Colossendeidae is a family of sea spiders (class Pycnogonida). This family includes more than 100 species distributed among six genera. These sea spiders inhabit the deep sea mostly but are also found in shallower waters, especially in Antarctic and Arctic waters.
Description
Most colossendeids can grow to giant sizes as adults, far larger than any other sea spiders. The largest sea spider, Colossendeis colossea, can reach a leg span of 70 cm, whereas the subfamily Hedgpethiinae includes tiny species with leg span of less than 1 cm. Like most sea spiders, colossendeids usually have four pairs of legs in adults, except for three species, Decolopoda australis, D. qasimi, and Pentacolossendeis reticulata, with five leg pairs, and one species, Dodecolopoda mawsoni, with six leg pairs.
Colossendeids typically have no chelifores, except in the extra-legged species which have robust pincers and 2-segmented scapes. The palps and ovigers are present in both males and females, they are elongated with basal processes located very close to each other. The palps have 8 or 9 segments (historically 9 or 10 as the basal process was once thought to be a segment) while the ovigers have 10 segments. Each oviger ends in a strong terminal claw and compound spines arranged in fields or rows on a well formed strigilis (hook-like distal segments). The proboscis is long, usually longer than the body. The body and legs are usually glabrous. The trunk segments are either segmented (Hedgpethiinae) or fused (Colossendeinae). The legs are very long, and the terminal main claws have no auxiliary claws. gonopores represented on some or all legs.
Taxonomy
- Colossendeinae <small>Jarzynsky, 1870</small>
- Colossendeis <small>Jarzinsky, 1870</small> (67 species)
- Colossendeis acuta <small>Stiboy-Risch, 1993</small>
- Colossendeis adelpha <small>Child, 1998</small>
- Colossendeis angusta <small>Sars, 1877</small>
- Colossendeis aperta <small>Turpaeva, 2005</small>
- Colossendeis arcuata <small>A. Milne-Edwards, 1885</small>
- Colossendeis australis <small>Hodgson, 1907</small>
- Colossendeis avida <small>Pushkin, 1970</small>
- Colossendeis belekurovi <small>Pushkin, 1993</small>
- Colossendeis bicincta <small>Schimkewitsch, 1893</small>
- Colossendeis brevirostris <small>Child, 1995</small>
- Colossendeis bruuni <small>Fage, 1956</small>
- Colossendeis clavata <small>Meinert, 1899</small>
- Colossendeis colossea <small>Wilson, 1881</small>
- Colossendeis concedis <small>Child, 1995</small>
- Colossendeis cucurbita <small>Cole, 1909</small>
- Colossendeis curtirostris <small>Stock, 1963</small>
- Colossendeis dalli <small>Child, 1995</small>
- Colossendeis drakei <small>Calman, 1915</small>
- Colossendeis elephantis <small>Child, 1995</small>
- Colossendeis enigmatica <small>Turpaeva, 1974</small>
- Colossendeis ensifer <small>Child, 1995</small>
- Colossendeis fijigrypos <small>Bamber, 2004</small>
- Colossendeis fragilis <small>Pushkin, 1993</small>
- Colossendeis gardineri <small>Carpenter, 1907</small>
- Colossendeis geoffroyi <small>Mane-Garzon, 1944</small>
- Colossendeis gibbosa <small>Mobius, 1902</small>
- Colossendeis glacialis <small>Hodgson, 1907</small>
- Colossendeis gracilis <small>Hoek, 1881</small>
- Colossendeis grassa <small>Pushkin, 1993</small>
- Colossendeis hoeki <small>Gordon, 1944</small>
- Colossendeis insolita <small>Pushkin, 1993</small>
- Colossendeis japonica <small>Hoek, 1898</small>
- Colossendeis korotkevitschi <small>Pushkin, 1984</small>
- Colossendeis kurtchatovi <small>Turpaeva, 1993</small>
- Colossendeis leninensis <small>Pushkin, 1993</small>
- Colossendeis leptorhynchus <small>Hoek, 1881</small>
- Colossendeis longirostris <small>Gordon, 1938</small>
- Colossendeis losinskii <small>Turpaeva, 2002</small>
- Colossendeis macerrima <small>Wilson, 1881</small>
- Colossendeis media <small>Hoek, 1881</small>
- Colossendeis megalonyx <small>Fry & Hedgpeth, 1969</small>
- Colossendeis melancholicus <small>Stock, 1975</small>
- Colossendeis mica <small>Pushkin, 1970</small>
- Colossendeis microsetosa <small>Hilton, 1943</small>
- Colossendeis minor <small>Schimkewitsch, 1893</small>
- Colossendeis minuta <small>Hoek, 1881</small>
- Colossendeis mycterismos <small>Bamber, 2004</small>
- Colossendeis nasuta <small>Hedgpeth, 1949</small>
- Colossendeis notialis <small>Child 1995</small>
- Colossendeis oculifera <small>Stock, 1963</small>
- Colossendeis peloria <small>Child, 1994</small>
- Colossendeis perforata <small>Turpaeva, 1993</small>
- Colossendeis pipetta <small>Stock, 1991</small>
- Colossendeis proboscidea <small>(Sabine)</small>
- Colossendeis robusta <small>Hoek, 1881</small>
- Colossendeis rostrata <small>Turpaeva, 1994</small>
- Colossendeis scoresbii <small>Gordon, 1932</small>
- Colossendeis scotti <small>Calman, 1915</small>
- Colossendeis sinuosa <small>Stock, 1997</small>
- Colossendeis spicula <small>Child, 1994</small>
- Colossendeis stramenti <small>Fry and Hedgpeth, 1969</small>
- Colossendeis subminuta <small>Schimkewitsch, 1893</small>
- Colossendeis tenera <small>Hilton, 1943</small>
- Colossendeis tenuipedis <small>Pushkin, 1993</small>
- Colossendeis tethya <small>Turpaeva, 1974</small>
- Colossendeis tortipalpis <small>Gordon, 1932</small>
- Colossendeis vityazi <small>Turpaeva, 1973</small>
- Colossendeis wilsoni <small>Calman, 1915</small>
- Decolopoda <small>Eights, 1835</small>
- Decolopoda australis <small>Eights, 1835</small> â Scotia Sea (littoral)
- Decolopoda quasimi <small>Sree, Sreepada & Parulekar, 1993</small>
- Dodecolopoda <small>Calman & Gordon, 1933</small>
- Dodecolopoda mawsoni <small>Calman & Gordon, 1933</small> â Scotia Sea
- Pentacolossendeis <small>Hedgpeth, 1943</small>
- Pentacolossendeis reticulata <small>Hedgpeth, 1943</small>
- Hedgpethiinae <small>Pu</small><small>shkin, 1990</small>
- Hedgpethia <small>Turpa</small><small>eva, 1973</small> (12 species)
- Rhopalorhynchus
- Rhopalorhynchus cinclus <small>Bamber, 2001</small> â Bodgaya Southern Rim Reef, Malaysia (on a sponge)
- Rhopalorhynchus claudus <small>Stock, 1975</small> â Barbados (muddy sand, shell debris, sponge bottom)
- Rhopalorhynchus clavipes <small>Carpenter, 1893</small> â Torres Strait (between reefs)
- Rhopalorhynchus filipes <small>Stock, 1991</small> â between Loyalty Islands and New Caledonia, near Chesterfield Island
- Rhopalorhynchus gracillimus <small>Carpenter, 1907</small> â Maldive Islands (on a black crinoid)
- Rhopalorhynchus kroeyeri <small>Wood-Mason, 1873</small> â Andaman Islands (on the bottom of filamentous algae)
- Rhopalorhynchus lomani <small>Stock, 1958</small> â Makassar Strait, Indonesia
- Rhopalorhynchus mortenseni <small>Stock, 1958</small> â off Jolo, Sulu Archipelago (on sand and corals)
- Rhopalorhynchus pedunculatum <small>Stock, 1957</small> â Red Sea (shore)
- Rhopalorhynchus sibogae <small>Stock, 1958</small> â Flores Sea, Indonesia (on mud and sand)
- Rhopalorhynchus tenuissimus <small>Haswell, 1884</small> â Port Denison, Australia
References
Sources