982</small> — Oriental
• Bicristella <small>Townes, 1966</small> — Neotropical
• Cryptanura <small>Brullé, 1846</small> — Nearctic, Neotropical
• Diloa <small>Cheesman, 1936</small> — Australian, Oriental
• Gotra <small>Cameron, 1902</small> — Ethiopian, Oriental
• Gyrolaba <small>Townes, 1970a</small> — Oriental
• Harpura <small>Townes, 1970a</small> — Neotropical
• Hercana <small>Townes, 1970a</small> — Neotropical
• Irabatha <small>Cameron, 1906</small> — Australian, Oriental
• Junctivena <small>Gauld, 1984</small> — Australian
• Mecistum <small>Townes, 1970a</small> — Neotropical
• Mesostenus <small>Gravenhorst, 1829</small> — worldwide
• Paranacis <small>Gauld, 1984</small> — Australian
• Polycyrtus <small>Spinola, 1840</small> — Nearctic, Neotropical
• Stiromesostenus <small>Cameron, 1911</small> — Australian, Oriental
• Tomagotras <small>Gauld, 1984</small> — Australian
• Tretobasis <small>Porter, 1973</small> — Neotropical
• Subtribe Osprynchotina
• Acroricnus <small>Ratzeburg, 1852</small> — Holarctic
• Iaria <small>Cheesman, 1936</small> — Australian, Oriental
• Messataporus <small>Cushman, 1929</small> — Nearctic, Neotropical
• Nematopodius <small>Gravenhorst, 1829</small> — Oriental, Palearctic
• Osprynchotus <small>Spinola, 1841</small> — Ethiopian
• Photocryptus <small>Viereck, 1913</small> — Neotropical
• Poecilopimpla <small>Morley, 1914</small>;
• Picardiella <small>Lichtenstein, 1920</small> — Ethiopian, Oriental, Palearctic
• Sphecoctonus <small>Seyrig, 1952</small> — Ethiopian
• Stenarella <small>Szépligeti, 1916</small> — Australian, Ethiopian, Oriental
• Subtribe Sphecophagina
• Arthula <small>Cameron, 1900</small> — Australian, Oriental, Palearctic
• Latibulus <small>Gistel, 1848</small> — Oriental, Palearctic
• Sphecophaga <small>Westwood, 1840</small> — Holarctic
• Subtribe Vagenathina
• Stetholophus <small>Townes, 1970a</small> — Oriental
• Vagenatha <small>Cameron, 1901</small> — Oriental

Tribe Aptesini

• Aconias <small>Cameron, 1904</small> — Palearctic, Oriental
• Aptesis <small>Förster, 1850</small> — Ethiopian, Holarctic, Oriental
• Colocnema <small>Förster, 1869</small> — Palearctic
• Cratocryptus <small>Thomson, 1873</small> — Holarctic
• Cubocephalus <small>Ratzeburg, 1848</small> — Holarctic
• Demopheles <small>Förster, 1869</small> — Holarctic, Oriental
• Echthrus <small>Gravenhorst, 1829</small> — Holarctic
• Giraudia <small>Förster, 1869</small> — Holarctic
• Hemigaster <small>Brullé, 1846</small> — Oriental
• Javra <small>Cameron, 1903</small> — Holarctic, Oriental
• Listrocryptus <small>Brauns, 1905</small> — Palearctic
• Litochila <small>Momoi, 1965</small> — Oriental, Palearctic
• Livipurpurata <small>Wang & Yao, 1994</small> — Oriental
• Mansa <small>Tosquinet, 1896</small> — Ethiopian, Oriental
• Megaplectes <small>Förster, 1869</small> — Holarctic
• Notocampsis <small>Townes, 1970a</small> — Neotropical
• Oresbius <small>Marshall, 1867</small> — Holarctic
• Oxytaenia <small>Förster, 1869</small> — Nearctic, Neotropical
• Parmortha <small>Townes, 1962</small> — Holarctic
• Platymystax <small>Townes, 1970a</small> — Ethiopian, Neotropical, Oriental
• Plectrocryptus <small>Thomson, 1874</small> — Oriental, Palearctic
• Pleolophus <small>Townes, 1962</small> — Holarctic
• Polytribax <small>Förster, 1869</small> — Holarctic, Neotropical, Oriental
• Rhytura <small>Townes, 1962</small> — Nearctic
• Schenkia <small>Förster, 1869</small> — Holarctic
• Stomacis <small>Townes, 1970a</small> — Oriental

References

External links

• Tribe Cryptini Images. The Ecology of Commanster.