Carabus

Carabus is a genus of beetles in the family Carabidae, and was first described in 1758 by Carl Linnaeus.

The genus is highly diverse with 94 subgenera, over 900 species and 2300 subspecies, thus is the largest genus in the subfamily Carabinae. The vast majority are native to the Palearctic, but 16 Nearctic species are also known. Carabus species are long, and most species are wingless and often very colourful. These are nocturnal, predatory beetles that feed on snails, earthworms, and caterpillars. Most Carabus species were thought to have inhabited the Eurasian forest, but their low dispersal abilities altered the distribution of lineages within the genus.

Diet

Adult Carabus feed on both small live and dead animals such as slugs, snails, earthworms, and insects in all stages, sometimes dead vertebrates. The ways of feeding on snails are different for their adaptations, as macrocephalic beetles feed on snails by crushing the snail shell, and stenocephalic beetles feed on snails by inserting their heads into the snail shell.

Species

Subgenus Acathaicus <small>Reitter, 1896</small>

Carabus alexandrae <small>Semenov, 1887</small>

Subgenus Acoptolabrus <small>A.Morawitz, 1886</small>

Carabus changgeonleei <small>(Ishikawa & Kim, 1983)</small>

Carabus constricticollis <small>(Kraatz, 1886)</small>

Carabus gehinii <small>Fairmaire, 1876</small>

Carabus haeckeli <small>Brezina & Imura, 1997</small>

Carabus jingkeanus <small>A.Müller, 2007</small>

Carabus leechi <small>(Bates, 1888)</small>

Carabus lopatini <small>A.Morawitz, 1886</small>

Carabus mirabilissimus <small>(Ishikawa & Deuve, 1982)</small>

Carabus planicranion <small>Rapuzzi, 2015</small>

Carabus schrenckii <small>(Motschulsky, 1860)</small>

Subgenus Acoptopterus <small>Lapouge, 1927</small>

Carabus agamemnon <small>Breuning, 1943</small>

Carabus andreiianus <small>Deuve, 2007</small>

Carabus anhuinus <small>Imura, 1996</small>

Carabus baimanorum <small>Deuve, 1999</small>

Carabus baogai <small>Häckel & Sehnal, 2009</small>

Carabus battoniensis <small>Deuve, 1991</small>

Carabus bornianus <small>G.Hauser, 1922</small>

Carabus dabashanus <small>Imura, 1995</small>

Carabus daiyunshan <small>Kleinfeld, 1998</small>

Carabus dongchuanicus <small>Deuve, 1994</small>

Carabus donjuan <small>(Imura, 2011)</small>

Carabus dreuxioides <small>Deuve, 1998</small>

Carabus emanuelei <small>Imura, 1997</small>

Carabus fontellus <small>Deuve, 1997</small>

Carabus jinnanicus <small>Deuve & Tian, 2006</small>

Carabus koidei <small>Imura, 1996</small>

Carabus kucerai <small>Deuve, 1997</small>

Carabus kweitshauensis <small>Mandl, 1975</small>

Carabus lacertosus <small>Cavazzuti, 2008</small>

Carabus latipennis <small>Breuning, 1932</small>

Carabus ludingensis <small>Deuve & Vigna Taglianti, 1992</small>

Carabus maowenensis <small>Deuve, 2001</small>

Carabus mianningensis <small>Kleinfeld, 2000</small>

Carabus mirandus <small>Cavazzuti & Rapuzzi, 2005</small>

Carabus morphocaraboides <small>Deuve, 1989</small>

Carabus nanosomus <small>(G.Hauser, 1931)</small>

Carabus nestor <small>(Breuning, 1934)</small>

Carabus oblongior <small>Deuve, 1992</small>

Carabus ohomopteroides <small>Deuve & Tian, 2004</small>

Carabus paris <small>Breuning, 1932</small>

Carabus platyfluvius <small>Imura, 2012</small>

Carabus pseudolatipennis <small>Deuve, 1991</small>

Carabus relictus <small>Semenov, 1898</small>

Carabus roccaianus <small>Deuve, 2011</small>

Carabus tianbaoshan <small>Kleinfeld, 1998</small>

Carabus tieguanzi <small>Imura, 1990</small>

Carabus vigil <small>Semenov, 1898</small>

Carabus vigilax <small>Bates, 1890</small>

Carabus wutaishanicus <small>A.Müller, 2000</small>

Carabus xupuensis <small>Kleinfeld, 1998</small>

Carabus yunnanus <small>Fairmaire, 1886</small>

Subgenus Acrocarabus <small>Lapouge, 1930</small>

Carabus callisthenoides <small>Semenov, 1888</small>

Carabus guerini <small>Fischer von Waldheim, 1842</small>

Subgenus Alipaster <small>Reitter, 1896</small>

Carabus barovskii <small>Semenov & Znojko, 1932</small>

Carabus dengyuani <small>Deuve & Tian, 2012</small>

Carabus dshungaricola <small>Deuve, 1990</small>

Carabus eokirgisicus <small>Kabak, 1990</small>

Carabus eous <small>A.Morawitz, 1889</small>

Carabus eugeniellus <small>Obydov, 2000</small>

Carabus foreli <small>G.Hauser, 1922</small>

Carabus hiekei <small>Kabak & Kryzhanovskij, 1990</small>

Carabus infantulus <small>A.Morawitz, 1886</small>

Carabus karkarensis <small>Kabak & Ovtshinnikov, 1994</small>

Carabus khalyktauensis <small>Kabak, 2005</small>

Carabus kiritschenkoi <small>Breuning, 1934</small>

Carabus malkovskyi <small>Kabak, 1990</small>

Carabus manap <small>Brezina & Kabak, 1993</small>

Carabus pauliani <small>Deuve, 2004</small>

Carabus pupulus <small>A.Morawitz, 1889</small>

Carabus rueckbeili <small>Breuning, 1932</small>

Carabus semenovianus <small>Breuning, 1932</small>

Carabus semenoviellus <small>Breuning, 1932</small>

Carabus shirtalensis <small>Gottwald, 1990</small>

Carabus thianshanskii <small>Breuning, 1932</small>

Carabus tianshanicola <small>Deuve & Tian, 2003</small>

Carabus valikhanovi <small>Kabak, 1990</small>

Subgenus Apoplesius <small>Deuve, 1990</small>

Carabus omphreodes <small>Reitter, 1898</small>

Subgenus Apotomopterus <small>Hope, 1838</small>

Carabus adolescens <small>(G.Hauser, 1925)</small>

Carabus aeneocupreus <small>(G.Hauser, 1932)</small>

Carabus ajax <small>Breuning, 1933</small>

Carabus angermaieri <small>Kleinfeld, 2008</small>

Carabus anlongensis <small>Deuve & Tian, 2006</small>

Carabus antoniettae <small>Cavazzuti, 2002</small>

Carabus anxiensis <small>Kleinfeld, 1998</small>

Carabus arrowi <small>G.Hauser, 1913</small>

Carabus benardi <small>Breuning, 1931</small>

Carabus birmanus <small>Andrewes, 1929</small>

Carabus boulbenianus <small>Deuve, 1996</small>

Carabus bousquetellus <small>Deuve, 1998</small>

Carabus breuningianus <small>Lemoult, 1930</small>

Carabus candidiequus <small>(Imura, 2009)</small>

Carabus cantonensis <small>(G.Hauser, 1918)</small>

Carabus caoyutangicus <small>Deuve, 2009</small>

Carabus casaleianus <small>Deuve, 1994</small>

Carabus cavazzutiellus <small>Deuve, 2002</small>

Carabus cenwangensis <small>Deuve & Tian, 2002</small>

Carabus chadianus <small>Cavazzuti & Ratti, 1999</small>

Carabus cheni <small>Deuve, 1992</small>

Carabus chenianus <small>Deuve & Tian, 1999</small>

Carabus circe <small>Cavazzuti & Ratti, 1998</small>

Carabus clermontianus <small>Breuning, 1933</small>

Carabus cyanopterus <small>G.Hauser, 1919</small>

Carabus dabamontanus <small>Imura, 1996</small>

Carabus dapanshanicus <small>Deuve, 2013</small>

Carabus datianshanicus <small>Kleinfeld, 1997</small>

Carabus davidis <small>H.Deyrolle, 1878</small>

Carabus dechambreianus <small>Deuve & Li, 2000</small>

Carabus delavayi <small>Fairmaire, 1886</small>

Carabus dilatatipennis <small>Rapuzzi & Cavazzuti, 2006</small>

Carabus ebian <small>Cavazzuti & Rapuzzi, 2010</small>

Carabus eccoptopteroides <small>Rapuzzi, 2012</small>

Carabus eccoptopterus <small>(Kraatz, 1894)</small>

Carabus elektra <small>Kleinfeld, 2004</small>

Carabus elliptipennis <small>Deuve, 1995</small>

Carabus fallettiellus <small>Deuve, 1998</small>

Carabus fanjinensis <small>Deuve & Tian, 2001</small>

Carabus feae <small>Gestro, 1888</small>

Carabus flavihervosus <small>(Imura, 2009)</small>

Carabus foveatus <small>(G.Hauser, 1913)</small>

Carabus francottei <small>Rapuzzi, 2012</small>

Carabus fushuangensis <small>Deuve, 1997</small>

Carabus gracilithorax <small>Deuve, 1989</small>

Carabus grossefoveatus <small>(G.Hauser, 1913)</small>

Carabus guangxicus <small>Deuve, 1989</small>

Carabus guibeicus <small>Deuve & Tian, 1999</small>

Carabus guzhangensis <small>Kleinfeld, 1998</small>

Carabus hera <small>Kleinfeld, 2000</small>

Carabus hiekeianus <small>Deuve, 1991</small>

Carabus huangianus <small>Deuve & Tian, 2003</small>

Carabus hubeicus <small>Deuve, 1991</small>

Carabus ichangensis <small>Bates, 1889</small>

Carabus inagakii <small>Deuve, 1991</small>

Carabus infirmior <small>(G.Hauser, 1924)</small>

Carabus inventoides <small>Deuve & Li in Deuve, 2000</small>

Carabus inventus <small>Cavazzuti, 1999</small>

Carabus jingliae <small>Deuve & Tian, 2019</small>

Carabus jingzhongensis <small>Deuve & Tian, 2005</small>

Carabus kafka <small>Kleinfeld & Reuter, 2010</small>

Carabus keithi <small>Deuve, 1995</small>

Carabus koiwayai <small>Deuve & Imura, 1990</small>

Carabus korellianus <small>Kleinfeld, 2002</small>

Carabus kouanping <small>Maindron, 1906</small>

Carabus kouichii <small>Imura & Mizusawa, 1997</small>

Carabus kryzhanovskianus <small>Deuve, 1992</small>

Carabus lamarcki <small>Deuve, 1994</small>

Carabus lambrechti <small>Kleinfeld, 2006</small>

Carabus laoshanicus <small>Imura, 1995</small>

Carabus leda <small>Kleinfeld, 2000</small>

Carabus lizizhongi <small>Deuve & Tian, 2007</small>

Carabus longeantennatus <small>(G.Hauser, 1931)</small>

Carabus ludivinae <small>Deuve, 1996</small>

Carabus luschanensis <small>(G.Hauser, 1919)</small>

Carabus madefactus <small>Cavazzuti, 1997</small>

Carabus malaisei <small>Breuning, 1947</small>

Carabus maoershanicus <small>Cavazzuti, 1995</small>

Carabus marcilhacianus <small>Deuve & Tian, 2010</small>

Carabus masuzoi <small>(Imura & Satô, 1989)</small>

Carabus mecynodes <small>Bates, 1890</small>

Carabus melli <small>(Born, 1923)</small>

Carabus menelaus <small>Breuning, 1951</small>

Carabus microtatos <small>Cavazzuti, 1997</small>

Carabus ngi <small>Deuve, 1994</small>

Carabus nigrovagans <small>Deuve, 2016</small>

Carabus novenumus <small>Deuve, 1995</small>

Carabus odysseus <small>Breuning, 1932</small>

Carabus pangi <small>Deuve & Tian, 1999</small>

Carabus patroclus <small>Breuning, 1933</small>

Carabus penelope <small>Kleinfeld, 1997</small>

Carabus pineticola <small>Deuve & Mourzine, 2004</small>

Carabus prattianus <small>Bates, 1890</small>

Carabus prodigus <small>Erichson, 1834</small>

Carabus protenes <small>Bates, 1889</small>

Carabus pseudocenwangensis <small>Deuve & Tian, 2005</small>

Carabus pseudoguizhouensis <small>Kleinfeld, 2006</small>

Carabus pseudotorquatus <small>Deuve, 1995</small>

Carabus rapuzzi <small>Kleinfeld, 2000</small>

Carabus reni <small>Deuve & Tian, 2009</small>

Carabus rugulothorax <small>Deuve, 2016</small>

Carabus saga <small>Cavazzuti, 1997</small>

Carabus sauteri <small>Roeschke, 1912</small>

Carabus schuetzei <small>Kleinfeld, 1998</small>

Carabus semelai <small>Deuve, 1995</small>

Carabus shun <small>Deuve, 1995</small>

Carabus solidior <small>Deuve & Imura, 1990</small>

Carabus tanakaianus <small>(Imura, 2005)</small>

Carabus tarasovinus <small>Deuve, 2009</small>

Carabus telemachus <small>(G.Hauser, 1925)</small>

Carabus tengchongicola <small>Deuve, 1999</small>

Carabus thilliezi <small>Deuve, 1994</small>

Carabus thoraciculus <small>Cavazzuti, 1998</small>

Carabus tiani <small>Deuve, 2013</small>

Carabus tibeticus <small>Cavazzuti & Rapuzzi, 2005</small>

Carabus tientei <small>J.Thomson, 1857</small>

Carabus tonkinensis <small>Deuve, 1990</small>

Carabus torquatus <small>Cavazzuti, 1995</small>

Carabus toulgoeti <small>Deuve, 1989</small>

Carabus tuxeni <small>Mandl, 1979</small>

Carabus vitalisi <small>Lapouge, 1918</small>

Carabus vogtae <small>Beheim & Breuning, 1943</small>

Carabus wangziensis <small>Deuve & Tian, 2005</small>

Carabus wumingensis <small>Deuve, 1992</small>

Carabus xianhensis <small>Deuve & Tian, 2007</small>

Carabus xiaoxiangensis <small>Deuve, 1995</small>

Carabus yaophilus <small>Deuve, 1990</small>

Carabus yaorenensis <small>Deuve, 2011</small>

Carabus yinjiangicus <small>Deuve & Tian, 2001</small>

Carabus yonganensis <small>Deuve, 2008</small>

Carabus yuae <small>Deuve, 1989</small>

Carabus yuanbaoensis <small>Deuve, 1994</small>

Carabus yuanshanensis <small>Kleinfeld, 1996</small>

Carabus yunanensis <small>(Born, 1905)</small>

Carabus yundongbeicus <small>Deuve, 2002</small>

Carabus yunlingensis <small>Deuve, 1991</small>

Carabus yunnanicola <small>Deuve, 1989</small>

Carabus zengae <small>Deuve & Tian, 2000</small>

Subgenus Archicarabus <small>Seidlitz, 1887</small>

Carabus alysidotus <small>Illiger, 1798</small>

Carabus monticola <small>Dejean, 1826</small>

Carabus montivagus <small>Palliardi, 1825</small>

Carabus nemoralis <small>O.F.Müller, 1764</small>

Carabus phoenix <small>Lapouge, 1925</small>

Carabus pseudomonticola <small>Lapouge, 1908</small>

Carabus rossii <small>Dejean, 1826</small>

Carabus steuartii <small>A.Deyrolle, 1852</small>

Carabus victor <small>Fischer von Waldheim, 1836</small>

Carabus wiedemanni <small>Ménétriés, 1836</small>

Subgenus Aristocarabus <small>Semenov, 1897</small>

Carabus romanowi <small>Semenov, 1897</small>

Carabus viridifossulatus <small>Fairmaire, 1887</small>

Subgenus Aulonocarabus <small>Reitter, 1896</small>

Carabus canaliculatus <small>M.Adams, 1812</small>

Carabus gaschkewitschi <small>Motschulsky, 1859</small>

Carabus gossarei <small>Haury, 1879</small>

Carabus kabakovi <small>Lafer, 1989</small>

Carabus kolymensis <small>Lafer, 1989</small>

Carabus kurilensis <small>Lapouge, 1913</small>

Carabus mouthiezianus <small>Deuve, 1991</small>

Carabus nangnimicus <small>Blumenthal & Deuve, 1984</small>

Carabus pseudokoreanus <small>Breuning, 1932</small>

Carabus truncaticollis <small>Eschscholtz, 1833</small>

Subgenus Axinocarabus <small>A.Morawitz, 1886</small>

Carabus fedtschenkoi <small>Solsky, 1874</small>

Subgenus Calocarabus <small>Semenov, 1887</small>

Carabus aristochroides <small>Deuve, 1992</small>

Carabus aurocostulus <small>Deuve & Kalab, 2018</small>

Carabus benpo <small>Kalab, 2007</small>

Carabus deuvianus <small>Cavazzuti & Casale, 2006</small>

Carabus dietererberi <small>Heinz, 2001</small>

Carabus gratus <small>Semenov, 1887</small>

Carabus guinanensis <small>Deuve, 1991</small>

Carabus janatai <small>Brezina, 1996</small>

Carabus juengerianus <small>Kleinfeld, 1995</small>

Carabus kalabi <small>Deuve, 1990</small>

Carabus linxiaensis <small>Deuve, 1992</small>

Carabus luzeicola <small>Kalab, 2005</small>

Carabus mandarin <small>Kalab, 2002</small>

Carabus miraculix <small>Kleinfeld & Puchner, 2011</small>

Carabus nevestimus <small>(Imura, 2005)</small>

Carabus przewalskii <small>A.Morawitz, 1886</small>

Carabus punctatocostulus <small>Deuve & Kalab, 2018</small>

Carabus sementivus <small>(Imura, 2005)</small>

Carabus sifanicus <small>Semenov, 1898</small>

Carabus trichothorax <small>Brezina & Imura, 1997</small>

Carabus turnaianus <small>Deuve, 1995</small>

Carabus worahniki <small>Kleinfeld & Puchner, 2011</small>

Subgenus Carabus <small>Linnaeus, 1758</small>

Carabus angustus <small>Roeschke, 1898</small>

Carabus arcensis <small>Herbst, 1784</small>

Carabus billbergi <small>Mannerheim, 1827</small>

Carabus cartereti <small>Deuve, 1982</small>

Carabus catenulatus <small>Scopoli, 1763</small>

Carabus cumanus <small>Fischer von Waldheim, 1823</small>

Carabus deyrollei <small>Gory, 1839</small>

Carabus goryi <small>Dejean, 1831</small>

Carabus granulatus <small>Linnaeus, 1758</small>

Carabus italicus <small>Dejean, 1826</small>

Carabus leplati <small>Deuve & Li, 2008</small>

Carabus manfredeggeri <small>A.Müller, 2012</small>

Carabus manifestus <small>Kraatz, 1881</small>

Carabus menetriesi <small>Hummel, 1827</small>

Carabus mianyangensis <small>Deuve & Li, 1998</small>

Carabus namhaedoensis <small>Kwon & Lee, 1984</small>

Carabus nitididorsus <small>Ishikawa & Kim, 1983</small>

Carabus obsoletus <small>Sturm, 1815</small>

Carabus paraysi <small>Palliardi, 1825</small>

Carabus pawlowskianus <small>Deuve, 1989</small>

Carabus pseudomanifestus <small>Rapuzzi, 2009</small>

Carabus sculpturatus <small>Ménétriés, 1832</small>

Carabus sternbergi <small>Roeschke, 1898</small>

Carabus stscheglowi <small>Mannerheim, 1827</small>

Carabus szeli <small>Deuve, 1994</small>

Carabus ulrichii <small>Germar, 1823</small>

Carabus vanvolxemi <small>Putzeys, 1875</small>

Carabus vinctus <small>(Weber, 1801)</small>

Carabus xiuyanensis <small>Deuve & Li, 1998</small>

Subgenus Cathaicus <small>Bates, 1870</small>

Carabus brandti <small>Faldermann, 1835</small>

Subgenus Cathoplius <small>C.G.Thomson, 1875</small>

Carabus aliai <small>(Escalera, 1944)</small>

Carabus asperatus <small>(Dejean, 1826)</small>

Carabus stenocephalus <small>Lucas, 1866</small>

Subgenus Cechenochilus <small>Motschulsky, 1850</small>

Carabus adangensis <small>Gottwald, 1983</small>

Carabus boeberi <small>M.Adams, 1817</small>

Carabus heydenianus <small>(Starck, 1889)</small>

Carabus kokujewi <small>Semenov, 1898</small>

Carabus lazorum <small>Belousov & Zamotajlov, 1999</small>

Subgenus Cechenotribax <small>Semenov & Znojko, 1932</small>

Carabus petri <small>Semenov & Znojko, 1932</small>

Subgenus Cephalornis <small>Semenov, 1889</small>

Carabus potanini <small>(Semenov, 1887)</small>

Subgenus Chaetocarabus <small>C.G.Thomson, 1875</small>

Carabus arcadicus <small>Gistel, 1848</small>

Carabus intricatus <small>Linnaeus, 1760</small>

Carabus krueperi <small>(Reitter, 1896)</small>

Carabus lefebvrei <small>Dejean, 1826</small>

Carabus merlini <small>Schaum, 1861</small>

Subgenus Chaetomelas <small>C.G.Thomson, 1875</small>

Carabus piochardi <small>(Géhin, 1883)</small>

Subgenus Chrysocarabus <small>C.G.Thomson, 1875</small>

Carabus auronitens <small>Fabricius, 1792</small>

Carabus basilicus <small>Chevrolat, 1836</small>

Carabus hispanus <small>Fabricius, 1787</small>

Carabus olympiae <small>Sella, 1855</small>

Carabus punctatoauratus <small>Germar, 1823</small>

Carabus rutilans <small>Dejean, 1826</small>

Carabus solieri <small>Dejean, 1826</small>

Carabus splendens <small>Olivier, 1790</small>

Carabus strasseri <small>(Lauffer, 1905)</small>

Subgenus Cratocarabus <small>Reitter, 1896</small>

Carabus gorodinskii <small>Obydov, 1998</small>

Carabus kryzhanovskii <small>Bogachev, 1965</small>

Carabus puer <small>A.Morawitz, 1886</small>

Subgenus Cratocechenus <small>Reitter, 1896</small>

Carabus akinini <small>A.Morawitz, 1886</small>

Carabus jacobsoni <small>Semenov, 1908</small>

Carabus ovtshinnikovi <small>Gottwald, 1987</small>

Carabus redikortzevi <small>Semenov & Breuning, 1931</small>

Carabus tshistjakovae <small>Kabak, 2001</small>

Carabus znojkoi <small>Semenov & Breuning, 1931</small>

Subgenus Cratocephalus <small>Kirsch, 1859</small>

Carabus balassogloi <small>C.A.Dohrn, 1881</small>

Carabus chan <small>Breuning, 1932</small>

Carabus cicatricosus <small>Fischer von Waldheim, 1842</small>

Carabus corrugis <small>C.A.Dohrn, 1882</small>

Carabus narinensis <small>Csiki, 1927</small>

Carabus solskyi <small>(Ballion, 1878)</small>

Subgenus Cratophyrtus <small>Reitter, 1896</small>

Carabus hauseri <small>(Reitter in F.Hauser, 1894)</small>

Carabus katajevi <small>Gottwald, 1989</small>

Carabus kaufmanni <small>Solsky, 1874</small>

Carabus nikolajevi <small>Kabak, 1998</small>

Carabus turcosinensis <small>Mandl, 1955</small>

Subgenus Cryptocarabus <small>Reitter, 1896</small>

Carabus iliensis <small>Kabak, 1994</small>

Carabus kadyrbekovi <small>Kabak, 1994</small>

Carabus lindemanni <small>Ballion, 1878</small>

Carabus merkensis <small>Kabak, 1992</small>

Carabus mullerellus <small>Beheim & Breuning, 1943</small>

Carabus munganasti <small>Reitter, 1909</small>

Carabus sacarum <small>Kabak, 1998</small>

Carabus sororius <small>A.Morawitz, 1886</small>

Carabus subparallelus <small>Ballion, 1878</small>

Carabus tsharynensis <small>Kabak, 1994</small>

Carabus turkestanus <small>Breuning, 1928</small>

Subgenus Ctenocarabus <small>C.G.Thomson, 1875</small>

Carabus galicianus <small>Gory, 1839</small>

Carabus melancholicus <small>Fabricius, 1798</small>

Subgenus Cupreocarabus <small>Deuve, 1997</small>

Carabus balangicus <small>Cavazzuti, 2002</small>

Carabus brezinai <small>Deuve, 1994</small>

Carabus concursans <small>Deuve & Kalab, 2010</small>

Carabus drahoslavae <small>Brezina & Häckel, 2006</small>

Carabus huangi <small>Deuve, 1992</small>

Carabus laevithorax <small>Breuning, 1935</small>

Carabus lixianensis <small>Deuve, 1990</small>

Carabus melii <small>Cavazzuti & Rapuzzi, 2005</small>

Carabus minimocupreus <small>Deuve, 2003</small>

Carabus miroslavi <small>Deuve, 2000</small>

Carabus mollardianus <small>Deuve, 2016</small>

Carabus pseudohuangi <small>Deuve, 1995</small>

Carabus quintus <small>Cavazzuti & Rapuzzi, 2009</small>

Carabus sackenioides <small>Deuve, 1991</small>

Carabus sculptior <small>Deuve, 1992</small>

Carabus siguniangensis <small>Deuve, 1997</small>

Carabus verecundus <small>Deuve & Kalab, 2010</small>

Carabus xichang <small>Kleinfeld & Puchner, 2013</small>

Carabus zhegushanus <small>Cavazzuti & Rapuzzi, 2005</small>

Subgenus Cychrostomus <small>Reitter, 1896</small>

Carabus anchocephalus <small>Reitter, 1896</small>

Carabus pseudoprosodes <small>Semenov & Znojko, 1932</small>

Subgenus Cyclocarabus <small>Reitter, 1896</small>

Carabus aulacocnemus <small>Semenov, 1897</small>

Carabus belousovi <small>Kabak, 1992</small>

Carabus brosciformis <small>Semenov, 1897</small>

Carabus karaterekensis <small>Kalab, 1996</small>

Carabus mniszechii <small>Chaudoir, 1852</small>

Carabus namanganensis <small>Heyden, 1886</small>

Carabus pseudolamprostus <small>Kalab, 1996</small>

Carabus pskemicus <small>Deuve & Kalab, 1993</small>

Carabus pullus <small>Semenov & Znojko, 1932</small>

Carabus zarudnyi <small>Semenov & Znojko, 1932</small>

Subgenus Damaster <small>Kollar, 1836</small>

Carabus augustus <small>Bates, 1888</small>

Carabus blaptoides <small>(Kollar, 1836)</small>

Carabus elysii <small>J.Thomson, 1856</small>

Carabus formosus <small>Semenov, 1887</small>

Carabus fruhstorferi <small>(Roeschke, 1900)</small>

Carabus gemmifer <small>(Fairmaire, 1887)</small>

Carabus guerryi <small>Born, 1903</small>

Carabus ignigena <small>(G.Hauser, 1914)</small>

Carabus ignimitella <small>Bates, 1888</small>

Carabus ishizukai <small>Deuve & Ohshima, 1989</small>

Carabus jankowskii <small>(Oberthür, 1883)</small>

Carabus kubani <small>Deuve, 1990</small>

Carabus lafossei <small>Feisthamel, 1845</small>

Carabus liianus <small>Deuve, 2008</small>

Carabus monilifer <small>Tatum, 1847</small>

Carabus nankotaizanus <small>Kano, 1932</small>

Carabus osawai <small>Imura; Zhou & Su, 1999</small>

Carabus principalis <small>Bates, 1889</small>

Carabus pustulifer <small>Lucas, 1869</small>

Carabus smaragdinus <small>Fischer, 1823</small>

Subgenus Deroplectes <small>Reitter, 1895</small>

Carabus arcanus <small>Semenov, 1898</small>

Carabus dokhtouroffi <small>Ganglbauer, 1887</small>

Carabus klapperichianus <small>Mandl, 1955</small>

Carabus sphinx <small>Reitter, 1895</small>

Carabus staudingeri <small>Ganglbauer, 1886</small>

Subgenus Diocarabus <small>Reitter, 1896</small>

Carabus aurocinctus <small>Motschulsky, 1844</small>

Carabus beybienkoi <small>Kryzhanovskij, 1973</small>

Carabus caustomarginatus <small>Imura & Mizusawa, 1994</small>

Carabus chamissonis <small>Fischer von Waldheim, 1820</small>

Carabus dorogostaiskii <small>Shilenkov, 1983</small>

Carabus fraterculus <small>Reitter, 1895</small>

Carabus loschnikovii <small>Fischer von Waldheim, 1823</small>

Carabus lucepunctus <small>Cavazzuti & Rapuzzi, 2005</small>

Carabus massagetus <small>Motschulsky, 1844</small>

Carabus opaculus <small>Putzeys, 1875</small>

Carabus slovtzovi <small>Mannerheim, 1849</small>

Subgenus Eccoptolabrus <small>Semenov, 1898</small>

Carabus exiguus <small>Semenov, 1898</small>

Carabus sunwukong <small>Imura, 1993</small>

Subgenus Euleptocarabus <small>Nakane, 1955</small>

Carabus porrecticollis <small>Bates, 1883</small>

Subgenus Eupachys <small>Chaudoir, 1857</small>

Carabus acutithorax <small>Deuve, 1989</small>

Carabus glyptopterus <small>Fischer von Waldheim, 1828</small>

Subgenus Eurycarabus <small>Géhin, 1876</small>

Carabus faminii <small>Dejean, 1826</small>

Carabus favieri <small>Fairmaire, 1859</small>

Carabus genei <small>Gené, 1839</small>

Subgenus Fulgenticarabus <small>Deuve & Li, 1998</small>

Carabus flutschi <small>Deuve & Li, 1998</small>

Subgenus Gnathocarabus <small>Deuve, 1991</small>

Carabus kusnetzovi <small>Semenov, 1903</small>

Subgenus Goniocarabus <small>Géhin, 1885</small>

Carabus caerulans <small>A.Morawitz, 1886</small>

Carabus grombczewskii <small>Semenov, 1891</small>

Carabus gussakovskii <small>Kryzhanovskij, 1971</small>

Carabus perelloi <small>Casale, 1979</small>

Carabus sogdianus <small>Semenov, 1898</small>

Carabus stackelbergi <small>Kryzhanovskij, 1971</small>

Carabus tadzhikistanus <small>Kryzhanovskij, 1968</small>

Subgenus Hemicarabus <small>Géhin, 1876</small>

Carabus macleayi <small>Dejean, 1826</small>

Carabus nitens <small>Linnaeus, 1758</small>

Carabus serratus <small>Say, 1823</small>

Carabus tuberculosus <small>Dejean, 1829</small>

Subgenus Heterocarabus <small>A.Morawitz, 1886</small>

Carabus bischoffii <small>Chaudoir, 1848</small>

Carabus marietti <small>Cristofori & Jan, 1837</small>

Carabus muchei <small>Breuning, 1961</small>

Subgenus Homoeocarabus <small>Reitter, 1896</small>

Carabus maeander <small>Fischer von Waldheim, 1820</small>

Subgenus Hygrocarabus <small>C.G.Thomson, 1875</small>

Carabus nodulosus <small>Creutzer, 1799</small>

Carabus variolosus <small>Fabricius, 1787</small>

Subgenus Hypsocarabus <small>Semenov, 1898</small>

Carabus laotse <small>Breuning, 1943</small>

Carabus latro <small>Semenov, 1898</small>

Carabus mikhaili <small>Deuve & Mourzine, 1997</small>

Carabus qinlingensis <small>Imura, 1993</small>

Carabus taibaiensis <small>Kleinfeld, 2001</small>

Carabus tewoensis <small>Deuve, 1992</small>

Subgenus Imaibius <small>Bates, 1889</small>

Carabus anami <small>(Ledoux, 1977)</small>

Carabus baronii <small>Heinertz, 1977</small>

Carabus barysomus <small>Bates, 1889</small>

Carabus boysii <small>Tatum, 1851</small>

Carabus caschmirensis <small>Kollar & L.Redtenbacher, 1844</small>

Carabus cavifrons <small>Mandl, 1974</small>

Carabus dardiellus <small>Bates, 1889</small>

Carabus epipleuralis <small>Semenov, 1907</small>

Carabus erberi <small>Heinz, 1983</small>

Carabus gandharae <small>Heinertz, 1978</small>

Carabus gridellii <small>Breuning, 1959</small>

Carabus isabellae <small>Lassalle, 1985</small>

Carabus kaghanensis <small>Heinertz, 1978</small>

Carabus leepai <small>Heinz, 1993</small>

Carabus nouristani <small>(Ledoux, 1977)</small>

Carabus olafi <small>Deuve & J.Schmidt, 2007</small>

Carabus pachtoun <small>(Ledoux, 1975)</small>

Carabus piffli <small>Mandl, 1961</small>

Carabus rostianus <small>Semenov, 1907</small>

Carabus stoliczkanus <small>Bates, 1878</small>

Carabus wittmerorum <small>Heinertz, 1978</small>

Subgenus Iniopachus <small>Solier, 1848</small>

Carabus auriculatus <small>Putzeys, 1872</small>

Carabus pyrenaeus <small>Audinet-Serville, 1821</small>

Subgenus Ischnocarabus <small>Kraatz, 1877</small>

Carabus cychropalpus <small>Peyron, 1858</small>

Carabus tenuitarsis <small>(Kraatz, 1877)</small>

Subgenus Isiocarabus <small>Reitter, 1896</small>

Carabus dargei <small>Deuve, 1988</small>

Carabus fiduciarius <small>J.Thomson, 1856</small>

Carabus gressittianus <small>Mandl, 1975</small>

Carabus heterodynamus <small>Deuve, 1991</small>

Carabus hienfoungii <small>J.Thomson, 1857</small>

Carabus hunanicola <small>Deuve & Yu, 1992</small>

Carabus kiukiangensis <small>Bates, 1888</small>

Carabus miaorum <small>Lassalle & Prunier, 1993</small>

Carabus pseudohunanicola <small>Deuve & Tian, 2009</small>

Carabus pustululatus <small>Deuve, 1993</small>

Carabus strandiellus <small>Breuning, 1934</small>

Subgenus Lamprostus <small>Motschulsky, 1866</small>

Carabus calleyi <small>Fischer von Waldheim, 1823</small>

Carabus cylindricior <small>Deuve, 1994</small>

Carabus erenleriensis <small>Schweiger, 1964</small>

Carabus guycolasianus <small>Deuve, 2001</small>

Carabus hemprichii <small>Dejean, 1826</small>

Carabus ledouxi <small>Deuve, 2001</small>

Carabus punctatus <small>Laporte, 1834</small>

Carabus renardi <small>Chaudoir, 1846</small>

Carabus robustus <small>E.Deyrolle, 1869</small>

Carabus rostandianus <small>Deuve, 2005</small>

Carabus saulcyi <small>Piochard de la Brûlerie, 1875</small>

Carabus seroulibkin <small>Cavazzuti, 2008</small>

Carabus spinolae <small>Cristofori & Jan, 1837</small>

Carabus staveni <small>Heinz, 2002</small>

Carabus syrus <small>Roeschke, 1898</small>

Carabus torosus <small>I.Frivaldszky von Frivald, 1835</small>

Subgenus Leptocarabus <small>Géhin, 1885</small>

Carabus arboreus <small>Lewis, 1882</small>

Carabus harmandi <small>Lapouge, 1909</small>

Carabus hiurai <small>Kamiyoshi & Mizoguchi, 1960</small>

Carabus koreanus <small>Reitter, 1895</small>

Carabus kumagaii <small>(Kimura & Komiya, 1974)</small>

Carabus kyushuensis <small>Nakane, 1961</small>

Carabus marcilhaci <small>Deuve, 1992</small>

Carabus procerulus <small>Chaudoir, 1862</small>

Carabus seishinensis <small>(Lapouge, 1932)</small>

Carabus semiopacus <small>Reitter, 1895</small>

Carabus vogtianus <small>Beheim & Breuning, 1943</small>

Carabus yokoae <small>Deuve, 1988</small>

Subgenus Leptoplesius <small>Reitter, 1898</small>

Carabus dolini <small>Deuve, 1992</small>

Carabus dolonicus <small>Obydov, 1996</small>

Carabus horoshanensis <small>Deuve, 2002</small>

Carabus itshkibashi <small>Kabak, 2004</small>

Carabus kleinfeldorum <small>Kabak & Putchkov, 1995</small>

Carabus marquardti <small>Reitter, 1898</small>

Carabus merzbacheri <small>G.Hauser, 1922</small>

Carabus shokalskii <small>Semenov & Breuning, 1931</small>

Carabus subtilistriatus <small>G.Hauser, 1922</small>

Subgenus Limnocarabus <small>Géhin, 1876</small>

Carabus clatratus <small>Linnaeus, 1760</small>

Subgenus Lipaster <small>Motschulsky, 1866</small>

Carabus gordius <small>Reitter, 1898</small>

Carabus stjernvalli <small>Mannerheim, 1830</small>

Subgenus Macrothorax <small>Chenu, 1851</small>

Carabus aumontii <small>Lucas, 1850</small>

Carabus meurguesianus <small>Ledoux, 1990</small>

Carabus morbillosus <small>Fabricius, 1792</small>

Carabus planatus <small>Chaudoir, 1843</small>

Carabus rugosus <small>Fabricius, 1792</small>

Subgenus Meganebrius <small>Kraatz, 1895</small>

Carabus alanstivelli <small>(Morvan, 1981)</small>

Carabus arunensis <small>Heinertz, 1980</small>

Carabus deliae <small>(Morvan, 1972)</small>

Carabus dilatotarsalis <small>Mandl, 1979</small>

Carabus epsteini <small>Heinertz, 1978</small>

Carabus everesti <small>Andrewes, 1929</small>

Carabus franzi <small>Mandl, 1974</small>

Carabus granulatocostatus <small>Mandl, 1965</small>

Carabus indicus <small>Fairmaire, 1889</small>

Carabus kadoudali <small>(Morvan, 1982)</small>

Carabus koganae <small>Colas, 1961</small>

Carabus lebretae <small>Colas, 1961</small>

Carabus montreuili <small>Deuve, 2003</small>

Carabus pseudoharmandi <small>Mandl, 1965</small>

Carabus queinneci <small>Deuve, 1983</small>

Carabus quinlani <small>Mandl, 1965</small>

Carabus salpansis <small>Deuve, 1984</small>

Carabus santostamangi <small>Deuve & J.Schmidt, 2017</small>

Carabus scheibei <small>Eidam, 1937</small>

Carabus swatensis <small>Heinertz, 1979</small>

Carabus tamang <small>Deuve, 1983</small>

Carabus thudamensis <small>Deuve & J.Schmidt, 2017</small>

Carabus tuberculipennis <small>Mandl, 1974</small>

Carabus wallichii <small>Hope, 1831</small>

Subgenus Megodontoides <small>Deuve, 1991</small>

Carabus erwini <small>Mandl, 1975</small>

Carabus promachus <small>Bates, 1891</small>

Carabus qiangding <small>Kleinfeld & Puchner, 2007</small>

Carabus thibetanus <small>Breuning, 1950</small>

Subgenus Megodontus <small>Solier, 1848</small>

Carabus aurolimbatus <small>Dejean, 1830</small>

Carabus avinovi <small>Semenov & Znojko, 1932</small>

Carabus blakistoni <small>Newman, 1858</small>

Carabus bonvouloirii <small>Chaudoir, 1863</small>

Carabus caelatus <small>Fabricius, 1801</small>

Carabus croaticus <small>Dejean, 1826</small>

Carabus danilevskii <small>Obydov, 1993</small>

Carabus exaratus <small>Quensel, 1806</small>

Carabus germarii <small>Sturm, 1815</small>

Carabus imperialis <small>Fischer von Waldheim, 1823</small>

Carabus kantaikensis <small>Géhin, 1885</small>

Carabus kolbei <small>Roeschke, 1897</small>

Carabus leachii <small>Fischer von Waldheim, 1823</small>

Carabus obovatus <small>Fischer von Waldheim, 1828</small>

Carabus persianus <small>Roeschke, 1896</small>

Carabus planicollis <small>Küster, 1846</small>

Carabus purpurascens <small>Fabricius, 1787</small>

Carabus schoenherri <small>Fischer von Waldheim, 1820</small>

Carabus septemcarinatus <small>Motschulsky, 1840</small>

Carabus stroganowi <small>Zoubkoff, 1837</small>

Carabus tuvensis <small>Shilenkov, 1996</small>

Carabus vietinghoffii <small>M.Adams, 1812</small>

Carabus violaceus <small>Linnaeus, 1758</small>

Subgenus Mesocarabus <small>C.G.Thomson, 1875</small>

Carabus dufourii <small>Dejean, 1829</small>

Carabus lusitanicus <small>Fabricius, 1801</small>

Carabus problematicus <small>Herbst, 1786</small>

Carabus riffensis <small>Fairmaire, 1872</small>

Subgenus Microplectes <small>Reitter, 1896</small>

Carabus convallium <small>(Starck, 1889)</small>

Carabus riedelii <small>Ménétriés, 1832</small>

Subgenus Mimocarabus <small>Géhin, 1876</small>

Carabus elbursensis <small>Breuning, 1946</small>

Carabus khorasanensis <small>Deuve, 1993</small>

Carabus maurus <small>(M.Adams, 1817)</small>

Carabus pumilio <small>Küster, 1846</small>

Carabus roseni <small>Reitter, 1897</small>

Subgenus Morphocarabus <small>Géhin, 1876</small>

Carabus aeruginosus <small>Fischer von Waldheim, 1820</small>

Carabus brinevi <small>Kabak, 2014</small>

Carabus chaudoirii <small>Gebler, 1847</small>

Carabus excellens <small>Fabricius, 1798</small>

Carabus gebleri <small>Fischer von Waldheim, 1817</small>

Carabus henningi <small>Fischer von Waldheim, 1817</small>

Carabus hummelii <small>Fischer von Waldheim, 1823</small>

Carabus karpinskii <small>Kryzhanovskij & Matveev, 1993</small>

Carabus kollari <small>Palliardi, 1825</small>

Carabus kozhantschikowi <small>Lutshnik, 1924</small>

Carabus mestscherjakovi <small>Lutshnik, 1924</small>

Carabus michailovi <small>Kabak, 1992</small>

Carabus monilis <small>Fabricius, 1792</small>

Carabus odoratus <small>Motschulsky, 1844</small>

Carabus praecellens <small>Palliardi, 1825</small>

Carabus regalis <small>Fischer von Waldheim, 1820</small>

Carabus rothii <small>Dejean, 1829</small>

Carabus scheidleri <small>Panzer, 1799</small>

Carabus spasskianus <small>Fischer von Waldheim, 1823</small>

Carabus tarbagataicus <small>Kraatz, 1878</small>

Carabus venustus <small>A.Morawitz, 1862</small>

Carabus versicolor <small>I.Frivaldszky von Frivald, 1835</small>

Carabus wulffiusi <small>A.Morawitz, 1862</small>

Carabus zawadzkii <small>Kraatz, 1854</small>

Carabus zhubajie <small>Imura, 1993</small>

Subgenus Neoplectes <small>Reitter, 1885</small>

Carabus chaudoirianus <small>Lapouge, 1909</small>

Carabus iberus <small>(Fischer von Waldheim, 1823)</small>

Carabus lafertei <small>Chaudoir, 1846</small>

Carabus martviliensis <small>Retezar & Djavelidze, 1992</small>

Carabus mellyi <small>Chaudoir, 1846</small>

Carabus prunierianus <small>Deuve, 2012</small>

Carabus szekelyii <small>Retezar, 2011</small>

Carabus titarenkoi <small>Zamotajlov & Fominykh, 2014</small>

Subgenus Neoplesius <small>Reitter, 1896</small>

Carabus alexpuchneri <small>Kleinfeld & Puchner, 2011</small>

Carabus alpherakii <small>Semenov, 1898</small>

Carabus borodini <small>Heinz, 1996</small>

Carabus bruggeianus <small>Deuve, 1992</small>

Carabus chomae <small>(Imura, 2002)</small>

Carabus chuandonzicus <small>(Imura, 2011)</small>

Carabus danae <small>Kalab, 1995</small>

Carabus ditomoides <small>Deuve, 1991</small>

Carabus edacis <small>Cavazzuti, 2006</small>

Carabus glinkai <small>Heinz, 1996</small>

Carabus hummelioides <small>Deuve, 1989</small>

Carabus kamensis <small>Semenov, 1903</small>

Carabus kaznakovi <small>Semenov & Znojko, 1932</small>

Carabus kozloviellus <small>Semenov & Znojko, 1932</small>

Carabus lama <small>Semenov, 1898</small>

Carabus leptoplesioides <small>Deuve, 1992</small>

Carabus mouthiezianoides <small>Deuve & Kalab, 1992</small>

Carabus nanschanicus <small>Semenov, 1898</small>

Carabus noctivagus <small>Deuve, 1992</small>

Carabus panda <small>Deuve, 1988</small>

Carabus paulusi <small>Kalab, 1995</small>

Carabus pseudomarkamensis <small>Deuve, 1992</small>

Carabus puetzi <small>Kleinfeld, 2000</small>

Carabus sinotibeticola <small>Mandl, 1975</small>

Carabus takashimai <small>Deuve & Imura, 1993</small>

Carabus tsogoensis <small>Deuve, 1997</small>

Carabus wagae <small>Fairmaire, 1882</small>

Carabus wrzecionkoianus <small>Deuve, 2007</small>

Carabus yushuensis <small>Deuve, 1991</small>

Carabus zhangjieae <small>Deuve & Li, 2020</small>

Subgenus Nesaeocarabus <small>Bedel, 1895</small>

Carabus abbreviatus <small>Brullé, 1835</small>

Carabus coarctatus <small>Brullé, 1836</small>

Carabus faustus <small>Brullé, 1836</small>

Carabus gomerae <small>A.Müller, 2004</small>

Subgenus Ohomopterus <small>Reitter, 1896</small>

Carabus albrechti <small>A.Morawitz, 1862</small>

Carabus arrowianus <small>Breuning, 1934</small>

Carabus chugokuensis <small>(Nakane, 1961)</small>

Carabus daisen <small>(Nakane, 1953)</small>

Carabus dehaanii <small>Chaudoir, 1848</small>

Carabus esakii <small>Csiki, 1927</small>

Carabus insulicola <small>Chaudoir, 1869</small>

Carabus iwawakianus <small>(Nakane, 1953)</small>

Carabus japonicus <small>Motschulsky, 1858</small>

Carabus kimurai <small>(Ishikawa, 1966)</small>

Carabus komiyai <small>(Ishikawa, 1966)</small>

Carabus lewisianus <small>Nakane, 1953</small>

Carabus maiyasanus <small>Bates, 1873</small>

Carabus sue <small>Imura, 2012</small>

Carabus uenoi <small>(Ishikawa, 1960)</small>

Carabus yaconinus <small>Bates, 1873</small>

Carabus yamato <small>(Nakane, 1953)</small>

Subgenus Ophiocarabus <small>Reitter, 1896</small>

Carabus aeneolus <small>A.Morawitz, 1886</small>

Carabus arshanicus <small>Kabak, 2005</small>

Carabus ballionis <small>Kraatz, 1879</small>

Carabus confinis <small>Semenov, 1888</small>

Carabus ernsti <small>Kabak, 2001</small>

Carabus imperfectus <small>Semenov, 1887</small>

Carabus iteratus <small>Breuning, 1934</small>

Carabus juldusanus <small>Breuning, 1933</small>

Carabus kalabellus <small>Deuve, 1993</small>

Carabus kurdaiensis <small>Kabak, 2016</small>

Carabus latiballioni <small>Deuve, 1993</small>

Carabus politulus <small>A.Morawitz, 1886</small>

Carabus praecox <small>Semenov, 1898</small>

Carabus regeli <small>A.Morawitz, 1886</small>

Carabus rufocuprescens <small>Deuve, 1993</small>

Carabus successor <small>Reitter, 1896</small>

Carabus tekesensis <small>Deuve & Tian, 2004</small>

Carabus variabilis <small>Ballion, 1878</small>

Carabus wusunshanicus <small>Kabak, 2015</small>

Subgenus Oreocarabus <small>Géhin, 1876</small>

Carabus ghilianii <small>LaFerté-Sénectère, 1847</small>

Carabus guadarramus <small>LaFerté-Sénectère, 1847</small>

Carabus luetgensi <small>Beuthin, 1886</small>

Carabus reitterianus <small>Breuning, 1932</small>

Subgenus Orinocarabus <small>Kraatz, 1878</small>

Carabus alpestris <small>Sturm, 1815</small>

Carabus bertolinii <small>(Kraatz, 1878)</small>

Carabus bremii <small>Stierlin, 1881</small>

Carabus carinthiacus <small>Sturm, 1815</small>

Carabus castanopterus <small>A. & G.B.Villa, 1833</small>

Carabus concolor <small>Fabricius, 1792</small>

Carabus fairmairei <small>C.G.Thomson, 1875</small>

Carabus linnei <small>Panzer, 1810</small>

Carabus pedemontanus <small>Ganglbauer, 1891</small>

Carabus sylvestris <small>Panzer, 1793</small>

Subgenus Oxycarabus <small>Semenov, 1898</small>

Carabus saphyrinus <small>Cristofori & Jan, 1837</small>

Subgenus Pachycarabus <small>Géhin, 1876</small>

Carabus imitator <small>Reitter, 1883</small>

Carabus koenigi <small>Ganglbauer, 1887</small>

Carabus roseri <small>Faldermann, 1836</small>

Carabus staehlini <small>M.Adams, 1817</small>

Carabus swaneticus <small>Reitter, 1883</small>

Subgenus Pachystus <small>Motschulsky, 1866</small>

Carabus cavernosus <small>I.Frivaldszky von Frivald, 1838</small>

Carabus cribratus <small>Quensel, 1806</small>

Carabus gemellatus <small>Ménétriés, 1832</small>

Carabus glabratus <small>Paykull, 1790</small>

Carabus graecus <small>Dejean, 1826</small>

Carabus hortensis <small>Linnaeus, 1758</small>

Carabus hungaricus <small>Fabricius, 1792</small>

Carabus morio <small>Mannerheim, 1830</small>

Carabus pisidicus <small>(Peyron, 1855)</small>

Carabus preslii <small>Dejean, 1830</small>

Carabus tamsii <small>Ménétriés, 1832</small>

Subgenus Pagocarabus <small>A.Morawitz, 1886</small>

Carabus caelestinus <small>(Imura, 2007)</small>

Carabus crassesculptus <small>Kraatz, 1881</small>

Carabus draco <small>Brezina, 1999</small>

Carabus hengduanicola <small>Deuve, 1996</small>

Carabus ludmilae <small>Deuve, 1992</small>

Carabus sichuanicola <small>Deuve, 1989</small>

Carabus trachynodes <small>Bates, 1891</small>

Carabus tryznai <small>Brezina & Häckel, 2006</small>

Subgenus Pantophyrtus <small>Thieme, 1881</small>

Carabus alajensis <small>Semenov, 1897</small>

Carabus brachypedilus <small>A.Morawitz, 1886</small>

Carabus debilis <small>Semenov, 1897</small>

Carabus evstigneevi <small>Obydov, 1997</small>

Carabus ferghanicus <small>Breuning, 1933</small>

Carabus longipedatus <small>Belousov & Kabak, 1993</small>

Carabus turcomanorum <small>(Thieme, 1881)</small>

Carabus validus <small>(Kraatz, 1884)</small>

Subgenus Piocarabus <small>Reitter, 1896</small>

Carabus blumenthaliellus <small>Deuve, 1988</small>

Carabus choui <small>Deuve, 1989</small>

Carabus kitawakianus <small>Imura, 1993</small>

Carabus nanwutai <small>Kleinfeld; Korell & Wrase, 1996</small>

Carabus ohshimaianus <small>Deuve, 1988</small>

Carabus sui <small>Imura & Zhou, 1998</small>

Carabus titanus <small>Breuning, 1933</small>

Carabus vladsimirskyi <small>Dejean, 1830</small>

Subgenus Platycarabus <small>A.Morawitz, 1886</small>

Carabus creutzeri <small>Fabricius, 1801</small>

Carabus cychroides <small>Baudi di Selve, 1864</small>

Carabus depressus <small>Bonelli, 1810</small>

Carabus fabricii <small>Panzer, 1810</small>

Carabus irregularis <small>Fabricius, 1792</small>

Subgenus Procerus <small>Dejean, 1821</small>

Carabus bulgharmaadensis <small>(E.Bodemeyer, 1915)</small>

Carabus caucasicus <small>M.Adams, 1817</small>

Carabus elbursianus <small>Mandl, 1958</small>

Carabus gigas <small>Creutzer, 1799</small>

Carabus scabrosus <small>Olivier, 1790</small>

Carabus syriacus <small>(Kollar in L.Redtenbacher, 1843)</small>

Subgenus Procrustes <small>Bonelli, 1810</small>

Carabus anatolicus <small>(Chaudoir, 1857)</small>

Carabus banonii <small>Dejean, 1829</small>

Carabus chevrolati <small>Cristofori & Jan, 1837</small>

Carabus clypeatus <small>M.Adams, 1817</small>

Carabus coriaceus <small>Linnaeus, 1758</small>

Carabus hybridus <small>(Ganglbauer, 1887)</small>

Carabus mulsantianus <small>A.Morawitz, 1886</small>

Carabus paiafa <small>White, 1845</small>

Carabus talychensis <small>(Ménétriés, 1832)</small>

Subgenus Pseudocoptolabrus <small>Reitter, 1896</small>

Carabus armiger <small>Imura, 1997</small>

Carabus belousovianus <small>(Imura, 2011)</small>

Carabus branaungi <small>Imura, 1999</small>

Carabus burmanensis <small>Breuning, 1932</small>

Carabus businskyi <small>Deuve, 1990</small>

Carabus chortenensis <small>Cavazzuti, 2005</small>

Carabus firmatus <small>Cavazzuti, 1997</small>

Carabus masahiroi <small>(Imura, 2006)</small>

Carabus nosei <small>Imura, 1997</small>

Carabus pseudochortenensis <small>Deuve & Tian, 2020</small>

Carabus taliensis <small>(Fairmaire, 1886)</small>

Carabus watanabei <small>(Imura, 2003)</small>

Subgenus Pseudocranion <small>Reitter, 1896</small>

Carabus aba <small>Kalab, 2002</small>

Carabus absonus <small>Cavazzuti & Rapuzzi, 2005</small>

Carabus baxianus <small>Cavazzuti & Rapuzzi, 2007</small>

Carabus brachygnathus <small>Deuve, 2002</small>

Carabus flavigenua <small>Cavazzuti, 2002</small>

Carabus fumigatus <small>Semenov, 1898</small>

Carabus gansuensis <small>Semenov, 1887</small>

Carabus gonggaicus <small>Deuve, 1989</small>

Carabus jiudingensis <small>Deuve, 1994</small>

Carabus kitawakiellus <small>Imura, 1995</small>

Carabus labrangicus <small>Deuve, 1992</small>

Carabus lazikouensis <small>Deuve, 1997</small>

Carabus meditabundus <small>Deuve, 1992</small>

Carabus pseudosackeni <small>Deuve, 1989</small>

Carabus sackeni <small>Semenov, 1898</small>

Carabus shuamaluko <small>Deuve, 1991</small>

Carabus sinicus <small>Breuning, 1950</small>

Carabus taibaishanicus <small>Deuve, 1989</small>

Carabus tibetanophilus <small>Deuve, 1991</small>

Carabus viatorum <small>Deuve, 1992</small>

Carabus wenxianicola <small>Deuve, 1996</small>

Carabus zhanglaensis <small>Deuve, 1991</small>

Subgenus Rhigocarabus <small>Reitter, 1896</small>

Carabus allegroi <small>Cavazzuti & Rapuzzi, 2007</small>

Carabus batangicoides <small>Deuve & Tian, 2011</small>

Carabus boanoi <small>Cavazzuti, 2003</small>

Carabus broukpytlik <small>Brezina & Häckel, 2004</small>

Carabus buddaicus <small>Semenov, 1887</small>

Carabus casanova <small>(Imura & Brezina, 2008)</small>

Carabus cateniger <small>A.Morawitz, 1886</small>

Carabus cechenoides <small>Deuve & Kalab, 2014</small>

Carabus dacatraianus <small>Deuve, 1995</small>

Carabus dubifer <small>Deuve & Kalab, 2014</small>

Carabus dungchen <small>Deuve & Kalab, 2019</small>

Carabus fubianensis <small>Deuve & Kalab, 2007</small>

Carabus gentleman <small>Brezina & Häckel, 2004</small>

Carabus gigolo <small>Heinz & Brezina, 1996</small>

Carabus gigoloides <small>Cavazzuti, 2000</small>

Carabus gracilicollis <small>Semenov, 1887</small>

Carabus handelmazzettii <small>Mandl, 1955</small>

Carabus humilior <small>Deuve & Kalab, 2012</small>

Carabus impavidus <small>Cavazzuti, 2008</small>

Carabus indigestus <small>Semenov, 1898</small>

Carabus itzingeri <small>(Breuning, 1934)</small>

Carabus jintangicus <small>Deuve, 2001</small>

Carabus jiulongensis <small>Deuve, 1994</small>

Carabus ladygini <small>Semenov, 1903</small>

Carabus legrandianus <small>Deuve & Tian, 2007</small>

Carabus maleki <small>Deuve, 1991</small>

Carabus mifan <small>Kalab, 2005</small>

Carabus morawitzianus <small>Semenov, 1887</small>

Carabus nianjuaensis <small>Imura, 2016</small>

Carabus paraxiei <small>Kleinfeld & Puchner, 2013</small>

Carabus pepek <small>(Imura & Kalab, 2006)</small>

Carabus poeta <small>Semenov, 1898</small>

Carabus propiorthais <small>Cavazzuti, 2000</small>

Carabus pseudopusio <small>Deuve, 1996</small>

Carabus pusioides <small>Cavazzuti & Rapuzzi, 2007</small>

Carabus quindecim <small>Cavazzuti, 2002</small>

Carabus rhododendron <small>Deuve & Imura, 1991</small>

Carabus roborowskii <small>Semenov, 1887</small>

Carabus sehnali <small>Brezina & Häckel, 2006</small>

Carabus shaluishan <small>Kleinfeld & Puchner, 2011</small>

Carabus sininensis <small>Semenov, 1898</small>

Carabus thais <small>Heinz, 1997</small>

Carabus turnai <small>Deuve, 1994</small>

Carabus wengdaensis <small>Kleinfeld & Puchner, 2009</small>

Carabus xiei <small>Deuve, 1992</small>

Subgenus Scambocarabus <small>Reitter, 1896</small>

Carabus auritus <small>Cavazzuti, 2000</small>

Carabus kruberi <small>Fischer von Waldheim, 1820</small>

Carabus modestulus <small>Semenov, 1887</small>

Carabus sculptipennis <small>Chaudoir, 1877</small>

Carabus shaanxiensis <small>Deuve, 1991</small>

Subgenus Semnocarabus <small>Reitter, 1896</small>

Carabus carbonicolor <small>A.Morawitz, 1886</small>

Carabus cicatricosulus <small>A.Morawitz, 1886</small>

Carabus erosus <small>Motschulsky, 1866</small>

Carabus perminimus <small>Deuve, 1994</small>

Carabus regulus <small>C.A.Dohrn, 1882</small>

Carabus rustemi <small>Kabak, 2010</small>

Carabus tekeliensis <small>Kabak, 2001</small>

Carabus transiliensis <small>Semenov, 1897</small>

Subgenus Shunichiocarabus <small>Imura, 1995</small>

Carabus uenoianus <small>Imura, 1995</small>

Subgenus Sphodristocarabus <small>Géhin, 1885</small>

Carabus adamsi <small>M.Adams, 1817</small>

Carabus biroi <small>Csiki, 1927</small>

Carabus bohemani <small>Ménétriés, 1832</small>

Carabus coruhnehriensis <small>Cavazzuti, 1990</small>

Carabus elegantulus <small>Motschulsky, 1850</small>

Carabus enigmaticus <small>Heinz, 1980</small>

Carabus georgia <small>Cavazzuti, 1984</small>

Carabus gilnickii <small>E.Deyrolle, 1869</small>

Carabus heinzi <small>Breuning, 1964</small>

Carabus karasudominus <small>Cavazzuti, 1991</small>

Carabus kurdicus <small>Heinz, 1975</small>

Carabus macrogonus <small>Chaudoir, 1847</small>

Carabus pavesii <small>Cavazzuti, 1992</small>

Carabus scovitzii <small>Faldermann, 1836</small>

Carabus separatus <small>Lapouge, 1907</small>

Carabus tokatensis <small>Roeschke, 1898</small>

Carabus varians <small>Fischer von Waldheim, 1823</small>

Subgenus Stephanocarabus <small>Imura, 1995</small>

Carabus fraterculoides <small>Breuning, 1961</small>

Subgenus Tachypus <small>Weber, 1801</small>

Carabus auratus <small>Linnaeus, 1760</small>

Carabus cancellatus <small>Illiger, 1798</small>

Carabus vagans <small>Olivier, 1795</small>

Subgenus Tanaocarabus <small>Reitter, 1896</small>

Carabus finitimus <small>Haldeman, 1852</small>

Carabus forreri <small>Bates, 1882</small>

Carabus hendrichsi <small>Bolivar y Pieltain; Rotger & Coronado, 1967</small>

Carabus sylvosus <small>Say, 1823</small>

Carabus taedatus <small>Fabricius, 1787</small>

Subgenus Teratocarabus <small>Semenov & Znojko, 1932</small>

Carabus azrael <small>Semenov & Znojko, 1932</small>

Subgenus Tmesicarabus <small>Reitter, 1896</small>

Carabus cristoforii <small>Spence, 1833</small>

Subgenus Tomocarabus <small>Reitter, 1896</small>

Carabus bessarabicus <small>Fischer von Waldheim, 1823</small>

Carabus convexus <small>Fabricius, 1775</small>

Carabus decolor <small>Fischer von Waldheim, 1823</small>

Carabus marginalis <small>Fabricius, 1794</small>

Carabus rumelicus <small>Chaudoir, 1867</small>

Carabus scabripennis <small>Chaudoir, 1850</small>

Subgenus Trachycarabus <small>Géhin, 1876</small>

Carabus besseri <small>Fischer von Waldheim, 1820</small>

Carabus coriaceipennis <small>Chaudoir, 1863</small>

Carabus estreicheri <small>Fischer von Waldheim, 1820</small>

Carabus latreillei <small>Fischer von Waldheim, 1820</small>

Carabus mandibularis <small>Fischer von Waldheim, 1828</small>

Carabus perrini <small>Dejean, 1831</small>

Carabus planarius <small>Obydov, 1994</small>

Carabus scabriusculus <small>Olivier, 1795</small>

Carabus sibiricus <small>Fischer von Waldheim, 1820</small>

Subgenus Tribax <small>Fischer von Waldheim, 1817</small>

Carabus adelphus <small>(Rost, 1892)</small>

Carabus agnatus <small>(Ganglbauer, 1889)</small>

Carabus apollo <small>(Zolotarev, 1913)</small>

Carabus apschuanus <small>(Rost, 1893)</small>

Carabus balkaricus <small>Belousov & Abdurakhmanov, 1991</small>

Carabus biebersteini <small>Ménétriés, 1832</small>

Carabus certus <small>(Reitter, 1896)</small>

Carabus circassicus <small>(Ganglbauer, 1886)</small>

Carabus compressus <small>Chaudoir, 1846</small>

Carabus curlettii <small>Cavazzuti, 1984</small>

Carabus daphnis <small>Kurnakov, 1962</small>

Carabus edithae <small>Reitter, 1893</small>

Carabus edmundi <small>Semenov, 1897</small>

Carabus jason <small>Semenov, 1898</small>

Carabus kasakorum <small>Semenov, 1897</small>

Carabus kasbekianus <small>Kraatz, 1877</small>

Carabus koltzei <small>(Rost, 1889)</small>

Carabus komarowi <small>Reitter, 1882</small>

Carabus kraatzi <small>Chaudoir, 1877</small>

Carabus kratkyi <small>Ganglbauer, 1891</small>

Carabus lailensis <small>Belousov, 1992</small>

Carabus lederi <small>Reitter, 1882</small>

Carabus macropus <small>Chaudoir, 1877</small>

Carabus merdeniki <small>Cavazzuti & Korell, 1992</small>

Carabus nothus <small>M.Adams, 1817</small>

Carabus osseticus <small>M.Adams, 1817</small>

Carabus plasoni <small>(Ganglbauer, 1886)</small>

Carabus polychrous <small>(Rost, 1892)</small>

Carabus prometheus <small>Reitter, 1887</small>

Carabus protensus <small>Schaum, 1864</small>

Carabus puschkini <small>M.Adams, 1817</small>

Carabus reitteri <small>(Retowski, 1885)</small>

Carabus satyrus <small>Kurnakov, 1962</small>

Carabus schamylii <small>Hampe, 1852</small>

Carabus shtchurovi <small>Belousov & Zamotajlov, 1993</small>

Carabus starcki <small>(Heyden, 1885)</small>

Carabus starckianus <small>(Ganglbauer, 1886)</small>

Carabus titan <small>(Zolotarev, 1913)</small>

Subgenus Ulocarabus <small>Reitter, 1896</small>

Carabus stschurowskii <small>Solsky, 1874</small>

Carabus theanus <small>Reitter, 1895</small>

Extinct, not assigned to subgenus

†Carabus ceresti <small>Nel, 1987</small>

†Carabus dzieduszyckii <small>Lomnicki, 1894</small>

†Carabus elongatus <small>Brodie, 1845</small>

†Carabus foveolatus <small>Piton & Theobald, 1935</small>

†Carabus jeffersoni <small>Scudder, 1900</small>

†Carabus maeandroides <small>Lomnicki, 1894</small>

†Carabus mecothoracus <small>Zhang: Liu & Shangguan, 1989</small>

†Carabus neli <small>Deuve, 1998</small>

†Carabus novalensis <small>Omboni, 1886</small>

†Carabus ovalis <small>Hong & Wang, 1986</small>

†Carabus praearvensis <small>Lomnicki, 1894</small>

†Carabus praeviolaceus <small>Lomnicki, 1894</small>

†Carabus westwoodii <small>Giebel, 1856</small>

†Carabus winkleri <small>Weyenbergh, 1869</small>

References

• Boleslav Brezina (1999) World catalogue of the genus Carabus L. (Coleoptera, Carabidae) Pensoft Series. Faunistica, No 15. Pensoft, Sofia - Moscow
• Thierry Deuve (2007) Liste Blumenthal 2007 (Liste de taxons valides du genre Carabus L., 1758). Musée national d'histoire naturelle, Paris

Further reading

External links

• Carabus.eu
• Key to the British species in tribe Carabini - Calsoma and Carabus in Keys for the identification of British Carabidae