Calosoma alternans, the reciprocal caterpillar hunter, is a species of ground beetle in the family Carabidae. It is found in Central and South America, where it may be found in various habitats, such as tropical dry and moist forests, grasslands and cultivated fields.
Adults prey on Alabama argillacea and Spodoptera frugiperda.
These two subspecies belong to the species Calosoma alternans: