Bursa is a genus of large sea snails, marine gastropod molluscs in the family Bursidae, the frog snails or frog shells.
Description
The ovate or oblong shell is compressed, and shows two rows of continuous varices, one on each side. The aperture is ova. The columella is arcuated, and ridged or crenulated. The siphonal canal is short and recurved. The outer lip is crenated.
The species of Bursa are mostly tropical, the majority being from the Eastern seas. They are found, when variegated and with nodose armature, in rocky places and on coral reefs. The winged species, with a smoother surface, are from deep water. They move about with considerable animation, and crawl rapidly.
Recognized species
Species within the genus Bursa include:
- Bursa asperrima <small>Dunker, 1862</small>
- Bursa bufonia <small>(Gmelin, 1791)</small>
- Bursa calcipicta <small>Dall, 1908</small>
- Bursa davidboschi <small> Beu, 1987 </small>
- Bursa grayana <small>Dunker, 1862</small> - elegant frogsnail
- Bursa humilis <small>Beu, 1981 </small>
- Bursa lamarckii <small> (G. P. Deshayes, 1853) </small>
- Bursa leo <small>Shikama, 1964</small>
- Bursa luteostoma <small> W. H. Pease, 1861 </small>
- Bursa natalensis <small> Matthews & Coelho, 1970 </small>
- â Bursa nodosa <small>(Borson, 1825)</small>
- Bursa rosa <small>Perry, 1811 </small>
- Bursa rugosa <small> G.B. Sowerby II, 1835 </small>
- â Bursa sangirana <small>Beu, 2005</small>
- â Bursa transeuntis <small>Forli & Smriglio, 2021</small>
- Bursa tuberosissima <small> L. A. Reeve, 1844 </small>
Species brought into synonymy
- Bursa affinis <small>(Broderip, W.J., 1832)</small>: synonym of Dulcerana affinis <small>(Broderip, 1833)</small>
- Bursa alfredensis <small>Turton, 1932</small>: synonym of Dulcerana alfredensis <small>(W. H. Turton, 1932)</small> (original combination)
- Bursa angioyorum <small>Parth, 1990</small>: synonym of Bursa lamarckii <small>(Deshayes, 1853)</small>
- Bursa awatii <small>Ray, 1949 </small>: synonym of Korrigania awatii <small>(Ray, 1948)</small> (original combination)
- Bursa barcellosi <small>Matthews, Rios & Coelho, 1973</small>: synonym of Ranella olearium <small>(Linnaeus, 1758)</small>
- Bursa bergeri <small>C. E. Tapparone-Canefri, 1880 </small>: synonym of Bursa rhodostoma (Sowerby II, 1835)
- Bursa bubo <small>Linnaeus</small>: synonym of Tutufa bubo <small>(Linnaeus, 1758)</small> represented as Tutufa (Tutufa) bubo <small>(Linnaeus, 1758)</small>
- Bursa bubo lissostoma <small>E. A. Smith, 1914</small>: synonym of Tutufa bubo <small>(Linnaeus, 1758)</small> represented as Tutufa (Tutufa) bubo <small>(Linnaeus, 1758)</small>
- Bursa bufonia dunkeri <small> Kira, 1959 </small>: synonym of Bursa bufonia <small> Kira, 1959 </small>
- Bursa bufo <small>(Bruguiere, 1792)</small>: spelling of Bursa bubo in <small>Spry 1961</small>
- Bursa caelata <small> W. J. Broderip, 1833 </small>: synonym of Bursa corrugata <small>(Perry, 1811)</small>
- Bursa calcipicta <small>Dall, 1908</small>: synonym of Bursa rugosa <small>(G.B. Sowerby II, 1835)</small>
- Bursa californica <small>(Hinds, 1843)</small>: synonym of Crossata ventricosa <small>(Broderip, 1833)</small>
- Bursa concinna <small>Dunker, 1862</small>: synonym of Gyrineum concinnum <small>(Dunker, 1862)</small>
- Bursa condita <small> J. F. Gmelin, 1791 </small>: synonym of Colubrellina condita <small>(Gmelin, 1791)</small>
- Bursa corrugata <small>Perry, 1811</small> - gaudy frogsnail: synonym of Alanbeuella corrugata <small>(Perry, 1811)</small>
- Bursa cruentata <small> G.B. Sowerby II, 1835 </small>: synonym of Lampasopsis cruentata <small>(G. B. Sowerby II, 1835)</small>
- Bursa corrugata ponderosa <small>L. A. Reeve, 1844</small>: synonym of Ranella ponderosa <small>Reeve, 1844,</small> in turn, synonym of Bursa corrugata <small>Perry, 1811</small>
- Bursa corrugata pustulosa <small>L. A. Reeve, 1844</small>: synonym of Bursa corrugata <small>Perry, 1811</small>
- Bursa crassa (Dillwyn, 1817): synonym of Marsupina bufo <small>(Bruguière, 1792)</small>
- Bursa crumena (Lamarck, 1816): synonym of Bufonaria crumena <small>(Lamarck, 1816)</small>
- Bursa crumenoides <small> Valenciennes, 1832 </small> : synonym of Bufo crumena <small>(Lamarck, 1816)</small>
- Bursa cubaniana <small>(d'Orbigny, 1841)</small>: synonym of Dulcerana cubaniana <small>(d'Orbigny, 1847)</small>
- Bursa cumingiana <small>Dunker, 1862</small>: synonym of Bursa granularis <small>(Röding, 1798)</small>: synonym of Dulcerana granularis <small>(Röding, 1798)</small> (junior synonym)
- Bursa dunkeri <small> Kira, 1962 </small>: synonym of Bursa bufonia <small>(Gmelin, 1791)</small>
- Bursa echinata <small>(Link, 1807)</small>: synonym of Bufonaria echinata <small>(Link, 1807)</small>
- Bursa elisabettae <small>Nappo, Pellegrini & Bonomolo, 2014</small>: synonym of Dulcerana elisabettae <small>(Nappo, Pellegrini & Bonomolo, 2014)</small>
- Bursa fijiensis <small> R. B. Watson, 1881 </small>: synonym of Bursina fijiensis <small>(Watson, 1881)</small>
- Bursa finlayi <small>Mcginty, 1962</small>: synonym of Bursa (Bufonariella) ranelloides <small>(Reeve, 1844)</small>
- Bursa fosteri <small>Beu, 1987</small>: synonym of Korrigania fosteri <small>(Beu, 1987)</small>
- Bursa fuscocostata <small>Dunker, 1862</small>: synonym of Gyrineum bituberculare <small>(Lamarck, 1816)</small>
- Bursa gibbosa <small>Röding, 1798</small>: synonym of Marsupina bufo <small>(Bruguière, 1792)</small>
- Bursa gnorima <small>Melvill, 1918</small>: synonym of Bursina gnorima <small>(Melvilll, 1918)</small>
- Bursa granifera <small>Röding</small>: synonym of Bursa granularis <small>(Röding, 1798)</small> accepted as Dulcerana granularis <small>(Röding, 1798)</small>
- Bursa granularis <small>Röding, 1798</small> - Cuba frogsnail: synonym of Dulcerana granularis <small>(Röding, 1798)</small> (original combination)
- Bursa kowiensis <small>Turton, 1932</small>: synonym of Bursa granularis <small>(Röding, 1798)</small>
- Bursa lamarkii <small>(Deshayes, 1853)</small>: synonym of Bursa lamarckii <small>(Deshayes, 1853)</small>
- Bursa lamellosa <small>Dunker, 1863</small>: synonym of Aspella producta <small>(Pease, 1861)</small>
- Bursa lampas <small> C. Linnaeus, 1758 </small>: synonym of Tutufa bubo <small>(Linnaeus, 1758)</small> represented as Tutufa (Tutufa) bubo <small>(Linnaeus, 1758)</small>
- Bursa latitudo <small>Garrard, 1961</small>: synonym of Tritonoranella latitudo <small>(Garrard, 1961)</small>
- Bursa lissostoma <small> E. A. Smith, 1914 </small>: synonym of Tutufa bufo <small>(Röding, P.F., 1798</small>)
- Bursa livida <small> L. A. Reeve, 1844 </small>: synonym of Bursa granularis <small>(Röding, 1798)</small>
- Bursa lucaensis <small> Parth, 1991 </small>: synonym of Lampasopsis lucaensis <small>(Parth, 1991)</small> (original combination)
- Bursa mammata <small> P. F. Röding, 1798 </small>: synonym of Bursa bufonia <small>(Gmelin, 1791)</small>
- Bursa marginata <small>(Gmelin, 1791)</small>: synonym of Aspa marginata <small>(Gmelin, 1791)</small>
- Bursa monitata <small> P. F. Röding, 1798 </small>: synonym of Bursa bufonia <small>(Gmelin, 1791)</small>
- Bursa muehlhaeusseri <small> Parth, 1990 </small>: synonym of Bursa lamarckii <small>(Deshayes, 1853)</small>
- Bursa nigrita Mulhauser & Blocher, 1979: synonym of Tutufa nigrita <small>Mühlhäusser & Blöcher, 1979</small> represented as Tutufa (Tutufella) nigrita <small>Mühlhäusser & Blöcher, 1979</small>
- Bursa nobilis <small>(Reeve, 1844)</small>: synonym of Bursina nobilis <small>(Reeve, 1844)</small>
- Bursa pacamoni <small> Matthews & Coelho, 1971 </small>: synonym of Bursa grayana <small>Dunker, 1862</small>
- Bursa pustulosa <small> L. A. Reeve, 1844 </small>; synonym of Bursa corrugata <small>(Perry, 1811)</small>
- Bursa pygmaea <small> S. Kosuge, 1979 </small>: synonym of Bursa (Bufonariella) ranelloides <small>(Reeve, 1844)</small>
- Bursa quirihorai <small> Beu, 1987 </small>: synonym of Korrigania quirihorai <small>(Beu, 1987)</small> (basionym)
- Bursa ranelloides <small>(Reeve, 1844)</small> - fine-sculpted frogsnail: synonym of Tritonoranella ranelloides <small>(Reeve, 1844)</small>
- Bursa rhodostoma <small>(G. B. Sowerby I or II, 1835)</small> - wine-mouth frog shell: synonym of Lampasopsis rhodostoma <small>(G. B. Sowerby II, 1835)</small>
- Bursa rhodostoma thomae <small> d'Orbigny, 1842 </small> - St. Thomas frog shell: synonym of Lampasopsis thomae <small>(d'Orbigny, 1847)</small>
- Bursa rubeta <small>(Linnaeus, 1758)</small>: synonym of Tutufa (Tutufella) rubeta <small>(Linnaeus, 1758)</small>
- Bursa (Tutufa) rubeta var. gigantea Smith, 1914: synonym of Tutufa (Tutufa) bardeyi <small>(Jousseaume, 1881)</small>
- Bursa scrobilator <small>Linnaeus, 1758</small>: synonym of Talisman scrobilator <small>(Linnaeus, 1758)</small>
- Bursa scrobilator coriaceum <small>L. A. Reeve, 1844 </small>: synonym of Talisman scrobilator coriaceum <small>(Reeve, 1844)</small>
- Bursa semigranosa <small> J. B. Lamarck, 1822 </small>: synonym of Bursa granularis <small>(Röding, P.F., 1798)</small>
- Bursa spinosa <small> J. B. Lamarck, 1816 </small>: synonym of Bufonaria echinata <small>(Link, 1807)</small>
- Bursa suensonii <small>Mörch, 1853</small>: synonym of Bufonaria echinata <small>(Link, 1807)</small>
- Bursa tenuisculpta <small>(Dautzenberg and Fischer, 1906)</small>: synonym of Bursa ranelloides tenuisculpta <small>Dautzenberg & Fischer, 1906 </small>
- Bursa thomae <small>(d'Orbigny, 1842)</small>: synonym of Bursa rhodostoma <small>(G. B. Sowerby I or II, 1835)</small>: synonym of Lampasopsis thomae <small>(d'Orbigny, 1847)</small>
- Bursa tumida <small>Dunker, 1862</small>: synonym of Argobuccinum tumidum <small>(Dunker, 1862)</small>
- Bursa venustula <small>(Reeve, 1844)</small>: synonym of Bursa rhodostoma <small>(G.B. Sowerby II, 1835)</small>
- Bursa verrucosa <small>(G. B. Sowerby I, 1825)</small>: synonym of Annaperenna verrucosa <small>(G. B. Sowerby I, 1825)</small>
References
- Gofas, S.; Le Renard, J.; Bouchet, P. (2001). Mollusca, in: Costello, M.J. et al. (Ed.) (2001). European register of marine species: a check-list of the marine species in Europe and a bibliography of guides to their identification. Collection Patrimoines Naturels, 50: pp. 180âÂÂ213
- Rolán E., 2005. Malacological Fauna From The Cape Verde Archipelago. Part 1, Polyplacophora and Gastropoda.