Convolutidae is a family of acoels, belonging to the phylum Xenacoelomorpha. This family contains more than a third of all known acoel species.
Description
The family Convolutidae includes acoels with a ventral mouth opening and a body-wall musculature composed both dorsally and ventrally of circular, longitudinal, and longitudinal crossover muscle fibers. The ventral body wall also has a group of U-shaped fibers. Most species are symbionts with algae. The anterior end has a cluster of frontal glands, a pair of eyes and a statocyst. The body has pigmented lens. Intestine and excretory system is absent. It is hermaphrodite but protandry is common.
Taxonomy
Genera
There are 26 genera in the family Convolutidae, of which 25 are extant. The type genus is Convoluta.
- â Gutticorpus <small>Knaust, 2021</small>'
- Achoerus <small>Beklemischev, 1914</small>
- Adenopea <small>Antonius, 1968</small>
- Amphiscolops <small>Graff, 1904</small>
- Antrosagittifera <small>Hooge & Tyler, 2001</small>
- Brachypea <small>Antonius, 1968</small>
- Conaperta <small>Antonius, 1968</small>
- Convoluta <small>ÃÂrsted, 1843</small>
- Convolutriloba <small>Hendelberg & Akesson, 1988</small>
- Haplodiscus <small>Weldon, 1888</small>
- Heterochaerus <small>Haswell, 1905</small>
- Neochildia <small>Bush, 1975</small>
- Oligochoerus <small>Beklemischev, 1963</small>
- Oxyposthia <small>Ivanov, 1952</small>
- Pelophila <small>Dörjes, 1968</small>
- Philachoerus <small>Dörjes, 1968</small>
- Polychoerus <small>Mark, 1892</small>
- Praesagittifera <small>Kostenko & Mamkaev, 1990</small>
- Pseudanaperus <small>Dörjes, 1968</small>
- Pseudoconvoluta <small>Beklemischev, 1929</small>
- Sagittifera <small>Kostenko & Mamkaev, 1990</small>
- Stomatricha <small>Hooge, 2003</small>
- Symsagittifera <small>Kostenko & Mamkaev, 1990</small>
- Thalassoanaperus <small>Hernandez, 2018</small>
- Waminoa <small>Winsor, 1990</small>
- Wulguru <small>Winsor, 1988</small>
Species
There are over 100 species recognised in the family Convolutidae:
- â Gutticorpus concentrica <small>Knaust, 2021</small>
- Achoerus caspius <small>Beklemischev, 1914</small>
- Achoerus ferox <small>Beklemischev, 1937</small>
- Achoerus pachycaudatus <small>Dörjes, 1968</small>
- Adenopea cenata <small>(Du Bois-Reymond Marcus, 1955)</small>
- Adenopea chuni <small>(Brauner, 1920)</small>
- Adenopea illardata <small>Löhner & Micoletzky, 1911</small>
- Adenopea illardatus <small>(Lohner & Micoletzky, 1911)</small>
- Amphiscolops bermudensis <small>Hyman 1939</small>
- Amphiscolops castellonensis <small>Steinböck, 1954</small>
- Amphiscolops cinereus <small>(Graff, 1874)</small>
- Amphiscolops evelinae <small>Marcus 1947</small>
- Amphiscolops fuligineus <small>Peebles 1913</small>
- Amphiscolops gemelliporus <small>Marcus 1954</small>
- Amphiscolops gerundensis <small>Steinböck, 1954</small>
- Amphiscolops japonicus <small>Kato 1947</small>
- Amphiscolops marinelliensis <small>Beltagi & Khafagi, 1984</small>
- Amphiscolops mosaicus <small>Kozloff 1998</small>
- Amphiscolops potocani <small>Achatz, 2008</small>
- Amphiscolops trifurcatus <small>(Beltagi 1983)</small>
- Amphiscolops zeii <small>Riedl 1956</small>
- Antrosagittifera corallina <small>Hooge & Tyler, 2001</small>
- Brachypea kenoma <small>Antonius 1968</small>
- Conaperta antonii <small>Achatz, Hooge & Tyler, 2007</small>
- Conaperta cirrata <small>Achatz, Hooge & Tyler, 2007</small>
- Conaperta flavibacillum <small>(Jensen, 1878)</small>
- Conaperta lineata <small>(Peebles, 1915)</small>
- Convoluta albomaculata <small>(Pereyaslawzewa, 1892)</small>
- Convoluta baltica <small>Meixner, 1938</small>
- Convoluta bimaculata <small>Graff, 1882</small>
- Convoluta bohmigi <small>(Brauner, 1920)</small>
- Convoluta borealis <small>Sabussow 1900</small>
- Convoluta boyeri <small>Bush 1984</small>
- Convoluta chiridotae <small>(Beklemischev, 1915)</small>
- Convoluta confusa <small>Graff 1904</small>
- Convoluta convoluta <small>(Abildgaard, 1806)</small>
- Convoluta enelitta <small>Antonius 1968</small>
- Convoluta furugelmi <small>Mamkaev, 1971</small>
- Convoluta henseni <small>Böhmig, 1895</small>
- Convoluta hipparchia <small>Pereyaslawzewa 1892</small>
- Convoluta kikaiensis <small>Yamasu 1982</small>
- Convoluta krana <small>Antonius, 1968</small>
- Convoluta lacazii <small>Graff 1891</small>
- Convoluta lacrimosa <small>Achatz, Hooge & Tyler, 2007</small>
- Convoluta marginalis <small>Ivanov 1952</small>
- Convoluta maris-alba <small>(Sabussow, 1899)</small>
- Convoluta marisalbi <small>Sabussov, 1900</small>
- Convoluta naviculae <small>Yamasu 1982</small>
- Convoluta niphoni <small>Achatz, 2008</small>
- Convoluta pelagica <small>Lohner & Micoletzky 1911</small>
- Convoluta philippinensis <small>Bush 1984</small>
- Convoluta punctata <small>Riedl, 1959</small>
- Convoluta pygopora <small>Antonius 1968</small>
- Convoluta schmidti <small>Czerniavsky, 1881</small>
- Convoluta schuelii <small>Achatz, 2008</small>
- Convoluta semperi <small>Graff, 1882</small>
- Convoluta sordida <small>Graff 1882</small>
- Convoluta thela <small>(Antonius, 1968)</small>
- Convoluta uljanini <small>Graff, 1904</small>
- Convoluta variabilis <small>(Pereyaslawzewa, 1892)</small>
- Convolutriloba hastifera <small>Winsor, 1990</small>
- Convolutriloba longifissura <small>Bartolomaeus & Balzer, 1997</small>
- Convolutriloba macropyga <small>Shannon & Achatz, 2007</small>
- Convolutriloba retrogemma <small>Hendelberg & Akesson, 1988</small>
- Haplodiscus acuminatus <small>Böhmig, 1895</small>
- Haplodiscus bocki <small>Dörjes 1970</small>
- Haplodiscus incola <small>(Leiper, 1902)</small>
- Haplodiscus obtusus <small>Böhmig, 1895</small>
- Haplodiscus ovatus <small>Böhmig, 1895</small>
- Haplodiscus piger <small>Weldon 1888</small>
- Haplodiscus ussovi <small>Sabussow, 1896</small>
- Haplodiscus weldoni <small>Böhmig, 1895</small>
- Heterochaerus australis <small>Haswell 1905</small>
- Heterochaerus blumi <small>(Achatz, Hooge & Tyler, 2007)</small>
- Heterochaerus carvalhoi <small>(Marcus, 1952)</small>
- Heterochaerus langerhansi <small>(Graff, 1882)</small>
- Heterochaerus sargassi <small>(Hyman 1939)</small>
- Neochildia fusca <small>Bush, 1975</small>
- Oligochoerus bakuensis <small>Beklemischev 1963</small>
- Oligochoerus chlorella <small>Beklemischev 1963</small>
- Oligochoerus conops <small>Beklemischev, 1963</small>
- Oligochoerus erythrophthalmus <small>Beklemischev 1963</small>
- Oligochoerus limnophilus <small>Ax & Doerjes 1966</small>
- Oligochoerus melanops <small>Beklemischev 1963</small>
- Oligochoerus xanthella <small>Beklemischev 1963</small>
- Oxyposthia praedator <small>Ivanov 1952</small>
- Pelophila lutheri <small>(Westblad, 1946)</small>
- Philachoerus johanni <small>Dörjes, 1968</small>
- Praesagittifera gracilis <small>(Yamasu, 1982)</small>
- Praesagittifera naikaiensis <small>(Yamasu, 1982)</small>
- Praesagittifera shikoki <small>Kostenko & Mamkaev 1990</small>
- Sagittifera sagittifera <small>(Ivanov, 1952)</small>
- Stomatricha hochbergi <small>Hooge, 2003</small>
- Thalassoanaperus australis <small>(Westblad, 1952)</small>
- Thalassoanaperus biaculeatus <small>(Boguta, 1970)</small>
- Thalassoanaperus gardineri <small>(Graff, 1911)</small>
- Thalassoanaperus ornatus <small>(Beltagi, 2001)</small>
- Thalassoanaperus rubellus <small>(Westblad, 1945)</small>
- Thalassoanaperus singularis <small>(Hooge & Smith, 2004)</small>
- Thalassoanaperus sulcatus <small>(Beklemischev, 1914)</small>
- Thalassoanaperus tvaerminnensis <small>(Luther, 1912)</small>
- Waminoa brickneri <small>Ogunlana, Hooge, Tekle, Benayahu, Barneah & Tyler, 2005</small>
- Waminoa litus <small>Winsor 1990</small>
- Wulguru cuspidata <small>Winsor 1988</small>
Notes
References