Aphodius

Aphodius is a genus of beetles in the family Scarabaeidae. In most species both the adults and larvae are coprophagous (dung feeding) although some species have herbivorous or saprophagous larvae. Aphodius species typically dominate dung beetle communities in north temperate ecosystems. Most species are functionally classified as endocoprids, also known as dwellers, because the larvae live and feed within the dung pat itself.

With a discovery of an earlier published version by Hellwig, the correct citation for this genus is now: Aphodius Hellwig, 1798 (p. 101; 24 January). Type species by subsequent designation by Latreille (1810): Scarabaeus fimetarius Linnaeus, 1758.

Species

These 44 species belong to the genus Aphodius, including 21 extinct species.

• Aphodius beloni <small>Mulsant & Godart, 1879</small>
• Aphodius calichromus <small>Balthasar, 1932</small>
• Aphodius cardinalis <small>Reitter, 1892</small>
• Aphodius clypeatus <small>Fischer von Waldheim, 1821</small>
• Aphodius coniugatus <small>(Panzer, 1795)</small>
• Aphodius corallifer <small>Koshantschikov, 1913</small>
• Aphodius crux <small>Wiedemann, 1823</small>
• Aphodius elegans <small>Allibert, 1847</small>
• Aphodius fasciger <small>Harold, 1881</small>
• Aphodius fimetarius <small>(Linnaeus, 1758)</small>
• Aphodius foetidus <small>(Herbst, 1783)</small>
• Aphodius guangdongensis <small>Maté, 2008</small>
• Aphodius irregularis <small>Westwood, 1839</small>
• Aphodius marginatus <small>Fischer von Waldheim, 1842</small>
• Aphodius micros <small>Walker, 1871</small>
• Aphodius minatorius <small>Péringuey, 1908</small>
• Aphodius pedellus <small>(Degeer, 1774)</small>
• Aphodius pedrosi <small>Wollaston, 1854</small>
• Aphodius plasoni <small>Käufel, 1914</small>
• Aphodius reginae <small>Král, 1997</small>
• Aphodius spadix <small>Schmidt, 1916</small>
• Aphodius swaneticus <small>Reitter, 1892</small>
• Aphodius thoracicus <small>Fischer von Waldheim, 1842</small>
• † Aphodius aboriginalis <small>Wickham, 1912</small>
• † Aphodius anteactus <small>Krell, 2000</small>
• † Aphodius bosniaskii <small>Handlirsch, 1907</small>
• † Aphodius brevipennis <small>Heer, 1862</small>
• † Aphodius charauxi <small>Piton, 1940</small>
• † Aphodius cretaceous <small>Nikolajev, 2008</small>
• † Aphodius florissantensis <small>Wickham, 1911</small>
• † Aphodius granarioides <small>Wickham, 1913</small>
• † Aphodius helvolus <small>Statz, 1952</small>
• † Aphodius inundatus <small>Wickham, 1914</small>
• † Aphodius krantzi <small>Heyden & Heyden, 1866</small>
• † Aphodius laminicola <small>Wickham, 1910</small>
• † Aphodius mediaevus <small>Wickham, 1914</small>
• † Aphodius meyeri <small>Heer, 1847</small>
• † Aphodius praeemptor <small>Wickham, 1913</small>
• † Aphodius precursor <small>Horn, 1876</small>
• † Aphodius schlickumi <small>Statz, 1952</small>
• † Aphodius senex <small>Wickham, 1914</small>
• † Aphodius shoshonis <small>Wickham, 1912</small>
• † Aphodius theobaldi <small>Krell, 2000</small>
• † Aphodius vectis <small>Krell, 2019</small>

The species of the following genera were formerly classified in Aphodius:

Acrossus <small>Mulsant, 1842</small>

Agoliinus <small>Schmidt, 1913</small>

Alloblackburneus <small>Bordat, 2009</small>

Ballucus <small>Gordon & Skelley, 2007</small>

Blackburneus <small>Schmidt, 1913</small>

Calamosternus <small>Motschulsky, 1859</small>

Caligodorus <small>Gordon & Skelley, 2007</small>

Cephalocyclus <small>Dellacasa et al., 1998</small>

Chilothorax <small>Motschulsky, 1859</small>

Cinacanthus <small>Schmidt, 1913</small>

Coelotrachelus <small>Schmidt, 1913</small>

Colobopterus <small>Mulsant, 1842</small>

Cryptoscatomaseter <small>Gordon & Skelley, 2007</small>

Dellacasiellus <small>Gordon & Skelley, 2007</small>

Dialytodius <small>Gordon & Skelley, 2007</small>

Drepanocanthoides <small>Schmidt, 1913</small>

Eupleurus <small>Mulsant, 1842</small>

Flaviellus <small>Gordon & Skelley, 2007</small>

Geomyphilus <small>Gordon & Skelley, 2007</small>

Haroldiellus <small>Gordon & Skelley, 2007</small>

Hornosus <small>Dellacasa, Dellacasa & Gordon, 2015</small>

Irrasinus <small>Gordon & Skelley, 2007</small>

Labarrus <small>Mulsant & Rey, 1869</small>

Lechorodius <small>Gordon & Skelley, 2007</small>

Liothorax <small>Motschulsky, 1859</small>

Luxolinus <small>Gordon & Skelley, 2007</small>

Maculaphodius <small>Gordon & Skelley, 2007</small>

Melinopterus <small>Mulsant, 1842</small>

Mendidius <small>Harold, 1868</small>

Merogyrus <small>Gordon & Skelley, 2007</small>

Neotrichonotulus <small>Dellacasa et al., 2004</small>

Nialaphodius <small>Kolbe, 1908</small>

Orodaliscoides <small>Schmidt, 1913</small>

Oscarinus <small>Gordon & Skelley, 2007</small>

Otophorus <small>Mulsant, 1842</small>

Oxyomoides <small>Dellacasa et al., 2016</small>

Pardalosus <small>Gordon & Skelley, 2007</small>

Phaeaphodius <small>Reitter, 1892</small>

Planolinellus <small>Dellacasa & Dellacasa, 2005</small>

Planolinoides <small>Dellacasa & Dellacasa, 2005</small>

Planolinus <small>Mulsant & Rey, 1869</small>

Pseudagolius <small>Schmidt, 1913</small>

Rugaphodius <small>Gordon & Skelley, 2007</small>

Scabrostomus <small>Gordon & Skelley, 2007</small>

Schaefferellus <small>Gordon & Skelley, 2007</small>

Setodius <small>Gordon & Skelley, 2007</small>

Stenotothorax <small>Schmidt, 1913</small>

Strigodius <small>Gordon & Skelley, 2007</small>

Tetraclipeoides <small>Schmidt, 1913</small>

Teuchestes <small>Mulsant, 1842</small>

Trichaphodioides <small>Paulian, 1942</small>

Trichonotulus <small>Bedel, 1911</small>

References