Columella is a genus of very small, air-breathing land snails, terrestrial pulmonate gastropod mollusks in the family Truncatellinidae, the whorl snails and allies.
Taxonomy
Columella is commonly classified within the Vertiginidae. However, as shown by Nekola & Coles (2016), the genus is not that closely related to the Vertiginidae, but may rather be affiliated with the Chondrinidae or form a pupilloid family of its own. A systematic revision is pending.
Species
Species within the genus Columella include:
- Columella acicularis <small>Almuhambetova, 1979</small>
- Columella aspera <small>Waldén, 1966</small>
- Columella columella <small>(G. v. Martens, 1830)</small>
- Columella edentula <small>(Draparnaud, 1805)</small> - toothless column
- Columella hartmutnordsiecki <small>Schlickum & Geissert, 1980</small>
- Columella hasta <small>(Hanna, 1911)</small>
- Columella intermedia <small>Schileyko & Almuhambetova, 1984</small>
- Columella microspora <small>(R.T. Lowe, 1852)</small>
- Columella nymphaepratensis <small>Hlaváà& Pokryszko, 2009</small>
- Columella polvonense <small>(Pilsbry, 1894)</small>
- Columella simplex <small>(Gould, 1840)</small>
- Columella talgarica <small>Schileyko & Rymzhanov, 2010</small>
- Columella wyciski <small>Schlickum & Strauch, 1972</small>
Species brought into synonymy:
- Columella inornata <small>(Michaud, 1831)</small>: synonym of Columella edentula <small>(Draparnaud, 1805)</small> (junior synonym)
- Columella ninagongonis <small>Pilsbry, 1935</small>: synonym of Truncatellina ninagongonis <small>(Pilsbry, 1935)</small> (original combination)
- Columella tridentata <small>A. B. Leonard, 1946</small>: synonym of Gastrocopta ruidosensis <small>(Cockerell, 1899)</small>
References