Butterfly ray

The butterfly rays are the rays forming the genus Gymnura and the family Gymnuridae. They are found in warm oceans worldwide, and occasionally in estuaries.

The body of butterfly rays is flattened and surrounded by an extremely broad disc formed by the pectoral fins, which merge in front of the head. They have a very short, thread-like, tail. They are up to in width.

McEachran et al. (1996) place the butterfly rays in the subfamily Gymnurinae of the family Dasyatidae, but this article follows FishBase and ITIS in treating them as a family.

Species

There are currently 14 species in this genus (others are considered synonyms):

• Gymnura afuerae <small>(Hildebrand, 1946)</small>
• Gymnura altavela <small>(Linnaeus, 1758)</small> – Spiny butterfly ray
• Gymnura australis <small>(E. P. Ramsay & Ogilby, 1886)</small> – Australian butterfly ray
• Gymnura crebripunctata <small>(W. K. H. Peters, 1869)</small> – Longsnout butterfly ray
• Gymnura hirundo <small>(Lowe, 1843)</small> – Madeira butterfly ray
• Gymnura japonica <small>(Temminck & Schlegel, 1850)</small> – Japanese butterfly ray
• Gymnura lessae <small>(Yokota & Carvalho, 2017)</small> – Lessa's butterfly ray
• Gymnura marmorata <small>(J. G. Cooper, 1864)</small> – California butterfly ray
• Gymnura micrura <small>(Bloch & J. G. Schneider, 1801)</small> – Smooth butterfly ray
• Gymnura natalensis <small>(Gilchrist & W. W. Thompson, 1911)</small> – Backwater butterfly ray
• Gymnura poecilura <small>(G. Shaw, 1804)</small> – Longtail butterfly ray
• Gymnura sereti <small>(Yokota & Carvalho, 2017)</small> – Seret's butterfly ray
• Gymnura tentaculata <small>(J. P. Müller & Henle, 1841)</small> – Tentacled butterfly ray
• Gymnura zonura <small>(Bleeker, 1852)</small> – Zonetail butterfly ray

• Gymnura bimaculata <small>(Norman, 1925)</small> - synonym of Gymnura japonica
• Gymnura crooki <small>Fowler, 1934</small> - synonym of Gymnura poecilura

Fossil taxa

The following fossil gymnurid genera are known:

• †Subathunura <small>Kumar & Loyal, 1987</small> (Eocene of India)
• †Jacquhermania <small>Cappetta, 1982</small> (Middle to Late Eocene of Europe and eastern North America)'
• †Ouledia <small>Cappetta, 1986</small> (Late Paleocene to Middle Eocene of North Africa, South America and South Asia)
• †Pachygymnura <small>Adnet et al., 2020</small> (Late Eocene of Egypt)