Amarygmini is a tribe of darkling beetles in the family Tenebrionidae. There are more than 80 genera in Amarygmini.
Description
Adult Amarygmini are active long-legged beetles that fly readily. They have an unhidden membrane between the front margin of the clypeus and the labrum. In males, the base of the aedeagus is asymmetric, more or less acutely drawn out, directed towards the right and fused with the parameres. The dorsal side of the aedeagus is directed towards the dorsum of the body.
Most larvae of Amarygmini have a regular, oval excavation on the ninth abdominal tergite. The Australian Chalcopteroides is an exception, instead having short urogomphi borne on an oblique projection.
Life cycle
Like other beetles, Amarygmini go through complete metamorphosis with the life stages of egg, larva, pupa and adult. Amarygmini are one of several tenebrionid groups to have long-lived larvae and comparatively short-lived adults.
Ecology
Most Amarygmini adults are associated with tree bark or dead wood, especially that which is covered/mixed with fungi or lichen. They occur in both natural (e.g. primary and secondary forests) and artificial (e.g. street trees, parks, gardens) habitats. They are nocturnal. A few species are attracted to artificial light. Larvae are believed to feed on wood (xylophagous) or fungi (mycophagous).
At least some larvae of Chalcopteroides live in soil. Some species of this genus are believed to be scavengers or predators, based on the presence of arthropod fragments in their guts.
Genera
These genera belong to the tribe Amarygmini:
- Alienoplonyx <small>Bremer, 2019</small> (Indomalaya)
- Alymon <small>Pascoe, 1866</small> (tropical Africa)
- Amarygmus <small>Dalman, 1823</small> (the Palearctic, Indomalaya, Australasia, and Oceania)
- Asthenochirus <small>Fairmaire, 1885</small> (tropical Africa)
- Asyleptus <small>Péringuey, 1896</small> (tropical Africa)
- Atropsorodes <small>Ardoin, 1963</small> (tropical Africa)
- Axynaon <small>Blackburn, 1897</small> (Australasia)
- Azarelius <small>Fairmaire, 1892</small> (Indomalaya)
- Bunamarygmus <small>Masumoto, 1988</small> (Indomalaya)
- Cantaloubeus <small>Ardoin, 1959</small> (tropical Africa)
- Caudamarygmus <small>Bremer, 2001</small> (Indomalaya)
- Cephalamarygmus <small>Bremer, 2001</small> (Indomalaya)
- Cerysia <small>Bremer, 2001</small> (Indomalaya and Australasia)
- Chalcoplonyx <small>Ardoin, 1963</small> (tropical Africa)
- Chalcopteroides <small>Gebien, 1948</small> (Australasia)
- Cheiroplus <small>Ardoin, 1963</small> (tropical Africa)
- Cleognathus <small>Gebien, 1921</small> (tropical Africa)
- Coccimarygmus <small>Ardoin, 1966</small> (tropical Africa)
- Crypsinous <small>Fairmaire, 1891</small> (tropical Africa)
- Cymatothes <small>Dejean, 1834</small> (North America and the Neotropics)
- Dalmanius <small>Bremer, 2001</small> (Indomalaya)
- Dasyplonyx <small>Bremer, 2014</small> (Indomalaya)
- Dichotymus <small>Fairmaire, 1891</small> (tropical Africa)
- Erycastus <small>Fairmaire, 1897</small> (tropical Africa)
- Euglyptonotus <small>Gestro, 1901</small> (tropical Africa)
- Eulytus <small>C.O. Waterhouse, 1882</small> (tropical Africa)
- Eumolpamarygmus <small>Pic, 1923</small> (Indomalaya)
- Eumolparamarygmus <small>Bremer, 2006</small> (Indomalaya)
- Eupezoplonyx <small>Pic, 1922</small> (Indomalaya)
- Eupezus <small>Dejean, 1834</small> (tropical Africa)
- Euspinamarygmus <small>Masumoto, 1989</small> (Indomalaya)
- Fahraeus <small>Ardoin, 1963</small> (tropical Africa)
- Falsastenochirus <small>Pic, 1938</small> (tropical Africa)
- Falsoplonyx <small>Ardoin, 1963</small> (tropical Africa)
- Falsosynopticus <small>Pic, 1936</small> (tropical Africa)
- Garambanus <small>Ardoin, 1964</small> (tropical Africa)
- Gonocnemis <small>J. Thomson, 1858</small> (tropical Africa and Indomalaya)
- Gonocnemocistela <small>Pic, 1935</small> (tropical Africa)
- Hesseodes <small>Ardoin, 1963</small> (tropical Africa)
- Hoplobrachium <small>Fairmaire, 1886</small> (tropical Africa and Indomalaya)
- Hoplonyx <small>J. Thomson, 1858</small> (tropical Africa)
- Hypamarygmus <small>Gebien, 1904</small> (tropical Africa)
- Insolitoplonyx <small>Bremer, 2014</small> (Indomalaya)
- Isopteroplonyx <small>Bremer, 2006</small> (Australasia)
- Javamarygmus <small>Pic, 1928</small> (Indomalaya)
- Lemoultia <small>Chatanay, 1913</small> (tropical Africa)
- Lobatopezus <small>Pic, 1952</small> (Indomalaya)
- Luzonoplonyx <small>Bremer, 2009</small> (Indomalaya)
- Macrosynopticus <small>Pic, 1922</small> (Indomalaya)
- Megacantha <small>Westwood, 1843</small> (tropical Africa)
- Meracantha <small>W. Kirby, 1837</small> (North America)
- Meroxys <small>Ardoin, 1963</small> (tropical Africa)
- Mimosynopticus <small>Pic, 1922</small> (tropical Africa)
- Neoplonyx <small>Ardoin, 1963</small> (tropical Africa)
- Nepaloplonyx <small>Bremer, 2014</small> (Indomalaya)
- Nesioticus <small>Westwood, 1843</small> (tropical Africa)
- Oplocheirus <small>Lacordaire, 1859</small> (tropical Africa)
- Overlaetia <small>Pic, 1937</small> (tropical Africa)
- Paragonocnemis <small>Kraatz, 1899</small> (tropical Africa and Indomalaya)
- Paramarygmus <small>Quedenfeldt, 1885</small> (tropical Africa)
- Pilosoplonyx <small>Bremer, 2014</small> (Indomalaya)
- Pimelionotus <small>Ardoin, 1963</small> (tropical Africa)
- Platypsorodes <small>Ardoin, 1963</small> (tropical Africa)
- Plegacerus <small>Gebien, 1921</small> (tropical Africa)
- Plesiophthalmus <small>Motschulsky, 1857</small> (North America, tropical Africa, and Australasia)
- Plinthochrous <small>Fairmaire, 1891</small> (tropical Africa)
- Podacamptus <small>Ardoin, 1964</small> (tropical Africa)
- Pontianacus <small>Fairmaire, 1898</small> (Indomalaya)
- Pseudalymon <small>Ardoin, 1969</small> (tropical Africa)
- Pseudoogeton <small>Masumoto, 1989</small> (the Palearctic and Indomalaya)
- Psilocastus <small>Ardoin, 1963</small> (tropical Africa)
- Psoroderes <small>Ardoin, 1962</small> (tropical Africa)
- Psorodes <small>Dejean, 1834</small> (tropical Africa)
- Psorophodes <small>Ardoin, 1963</small> (tropical Africa)
- Pterodes <small>Ardoin, 1963</small> (tropical Africa)
- Pubamarygmus <small>Pic, 1915</small> (Australasia)
- Reichenspergeria <small>Wasmann, 1921</small> (Indomalaya)
- Seorsoplonyx <small>Bremer, 2010</small> (Indomalaya)
- Singapura <small>Gebien, 1925</small> (Indomalaya)
- Spathulipezus <small>Gebien, 1921</small> (Australasia)
- Spinodietysus <small>Pic, 1927</small> (Indomalaya)
- Stemmoderus <small>Spinola, 1842</small> (tropical Africa)
- Sylvanoplonyx <small>Bremer, 2010</small> (Indomalaya)
- Timogebienus <small>Ardoin, 1963</small> (tropical Africa)
- Trichamarygmus <small>Carter, 1913</small> (Australasia)
- Umslatus <small>Péringuey, 1899</small> (tropical Africa)
- Vutsimus <small>Péringuey, 1899</small> (tropical Africa)
- Ziaelas <small>Fairmaire, 1892</small> (Indomalaya)
Gallery
References
Further reading